广西融水特色红心杉木优树材质性状变异规律研究

[目的]通过测定和分析广西融水种源红心杉木优树材质性状指标,以了解该地区红心杉木材性状变异规律。[方法]以广西融水种源60株20年生的红心杉木优树为研究材料,测定单株材积、红心率、基本密度、组织比量、管胞性状、微纤丝角等10个材质性状指标,分析各材性性状指标分布和变异规律及性状间的相关关系。[结果]表明:红心率、基本密度、木射线比量、管胞比量、管胞长、管胞宽、管胞长宽比等性状数据分布服从正态分布。融水两个地区红心杉优树轴向薄壁细胞比量的变异系数分别为35.08%和44.97%,变异较大。管胞比量变异系数分别为3.28%和3.56%,变异较小。红心率、木射线比量、管胞长、管胞宽、微纤丝角等性状差异极显著(P0.01),轴向薄壁细胞比量差异显著(P0.05)。10个材质性状间存在12对表型显著相关。[结论]测定的10个性状均是连续性数量性状。早、晚材的管胞长度、管胞宽度和管胞长宽比等性状数据呈从心材至边材逐渐增加的规律。作为重要经济性状的红心率与木射线比量呈极显著正相关(P0.01),单株材积与轴向薄壁细胞比量呈显著正相关(P0.05),而单株材积与基本密度的相关性不显著,这使红心杉木生长量与材性相结合的遗传改良成为可能。     

The influence of heartwood on the pulping properties of Acacia melanoxylon wood

The pulping wood quality of Acacia melanoxylon was evaluated in relation to the presence of heartwood. The sapwood and heartwood from 20 trees from four sites in Portugal were evaluated separately at 5% stem height level in terms of chemical composition and kraft pulping aptitude. Heartwood had more extractives than sapwood ranging from 7.4% to 9.5% and from 4.0% to 4.2%, respectively, and with a heartwood-to-sapwood ratio for extractives ranging from 1.9 to 2.3. The major component of heartwood extractives was made up of ethanol-soluble compounds (70% of total extractives). Lignin content was similar in sapwood and heartwood (21.5% and 20.7%, respectively) as well as the sugar composition. Site did not influence the chemical composition. Pulping heartwood differed from sapwood in chemical and optical terms: lower values of pulp yield (53% vs 56% respectively), higher kappa number (11 vs. 7), and lower brightness (28% vs 49%). Acacia melanoxylon wood showed an overall good pulping aptitude, but the presence of heartwood should be taken into account because it decreases the raw-material quality for pulping. Heartwood content should therefore be considered as a quality variable when using A. melanoxylon wood in pulp industries     

柚木种源生长和与材质有关特性的遗传变异及综合评价

对海南岛乐东尖峰岭27年生柚木6个种源的生长及与材质有关特性进行测定与分析,结果表明:种源间树皮厚度、心材宽度、基本密度差异极显著,树高、材积、枝下高、边材宽度差异显著,而胸径、边材年轮数及心材率差异不显著,说明在种源水平上对柚木进行生长和与材质有关特性的改良具有较大的潜力,尤其进行木材生长性状、基本密度和心材宽度的种...     

Variability of wood properties in two wild cherry clonal trials

There has been an increase in wild cherry cultivation over recent decades, revealing a need for improvements in planting material to be used for the production of high quality timber. To date, trial tests have been carried out mainly on growth and tree architectural traits, while no investigations have focused on wood properties. The present study investigated the variability of the growth traits and the physical and mechanical properties of wild cherry wood, both within the tree, within and among clones and between sites. The aims of the study are to provide useful information on the genetics of wood traits in valuable hardwoods and to discuss how within-tree variation can affect early selection of genotypes. The results suggest that site was the most important source of variation in growth traits and in the proportion of heartwood, although differences among clones were highly significant. The number of sapwood rings was very homogeneous both between sites and among clones. Concerning the wood traits, most variation was detected within the tree, rather than within or among clones or between sites. Within tree variability seems to be mainly due to heartwood/sapwood presence. However, since there is a high correlation between heartwood and sapwood properties, a quite efficient selection made on the basis of sapwood should provide an efficient means of selecting heartwood traits.     

Chemical characterization of Pinus halepensis sapwood and heartwood

ABSTRACT

This paper describes the chemical composition of sapwood (SW) and heartwood (HW) of Pinus halepensis Mill stem. Extractives were first isolated by accelerated solvent extraction and then analysed by gas chromatography-mass spectrometry (GC-MS). The cellulosic polysaccharide content present in the pre-extracted wood samples was determined with acid hydrolysis and GC. The hemicelluloses content was determined with acid methanolysis and GC. Free monomers were additionally analysed by GC. The amount of lignin was determined gravimetrically by the Klason lignin method and the acid-soluble lignin was determined by a UV method. Formic and acetic acids in wood were determined after alkaline hydrolysis and analysed by HP-SEC. It was found that lipophilic and hydrophilic extractives were more abundant in heartwood (1.6% and 2.5%) than in sapwood (1.1% and 1.8%). Celluloses content was higher in sapwood (42.5%) than in heartwood (39.7%), whereas lignin, hemicelluloses and sugar monomer contents were more abundant in heartwood (28.9%, 26.8% and 0.3%) than in sapwood (28.0%, 24.5% and 0.2%). The variation in acetic and formic acids and ash contents between sapwood (0.7%, 0.2% and 0.5%) and heartwood (0.6%, 0.1% and 0.4%) was small. The acetylation degrees were found to be slightly similar in sapwood (0.4) and heartwood (0.3).     

Genetic control of wood colour, density and heartwood ellagitannin concentration in European oak (Quercus petraea and Q. robur)

Pressler cores of young clones of Quercus petraea and Q. roburwere analysed from two sites in Germany. Variation of wood colour(defined by the CIELAB system), density and heartwood ellagitanninswas examined between clones, species and sites. Similar studieswere made of cores from another trial of 20 half-sib familiesof parent trees deriving from five German forests. The resultsfrom both the clonal and progeny trials indicate that heartwoodellagitannin content and wood density are under strong geneticcontrol, while sapwood and heartwood colour are less so. A largeproportion of between-clone variation in wood density and ellagitanninlevels occurred between the two species, while these propertiesvaried little among ramets of the same clone grown on two contrastingsites, despite a difference in growth rates. A large proportionof the total variation among progeny was attributed to forestorigins and could not be attributed clearly to either geneticor environmental causes because of limitations of the samplingdesign     

Resistance of Pinus leucodermis heartwood and sapwood against the brown-rot fungus Coniophora puteana

Abstract

This study assessed the decay resistance of Pinus leucodermis wood to the brown-rot fungus Coniophora puteana. Based upon the median weight losses of 30.65% for heartwood and of 34.68% for sapwood obtained in the biological tests, both the heartwood and sapwood material examined was classified as not durable (durability class 5) according to the CEN/TS 15083-1 classification. Total extractives were low, 3.93% in heartwood and 1.00% in sapwood, while lignin content was 22.60% and 25.41% in heartwood and sapwood, respectively. It is highly recommended to use protective treatments before using P. leucodermis wood in outdoor conditions.     

三倍体毛白杨新无性系木材气干密度和力学性质性状的遗传控制及对单板材育种的意义

该文利用 9个 9年生三倍体毛白杨无性系木材试样 ,研究了木材气干密度组成及力学性质的遗传变异规律 .结果表明 ,木材气干密度组成及力学性质在无性系间存在显著或极显著差异 ,并受到中等强度的遗传控制 ;气干密度的径向和纵向变异与多数研究结果一致 ;力学性质除抗弯弹性模量和端面硬度外其株内变异趋势也基本符合木材学理论 .在力学性质指标中 ,抗弯弹性模量和弦面硬度是遗传性很强的性状 ,其无性系重复力分别为 0 90和 0 80 ,抗弯强度、顺纹抗压强度和硬度的无性系重复力稍低 .遗传相关表明对单板材可通过木材密度与干形等形质指标进行优良无性系选择     

13种杨树木材物理力学性质的研究

采集1－69杨等10种黑杨的9年生试材，易县毛白杨等3种白杨的13年生试材，测试木材物理力学性能，分析心边材，不同树高位置，株间的变化规律，比较品种间木材性质的差异，试验结果表明，13种杨树全树木材心材率在14．6－34．1％，之间，生材含水率在86．2－148．5％之间，由生材干燥至含水率为12％所产生的干缩率在8．66％－11．96％这间，气干密度属低类或很低类，强度属弱类，硬度属软类或甚软类，冲击韧性中等；黑杨心材颜色深，心边材性质差异大，白杨心材颜色淡，心边材性质差异小，品种间木材物理力学性质差异较大，按其主要特点可将13个品种的木材分为4类。     

杉木无性系木材干缩性与吸水性的变异

以102个17年生杉木无性系为试材,对木材的干缩性与吸水性进行测定分析发现,木材干缩性、吸水性在杉木无性系间差异极显著;干缩性变异幅度为3.1%～20.0%,表型变异系数达59.8%;吸水性变异幅度在214.5%～338.1%间,表型变异系数为13.2%。木材干缩性与吸水性均具有明显的无性系方差分量,表明杉木木材干缩性与吸水性受遗传控制。进一步研究显示,杉木无性系木材干缩性遗传变异系数为38.6%,重复力为0.390,均属中低水平;吸水性遗传变异系数值虽较低,但重复力相对较高(0.664),属中上水平;在不同入选率(10%、20%、30%)下,木材干缩性与吸水性随入选率的降低其遗传增益值(绝对值)不断提高。研究还发现,杉木木材干缩性与木材基本密度、心材比间为极显著遗传负相关,与木材吸水性间则表现为极显著遗传正相关;木材吸水性在遗传水平上与胸径、材积、心材比间呈极显著正相关,而与木材基本密度间为极显著负相关。     

Variation of extractives content in heartwood and sapwood of Eucalyptus globulus trees

The variation in extractives content in sapwood and heartwood was investigated among 12 trees in each of four commercial plantations of Eucalyptus globulus in central Portugal. The study was carried out at the 15% height level and extractions used successively dichloromethane, ethanol and water. At all sites, heartwood had significantly more extractives than sapwood, on average 3.8 and 2.4%, respectively. Most extractives consisted of ethanol soluble material (on average 52% of total extractives). Among the sites, there was a statistically significant difference in the content of extractives but the most important source of variation was the within-tree variation between sapwood and heartwood. Differences in the content of extractives were also observed among trees. A strong relation between extractives content and heartwood proportion was found. The potential loss of pulp yield and problems associated with accumulation of extractives are directly related to the heartwood proportion in the eucalypt stems. Forest management should take into account heartwood development and selection for minimising heartwood extractives.     

Inter-clonal, intra-clonal, and single tree variations of wood anatomical properties and specific gravity of clonal ramets of Dalbergia sissoo Roxb.

Inter-clonal, intra-clonal and within tree variations in specific gravity and wood anatomical properties of 8-year-old grown ramets of Dalbergia sissoo Roxb. have been investigated. Radial and location-wise intra-clonal variations were non-significant for anatomical properties and specific gravity for all six clones at all three sites. However, inter-clonal variations in wood anatomical properties and specific gravity were significantly different. Inter-clonal variations in anatomical properties and specific gravity were also significant due to sites, which indicated that site-characteristics overshadowed the genetic priority of different clones for wood anatomical properties. Average fiber-characteristics of all clones showed the best performance at Lalkuan, Haldwani (site III), while average specific gravity performed well at Brandis Road, Dehradun (site I) followed by site III (Lalkuan, Haldwani) and site II (Lachchiwala, Dehradun). Within tree variations in anatomical properties like fiber length, fiber diameter, wall thickness, vessel member length and vessel member diameter due to vertical or radial direction and location (pith to periphery) are non-significant. Radial direction, location and height showed no impact on wood element variation. It indicated that there is no impact of juvenile wood, sapwood and heartwood ratio, and reaction wood on wood anatomical properties of 8-year-old ramet of D. sissoo. It further indicated that clone raised ramet of 8-year-old D. sissoo showed the characteristics of mature wood. Within tree variations in specific gravity were significant due to height, which may be related to differential sapwood and heartwood ratio in the vertical direction. Different wood elements viz. fiber length, fiber diameter, wall thickness, vessel member length and vessel member diameter showed significant correlations with each other and with specific gravity.     

Influence of site in the wood quality of Eucalyptus in plantations in Brazil

Genetic materials interact with the environment in specific ways, therefore their responses in wood need to be better understood. The present study sought to evaluate the influence of different climatic conditions in Minas Gerais, Brazil on Eucalyptus grandis × Eucalyptus urophylla wood. Two six-year-old clones were cultivated in Belo Oriente, Ferros, Santa Bárbara, Itabira and Peçanha. Five trees from each clone were sampled from the base to top direction. Wood density, heartwood/sapwood ratio, vessels, fibres, and extractive and lignin contents were determined, as well as their correlation with environmental variables. Vessel diameters, heartwood/sapwood ratio, and extractive and lignin contents all increased due to higher water availability in Santa Bárbara and Itabira. In both genetic materials, the variables linked to growth and wood quality were related to environmental factors.     

Variation analysis and fine family selection on half-sib progenies from clonal seed orchard of Pinus massoniana

Variation of traits that include height, diameter at breast height (DBH), stem volume, crown width, straightness, wood basic density, tracheid length and width of open-pollinated progenies from clonal seed orchard of Pinus massoniana was analyzed. Differences in traits such as growth, tree form and wood quality among families was significant, which indicated that the improvement potential was notable. Heritability of height and wood basic density reached 0.691 and 0.640, respectively. According to a correlation analysis between traits, stem straightness could be improved indirectly when growth traits are improved; growth traits were significantly correlated with wood basic density; and wood basic density was not significantly correlated with length and width of the tracheid. Under the selection ratio of 10%, genetic gains were 19.74% and 19.32% respectively, and selected families from the progeny test stands at the age of five years and eight years were the same. Genetic correlation of height, DBH and stem volume was also significant at the level of 0.01. These show that selection of the short rotation of P. massoniana for pulp and paper use would be efficient for five-year-old trees. According to the comparison of the heritability of all the traits and taking into account the result of canonical correlation analysis, height could be considered as the most important index for selection. Given wood quality of all the families, and flowering and fruit settings of the clones in a seed orchard, twelve fine families were selected at 10% selection ratio of height growth and stem volume. Translated from Scientia Silvae Sinicae, 2005, 41(6): 43–49 [译自: 林业科学]     

Profile of bordered pit aspiration in Cryptomeria japonica using confocal laser scanning microscopy: pit aspiration and heartwood color

Nine trees of Cryptomeria japonica from six elite tree clones with a broad range of heartwood colors were selected. The profiles of pit aspiration percentage (ASP) of earlywood and latewood from pith to bark for green and air-dry conditions were determined to study the relationship between heartwood color and pit aspiration. Confocal laser scanning microscopy (CLSM) observations showed that the ASP of earlywood was low in sapwood and high in heartwood in the green condition. Pit aspiration increased in intermediate wood when compared with sapwood. On the other hand, latewood pits did not aspirate during heartwood formation. Comparing the air-dry condition with the green condition, sapwood pits aspirated during drying in both earlywood and latewood; however, there was no significant difference in pit aspiration of heartwood. There was no significant difference between samples with red and black heartwoods for ASP. The difference in ASP between individual trees was larger than that by heartwood color. The general advantage of CLSM over light microscopy is that serial optical sections along the Z axis can be obtained for any moisture condition, without the need for thin sectioning or embedding.     

Radial patterns of carbon isotopes in the xylem extractives and cellulose of Douglas-fir

Heartwood extractives (nonstructural wood components) are believed to be formed from a combination of compounds present in the adjacent sapwood and materials imported from the phloem. The roles of local compounds and imported material in heartwood formation could have important implications for the wood quality of species having naturally durable wood. Stable isotope composition (delta(13)C) was analyzed to assess radial variation in sapwood extractives, and to estimate the relative importance of adjacent sapwood extractives and imported photosynthate in the formation of heartwood extractives. Cellulose and extractives from the outer 39 annual rings of six Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees were isolated and their delta(13)C composition determined. Although the extractives and the cellulose showed different absolute delta(13)C values, the patterns of change over time (as represented by the annual rings) were similar in most cases. Within an annual ring, carbon isotope ratios of extractives were correlated with the cellulose isotope ratio (R2 = 0.33 in sapwood, R2 = 0.34 in heartwood for aqueous acetone-soluble extractives; R2 = 0.41 in sapwood for hot-water-soluble extractives). These data suggest that some sapwood extractives are formed when the wood ring forms, and remain in place until they are converted to heartwood extractives many years later. Sapwood extractives appear to be important sources of materials for the biosynthesis of heartwood extractives in Douglas-fir.     

Heartwood and sapwood variation in Eucalyptus globulus Labill. trees at the end of rotation for pulp wood production

The amount of heartwood and sapwood in Eucalyptus globulus Labill. wood was determined in commercial pulpwood plantations at harvest age (12 to 15 years) in four ites in central portugal. twelve trees were sampled in each site at nine stem height levels. Heartwood was present in all the trees up to 82%–87% of the total tree height and amounted to 38%–41% of the total tree volume. The heartwood proportion in the stem cross-section decreased from the base upwards, representing 53.3% and 26.1% of the total area respectively at the base and 55% height level. The sapwood width remained relatively constant along the stem at a mean 38 mm up to the 55% height level. The trees within the site, the height level and their interaction were highly significant sources of variation for heartwood proportion and sapwood width. The heartwood diameter and sapwood width were positively correlated with radial tree growth.     

Clonal variation in chemical wood characteristics in Hanönü (Kastamonu) Günlüburun black pine (Pinus nigra Arnold. subsp. Pallasiana (Lamb.) Holmboe) seed orchard

ABSTRACT

Scientific evidence suggests forest cover across the world is gradually decreasing while the world’s population is rapidly increasing. In line with this, the demand on raw wood is also increasing, making it impossible for the already limited natural resources to meet human needs in proportion to the increasing population size. To counter this, there is a need to increase the variety and availability of timber through genetic improvement. This has been seen in genetic improvement studies in forestry which have resulted in seed orchard establishments; however, the genetic variation in these seed orchards continues to be a topic of debate. This study aims to determine the clonal variation in chemical characteristics in a Hanönü (Kastamonu) Günlüburun black pine (Pinus nigra Arnold. subsp. pallasiana (Lamb.) Holmboe) seed orchard. The analysis shows differences between the clones were seen at the 99.9% confidence level. Ash content, NaOH, extractive substance, and hot water solubility exhibited the highest heritability scores, according to the heritability estimates of the clones in the seed orchard. However, hemicellulose content showed an extremely low H₂ (0.1).     

Genetic variation in wood stiffness and strength properties of hybrid larch (Larix gmelinii var. japonica × L. kaempferi)

Genetic parameters for wood stiffness and strength properties were estimated in a 29-year-old hybrid larch stand (Larix gmelinii var. japonica × Larix kaempferi). The study included 19 full-sib larch families from Hokkaido, northern Japan. Implications of these genetic parameters in wood quality improvement are subsequently discussed. Traits included in the analyses were the dynamic modulus of elasticity of green logs (E _log), the modulus of elasticity (MOE), the modulus of rupture (MOR), compression strength parallel to the grain (CS) in small clear specimens, wood density (DEN), and diameter at breast height (DBH). DEN had the lowest coefficients of variation and MOE the highest. The narrow-sense heritability estimates of E _log, MOE, MOR, and CS were 0.61, 0.44, 0.60, and 0.43, respectively, and those of DEN and all mechanical properties increased from an inner to outer position within the stem. E _log and DEN had high positive phenotypic (0.52–0.83) and genetic (0.70–0.92) correlations with MOE, MOR, and CS. The mechanical properties of the inner position of the stem had rather high phenotypic and genetic correlations with those of the outer position and overall mean. The predicted gains in wood stiffness (E _log and MOE) were higher than those of the strength properties (MOR and CS). The predicted correlated responses in MOE, MOR, and CS when selecting for E _log and DEN were 72.6%–97.8% of a gain achievable from direct selection of these traits. DBH showed an insignificant correlation with all mechanical properties, although selection of this trait had a slightly negative effect on the mechanical properties.     

三倍体毛白杨新无性系木材干缩性的遗传分析

利用 9a生三倍体毛白杨测定林的 9个无性系为试材 ,着重研究了木材干缩率的遗传学问题。结果表明 :木材干缩率在无性系间的差异都达到了极显著水平 ,并受到强度遗传控制 ;木材干缩在株内纵向的变异模式是随树高增加而降低 ;全干体积 ,径向、弦向干缩率和气干体积 ,径向、弦向干缩率无性系重复力分别为 0 89,0 95 ,0 84 ,0 6 7,0 84和 0 6 8。