人为干扰对生物多样性的影响

从景观破碎、保护历史、种群行为机制和种群大小4个方面讨论了人为干扰对生物多样性的影响，以期为生物多样性保护和生态环境恢复提供依据。一般人为轻度干扰会导致生物多样性提高，对比研究武夷山人为干扰甜槠林与郁闭稳定甜槠林结果表明，人为干扰甜槠林高等植物物种丰富相对于郁闭稳定林分提高了76．32％；但人为干扰甜槠林的群落优势度却远低于郁闭林，故在演替过程中群落物种多样性的增加是生态系统对外界轻度干扰的一种适应，是恢复生态系统稳定性的一种对策，一旦干扰超过其调节能力，将难以恢复为原来的群落类型。生物在各自进化和适应的过程中由于生态位泛化或特化，许多物种仅局限于非常小的生态位空间，这些物种一旦遭遇到观破碎则很可能从地球上消失，如黄连和短萼黄连对阴湿条件的适应与要求，导致其濒临灭绝；另一些种类由于在群落中零星分布，如斑叶兰、羽萼悬钩子、浙江红花油茶、多种粗叶木（Lasianthus spp.)和一些紫金牛（Ardisia spp.)等，当生境缩小或受到干扰时则濒临灭绝。地史上古老的地区或自然保护历史较长的地区可能拥有更多的物种。保护历史较长的福建省南靖县和溪南亚热带雨林在1200m^2面积内有128种高等植物，同一地带的鼎湖山及和溪周边地区的南亚热带雨林物种丰富均低于此。许多植物是动物传布植物，动物物种多样性的减少影响了这些植物的传布，生境的改变也影响着植物的传布，另一些物种传布到新区域而不能萌发或完成其生活史。地方种群小于一定数量时其灭绝的可能性也增加。     

崀山丹霞地貌区天然常绿阔叶林群落物种多样性研究

 采用样方法采集数据,以重要值为测度指标,以丰富度指数D2、Simpsom指数D1、Shannon-Wiener指数H′、Sheldon均匀度指数JSW和Pielou均匀度指数E为测度指数,研究了崀山丹霞地貌14个天然常绿阔叶林群落物种多样性。结果表明:1)在相似生境中,随着群落演替至亚顶极阶段,物种多样性的变化趋势是增加的,顶极群落中,生境变优,灌、草层物种多样性的变化趋势为增加,而乔木层无明显变化规律;2)群落物种多样性随海拔的变化,除一般常见的变化模式外,还出现一种双峰曲线模式;3)H′较D1,E较JSW具有较高的敏感度。D1与H′在描述群落乔、灌、草,H′与E、D1与E、JSW与E在描述乔木,JSW与E在描述草本之间物种生物多样性上有相似的作用。表明:丹霞地貌中海拔、生境和演替影响植物群落物种多样性,H′、D1JSW与E能单独表达丹霞地貌群落物种多样性。     

人为干扰对常绿阔叶林主要种群分布格局的影响

通过对甜槠群落几个主要种群甜槠、深山含笑、木荷、赤楠分布格局的研究,探讨常绿阔叶林在受到人为干扰后的种群变化动态。结果表明,当群落受不同程度外界扰动后,各种群分布格局呈不同的变化趋势。人为干扰强度越大,甜槠、深山含笑的聚集性越强,而木荷、赤楠的聚集性则下降。在低立木级上,轻度干扰更有利于形成种群的集群分布特征;在中间立木级上,随立木级的增大,种子萌发和萌条能力强的树种分布格局趋势是以重度干扰群落中的聚集性更强;在高立木级上,各种群的分布格局均呈随机分布。     

围封对荒漠草原物种多样性和功能多样性的影响

为揭示荒漠草原区域围封对不同土壤生境物种多样性和功能多样性的影响,以荒漠草原不同土壤生境下围封（15年、10年、6年）和放牧（15年、6年、5年）的草地植物群落为研究对象,基于方差分析和主成分分析的基础上,通过函数拟合分析,探讨植物物种多样性与功能多样性之间的关系。结果表明：（1）围封相对于自由放牧条件,灰钙土生境植物功能多样性显著增加（P<0.05）。随着封育年限的增加,物种多样性呈显著降低趋势（P<0.05）,功能多样性未发生显著变化（P>0.05）;风沙土生境植物功能多样性和物种多样性未发生显著变化（P>0.05）。（2）功能离散度表征的功能多样性与物种丰富度关系可用二次多项式Y=aX²+bX+c形式表示,随着封育年限的增加,二者的相关系数在灰钙土和风沙土生境均呈增加趋势。荒漠草原物种多样性与功能多样性整体正相关,且符合二次函数关系,物种多样性可以用来反映植物群落功能多样性的水平高低,但这种替代作用受封育年限和放牧的影响,随着封育年限增加,物种多样性反映功能多样性水平的能力增加,在未来荒漠草原多样性与生态系统功能关系及维持机制中,需要同时考虑多样性的多维度特征以及不同土壤生境类型。     

天然甜槠林物种多样性的研究

本文运用物种多样性、物种丰富度、物种均匀度等分析方法对福建省周宁县天然甜槠林的群落组成进行研究。结果表明:甜槠群落物种多样性指数和均匀度值均较大,群落结构复杂、物种丰富。研究结果为保护该地域森林植被提供了理论依据。     

古田山常绿阔叶林木本植物与生境的相关性

 用来解释物种丰富的热带雨林物种共存机制的生态位理论和中性理论已经引起了生态学家广泛的关注,而对亚热带常绿阔叶林物种共存机制及物种多样性维持的研究很少。笔者采用卡方检验和典范对应分析方法,对古田山典型常绿阔叶林24hm2样地中胸径不小于10mm、个体数大于18的106种木本植物与平均海拔、坡度和凸度3个小地形生境指标进行生境关联分析。卡方检验结果表明,106种木本植物中,84种(79%)至少与一种生境显著相关(P<0.05);CCA及随机化排列测验方法分析结果表明,大部分物种的分布受生境因子限制;因此,古田山24hm2样地木本植物的分布与生境显著相关,生境特化是亚热带典型常绿阔叶林在局域尺度上木本植物多样性维持的主要机制。     

太行山低山丘陵区不同植物群落物种多样性研究

 通过样地法对小浪底水库库区山地生态系统不同群落间的Shannon-Wiener指数(H′)、Simpson指数(D1)、Margalef物种丰富度(D2)、种间相遇概率(P)和Pielou均匀度指数(Jsw)等多样性指数进行研究,利用统计方法分析不同群落的物种多样性指数与其所处环境的关系。结果表明:1)物种多样性指数和丰富度指数与群落类型有密切关系,同一类型的群落中,多样性指数随演替时间表现出复杂的关系;2)土壤含水量对H′、P、Jsw、D1等呈极显著的正相关性,土壤厚度与H′、D1、P呈显著正相关性;多样性指数与坡度无相关性;土壤有机质、速效氮、速效磷等养分指标与多样性指数H′、D1有显著的正相关;3)多样性指数H′与乔木、灌木群落生物量表现为正相关关系,草本群落与D1相关性不显著;4)乔木林群落的恢复时间与Jsw、H′呈正相关关系,其余多样性指数(P、D1、D2)与恢复时间关系不显著(P>0.05);恢复时间每增加10a,群落物种数平均增加约6种;5)物种多样性指数H′与灌木的盖度和高度呈极显著正相关性;有些单优群落随高度增加和盖度提高,多样性指数反而表现出降低;6)草本植物群落的高度和盖度与物种丰富度指数D2与呈显著的负相关关系,与H′相关性不显著。     

中国特有植物猪血木的濒危原因及保护对策

猪血木（Euryodendron excelsum H.T.Chang）为中国特有的山茶科单型属猪血木属的珍稀濒危植物,现仅分布于广东省阳春县八甲镇地区,且仅残存一个居群,居群内个体数量仅100余株,该物种已被列为国家二级保护植物。物种濒危原因分析是生物多样性保护的关键环节,文章从猪血木的地理分布、生活史特性、种群分布现状及生态环境因素影响等方面分析了该物种的濒危现状,猪血木种群地理分布的局限性限制了其空间拓展;种子萌发和幼苗生长对生境要求特殊和种群现状不利于其更新和发展,人为干扰对猪血木影响严重。其濒危的主要原因是人为因素引起的生境破碎化和过度砍伐所导致的个体数量下降,文章认为现存种群和生境的保护是濒危物种保护的最有效手段,针对猪血木的濒危现状和原因,提出了就地保护,促进自然种群的繁衍更新,建立猪血木繁育基地进行迁地保护和回归引种的保护策略,以期为猪血木的保护和种群恢复提供科学的理论指导,也为中国珍稀濒危物种和生物多样性的保护提供基础资料。     

武夷山黄山松群落物种多样性与种群空问格局的研究

对武夷山黄山松群落物种多样性和种群空间格局的研究结果表明 ,低海拔地区由于受人为干扰影响较大而物种多样性与种群空间分布格局未表现出明显规律性 ,高海拔地区群落物种多样性指数、丰富度和均匀度均有不同程度下降 ,黄山松种群空间格局均呈集群分布 ,且随海拔高度升高而黄山松种群生长受到一定抑制     

宁夏盐池县不同荒漠化治理措施植物多样性研究

 分析不同荒漠化治理措施下植物多样性变化,可以为制定防沙治沙对策和措施提供基础数据。采用α多样性指数(ShannonWiener指数、Simpson指数、群落均匀度指数Jsw和Js)和β-多样性指数(相似性系数Cs),研究宁夏盐池县草原群落在不同荒漠化治理措施(人工封育、退耕还林、撂荒地)固定样地植物多样性变化。对比2002和2003年不同类型固定样地植物多样性观测数据表明,到2003年,各实验样地的α-多样性指数均有不同程度的提高;β-多样性的研究结果同样表明,不同治理措施实验样地的种类组成,已经发生了一定的变化。此外,还进一步分析了引起植物多样性变化的主要原因。     

北京昌平区农业景观野生蜂多样性的时空动态分布

近年来,由于生境质量的下降和生境丧失,野生蜂多样性急剧下降,严重威胁传粉服务和农业的可持续发展。为揭示野生蜂在农业景观镶嵌体中的时空分布格局及不同生境对野生蜂保护的重要性,本研究于2016年4—9月采用网捕法调查了北京昌平农业景观4种主要生境类型(人工林、自然灌木林、荒草地和桃园)中的野生蜂,分析不同生境类型中野生蜂多样性及其时间动态变化特征。结果表明:野生蜂及其中的大体长蜂和独居蜂的物种数和个体数,以及中体长蜂的个体数均在自然灌木林中最高,人工林中小体长蜂个体数最多,野生蜂总个体数和总物种数均在桃园中最低。在不同月份,所有功能群野生蜂均在自然或半自然生境中最高,5月桃园中最低,原因可能与桃园中对地表杂草的集约化管理有关。多度最大的4个优势物种依次为黄胸彩带蜂、铜色隧蜂、黄芦蜂和隧淡脉隧蜂,其在生境间的时间动态特征不同。调查的4种生境均为野生蜂提供了可利用资源,但自然生境和半自然生境的作用更大;其中,自然灌木林维持了较多具有较高潜在传粉效率的中体长和大体长野生蜂,具有更高的保护价值。为了促进该地区农业景观中不同功能群野生蜂多样性及传粉服务,需提高生境类型的多样性,同时在野生蜂活动高峰季节采取低集约化的生境管理方式。     

人类活动对大明山区生物多样性的影响

人类经济活动对生物多样性的影响具有正负效应。强度干扰 ,包括大面积采伐森林、物种的过度利用、大面积的森林火灾和炼山造林、毁林开荒和大规模引入新的物种、农药的施用和环境污染、林区采矿等经济活动导致生物多样性的下降。低强度干扰 (小面积皆伐、择伐和小面积火灾 )增加生物多样性 ,尤其在增加生态系统和景观多样性方面较明显。     

Diversity erosion beyond the species level: Dramatic loss of functional diversity after selective logging in two tropical amphibian communities

Assessing the effects of logging on different aspects of biodiversity and general ecosystem properties is of prime importance if the few remaining areas of intact tropical forest are to be efficiently protected. Commonly used measures of biodiversity may only inadequately reflect actual disturbance after logging and studies restricted to only one specific eco-region do not allow for generalizations of results. We hence measured the impact of selective logging on different levels of diversity of two tropical anuran communities in two geographically distinct eco-regions. Species-diversity patterns were incoherent both, within and between studies. In West Africa, species richness did not differ between primary and exploited forest sites, whereas South American anuran communities exhibited higher species richness in primary sites. Yet, in both eco-regions, functional diversity (FD) was higher in primary forest communities. Absolute values of FD were higher in South American anuran communities, despite higher species richness in West African communities. FD was higher in older recovery, as compared to younger recovery states, even though species-diversity did not differ significantly. Three major conclusions can be drawn from our results. 1. Scale matters: it is important to monitor different levels of biodiversity in order to reveal its actual loss after anthropogenic disturbance. 2. Time matters: the disturbance history of a site is important in order to detect patterns that otherwise remain unnoticed. 3. Geographic history matters at the local scale: whereas general patterns at higher diversity levels were identical in both eco-regions, species richness, species diversity and turnover patterns differed.     

Old manor parks as potential habitats for forest flora in agricultural landscapes of Estonia

Forest biodiversity conservation in intensively managed agricultural landscapes is a constant concern. The dispersal ability of forest plants is, hypothetically, the major limiting factor in fragmented forest landscapes and, therefore, we tested the validity of the theoretical dispersal scheme for plants in fragmented landscapes: ancient forest > woody corridor > emerging forest patch. To this end, we examined the distribution pattern of forest-dwelling plant species in rural landscapes, specifically the occurrence of common forest plant species in old historical forests and in closed-canopy stands of rural ornamental parks, planted on an agricultural land one–two centuries ago.We found that (i) the shade tolerant plant flora in parks’ stands more resemble forests than woody linear habitats (corridors), (ii) nearly 50% of the local forest species pool was present in parks, (iii) the abundance of seed source habitats and the ecological quality of the target habitat determine success rate of colonization. Models predicted that optimal stand characteristics for forest herbs are a minimum area 2.5 ha, canopy closure 0.7–0.8, basal area of trees 10–20 m² ha^?1 and the presence of moderate understory management.We conclude that only patch-type habitats provide suitable environmental conditions for forest-specific plant species. Many common forest plant species are capable of long-distance dispersal between habitat patches across hostile agricultural land, and accordingly, their dispersal follows a modified scheme of the island biogeography, without intermediary role of corridor habitats. Old rural manor parks provide an ecosystem service for nature conservation by harbouring forest biodiversity, and should be considered as potential refugium habitats.     

晋西黄土丘陵沟壑区人工林下草本植物生物多样性研究

以长武王东沟小流域径流小区为试验点,研究了于黄土高原高塬沟壑区坡面表层土壤水分特征,分析了降雨量、植被覆盖度对地表土壤水分的影响,旨在为该区植被恢复和水土流失治理提供科学的理论依据。研究结果表明,0—30cm土层土壤含水量呈现中等变异。15°阳坡土壤含水量小于35°半阳坡土壤含水量。土壤水分的变化趋势与降雨量的变化趋势基本一致,0—10cm土层土壤含水量与降雨量具有良好的同步性,而10—20cm土层和20—30cm土层土壤含水量的同步性较差。植被覆盖度越高,其土壤平均含水量就越高,植被覆盖度达到40%时,中坡和下坡的剖面各层土壤含水量随深度增加而减少的趋势更加明显。土壤平均含水量从坡顶到坡底逐渐增加。0—30cm土层土壤含水量随深度增加而减少。     

东北平原典型玉米种植区农业景观植物多样性研究——以昌图县为例

非耕作生境是农业景观中的重要组成部分,为生物提供食物和栖息地,对生物多样性保护具有重要意义。本文以辽宁省昌图县为例,选取了20个非耕作生境比例为0~50%的网格(1 km?1 km)为样区,采用典型样地法对调查区域内农业景观中的林地、田间路、草地、沟渠、果园5类主要非耕作生境中的植物群落进行调查,探讨不同非耕作生境类型和比例中植物多样性及其与景观异质性之间的关系。研究结果表明,随着非耕作生境比例增加,植物物种多样性和均匀度先增加后减小,在20%~30%时达到峰值;非耕作生境比例在0~40%范围内时,植物物种丰富度同样呈现先增加后减小,且在20%~30%时达到峰值;非耕作生境比例高于40%时,植物物种丰富度再次出现峰值且高于20%~30%。随着非耕作生境比例的增加,农业景观香浓多样性、香浓均匀度指数和景观斑块形状复杂程度逐渐增加,蔓延度恰好相反。不同生境中重要值最高的均为禾本科植物,但植物种不同;优势种优势度为果园沟渠草地田间路林地。果园、草地和沟渠的物种多样性和均匀度指数显著高于林地和田间路,丰富度指数为草地林地沟渠果园田间路,且草地、果园和田间路的不同调查区域间植物群落指数存在较大的变异性。草地、果园、沟渠、田间路和林地中景观异质性与物种多样性相关性顺次降低。研究结论表明,东北平原典型玉米种植区农业景观中,最适宜非耕作生境比例为20%~30%;非耕作生境中林地、沟渠对物种多样性维持、湿生植物多样性保护具有积极作用;不同生境干扰类型和方式不同,景观异质性与植物物种多样性相关性程度也不同。在今后农业景观生物多样性保护中应综合考虑非耕作生境的类型和比例,干扰的类型和方式等多种因素对生物多样性、生态服务及农业产量的影响。     

乌兰布和—西鄂尔多斯荒漠草原过渡带典型灌木群落及其多样性

[目的] 研究乌兰布和沙漠—西鄂尔多斯荒漠草原生态过渡带典型灌木群落的多样性,为该区生物多样性保护和生态环境建设提供理论依据。[方法] 通过重点精查的方式对该地区代表性群落类型设置样点进行植被调查,分析了该过渡带主要灌木群落的类型、物种组成、群落特征和物种多样性。[结果] 该区共有12科30属36种植物。豆科、菊科、藜科和禾本科植物比例较高,单种属较多。多年生草本和灌木居多,以旱生植物为主。灌木群落包括分布在西鄂尔多斯荒漠草原的驼绒藜群落、霸王群落和四合木群落,分布在乌兰布和沙漠的白刺群落、油蒿群落和沙冬青群落。西鄂尔多斯荒漠草原灌木群落的植株密度较高,乌兰布和沙漠灌木群落的建群种重要值、盖度和地上生物量较高。西鄂尔多斯荒漠草原灌木群落的物种多样性普遍高于乌兰布和沙漠。大部分群落的相似性较低。[结论] 乌兰布和—西鄂尔多斯荒漠草原过渡带植物数量比较贫乏,多年生草本和灌木是该区植物的主要部分,植物旱生化明显,西鄂尔多斯荒漠草原植物多样性变化幅度较小且保持在较高水平,不同生境下群落相似性较低。     

Patterns of species change in anthropogenically disturbed forests of Madagascar

Five main conclusions arise from this review of the responses of species to anthropogenic disturbance in Madagascar: First, species’ reactions to anthropogenic disturbance are generally negative, but remain poorly known. Our knowledge is patchy among and within higher taxonomic groups; we are still largely gathering case studies. Second, taxonomic groups vary considerably in which proximate factors are most important. Third, several groups show differing responses within different ecoregions. Whether these differences are consistent across groups requires further testing. Fourth, related species often have divergent reactions to disturbance, even within lower taxonomic groupings (families or genera). Thus, we cannot rely on phylogenetic relatedness or even ecological similarity to infer similarity in responses. Finally, disturbance typically reduces species diversity (especially of native and/or endemic species), but also causes species turnover, typically with forest specialists replaced by grassland generalists, and endemics replaced by non-endemics (including invasives). Given these knowledge gaps, we stress the urgency of applied studies that assess species’ ecology, behaviour and health across disturbance gradients, including purely anthropogenic landscapes. Remaining natural vegetation and protected areas will be unable to preserve Madagascar’s biodiversity under the impact of climatic change; we must understand responses of plants and animals to disturbance in order to create buffer zones and corridors combining secondary, degraded and natural habitats.     

Oribatid communities (Acari: Oribatida) inhabiting saxicolous mosses and lichens in the Krkonoše Mts. (Czech Republic)

Composition, structure and diversity of oribatid communities inhabiting saxicolous mosses and lichens were studied in the Krkonoše Mts. (Czech Republic) along an altitudinal gradient reaching from submontane to alpine belt. Samples of various saxicolous mosses and lichens from 197 stands were collected. Impact of altitude and dominant moss or lichen species on community structure were analysed. Data were evaluated using divisive cluster analysis and direct ordination analysis. Altogether 104 oribatid species were recorded. Four groups of saxicolous habitats, which differ in the composition of their oribatid communities, were distinguished:1. Mosses below the upper forest line with accessory higher plants growing on their surface. Higher plants indicate favourable moisture conditions and an appreciable degree of humus layer development. Their oribatid mite community is rich in number of frequent species and consists of ubiquitous species, ubiquitous species with higher requirements for moisture and amount of decaying organic matter, a high number of soil dwelling species and several hygrophilous species.2. Mosses below the upper forest line without accessory higher plants. They predominantly include mosses with no or only a weakly developed humus layer. Their oribatid mite community is composed mainly of ubiquitous species and a few soil dwelling species.3. Mosses in open areas above the upper forest line. Their humus layer is not developed at all or only weakly. Humidity and temperature fluctuations are here much higher compared with mosses below the upper forest line, which are protected by the specific forest microclimate. Their oribatid mite community is poor in number of frequent and dominant species and consists predominantly of Oribatula cf. pallida and two specialised species living exclusively in mosses and lichens (Mycobates tridactylus and Trichoribates monticola).4. Saxicolous lichens. Their oribatid community comprises ubiquitous species, species frequent both in mosses and lichens and several species with a strong affinity to lichens (Mycobates carli and five species of the genus Carabodes).     

Ground beetles (Carabidae) and edge effect in oak-hornbeam forest and grassland transects

We evaluated ground beetle diversity in relation to forest edge between an oak-hornbeam forest and adjacent herbaceous grassland. To test our hypothesis that the diversity of ground beetles was higher in the forest edge than the interior, pitfall trap samples were taken along two forest-grassland transects in northern Hungary. The diversity of ground beetles was significantly higher at the forest edge and in the grassland than in the forest interior. Ground beetle assemblages in the forest interior, forest edge and grassland could be separated from each other by ordination. Indicator species analysis detected five groups of species: habitat generalists, grassland-associated species, forest generalists, forest specialists, and edge-associated species. Rank correlation indicated leaf litter, herb, canopy cover, and prey abundance as the most important factors influencing carabid diversity. The high diversity of the forest edge resulted from the presence of edge-associated species and of species characteristic of adjacent habitats. Forest edges seem to play an important role in maintaining diversity. Serving as source habitats, edges also contribute to the recolonisation by ground beetles after habitat destruction or other disturbance in the adjacent habitats.