蒸腾速率、阻力与叶内外水势和光强关系的研究

本文采用ＬＩ一１６００稳态气孔计和压力室测定了杨柴的蒸腾速率、阻力、叶水势及其环境因子的日变化，分析结果表明：当０～２ｍ土层内土壤平均水势在－０．０４～－０．２０ＭＰａ时，光照强度是影响杨柴等树木蒸腾速率的主要因子，它主要是通过影响蒸腾阻力所致。光照强度（Ｌｉ）与蒸腾阻力（Ｄｒ）的关系符合方程Ｄｒ＝ａｅｂ／Ｌｉ。这样，根据光照强度对阻力的影响过程，把光照强度与蒸腾速率（Ｔｒ）的关系分成３个区段：①弱光照区，Ｔｒ＝Ｋ／Ｄｒ；②过渡区，Ｔｒ＝Ｋ·ψＬ－Ａ／Ｄｒ；③光饱和区，Ｔｒ＝ａ·ψＬ－Ａ＋ｂ，其中ａ、ｂ和Ｋ是系数。     

杉木人工林针叶光合与蒸腾作用的时空特征

对3个林分(密度A1 667株·hm-2;B3 233株·hm-2;C9 767株·hm-2)的测定和分析表明,树冠不同部位的同龄叶的光补偿点、光饱和点及其最大光合速率不同;树冠同一层内,成熟针叶光合速率依当年生叶、1 a生叶、2 a生叶递减.强光下,中层叶的蒸腾速率明显小于上层叶.弱光下,中层和下层叶具有较高的蒸腾速率,几乎等于甚至超过上层叶.一般而言,杉木针叶光合作用的光饱和点为1 000～1 900 μmol·m-2s-1,光补偿点很低,大致在4.780 ～30.114 μmol·m-2s-1范围.同一枝条不同部位不同年龄针叶光合和蒸腾的能力差异明显.枝条后部针叶净光合速率和蒸腾速率明显小于处于枝中、前部的同龄叶,却明显具有较高的量子利用效率或对弱光(以散射光为主)的利用效率,分别为当年生叶0.006 60 μmolCO2·μmol-1PAR;1 a生叶0.017 94 μmolCO2·μmol-1PAR;2 a生叶0.012 97 μmol CO2·μmol-1PAR. 针叶最大光合速率可达13.335 μmolCO2·m-2s-1 .同一枝条不同部位的杉木针叶的蒸腾效率变化于0.003 5 ～0.007 gCO2·g-1H2O,越靠近枝条后部,蒸腾效率越低.一般而言,阳生叶的光合能力是阴生叶的2～4倍.生长季中当年生针叶光合和蒸腾速率的日变化均有2个峰值,中部和下部当年生叶的变化幅度小于上部针叶.辐射强度、空气湿度或者叶片周围空气的水汽压,是决定叶片气孔导度、光合与蒸腾速率的主要环境因子.呼吸作用白天明显高于夜间,多呈中午单峰型;日变化规律主要受温度、叶片周围空气的水汽压大小控制.蒸腾和呼吸呈紧密线性负相关.3个林分中,无论是2 a生叶,1 a生叶或当年生叶,光合强度均是密度小的最大,密度大的最小.密度越大,叶平均N素含量越低,光饱和点越低,而光补偿点越高,偏向于利用较弱的光,生长也较弱.     

山苍子光合速率、蒸腾速率的研究

在晴天适宜条件下,使用LI-6400 p便携式光合测定系统测定不同年龄阶段的山苍子(Litsea cubeba)连体叶片的光合速率和蒸腾速率。结果表明:山苍子的光合作用日变化均呈双峰型,存在光合午休现象,并且3 a生山苍子的光合速率高于2 a生的,1 a生的最低。且蒸腾速率与光合速率间存在相同的变化趋势。     

黄土半干旱区集水造林条件下林木生长适宜的土壤水分环境

2000年在山西方山，采用LI－6200光合测定系统和LI－1600稳态气孔仪，对不同集水措施试验地的刺槐、侧柏和苹果的生理指标进行了实地观测，并辅以相同树种实地盆栽苗木水分胁迫条件下的林木生理指标观测，研究了林木净光合速率、羧化效率、水分利用效率与土壤含水量的定量关系。结果表明：维持刺槐、侧柏和苹果净光合作用的最佳土壤含水量分别为17.13%、15.9%和16.11%，三树种维持叶片水分利用效率的最佳土壤含水量依次为13.23%、10.66%和12.38%；羧化效率最高时的土壤含水量临界值依次为15.7%、15.5%和15.8%；土壤水合补偿点分别为4.55%、3.91%和4.49%。上述三树种蒸腾速率所适宜的土壤含水量分别为18.82%、19.70%和16.33%。根据集水造林的基本构思，可选取维持最佳叶片水分利用效率的土壤含水量临界值作为集水造林林木密度调控的土壤水分阈值和林木需水量的低限计算条件。这一指标可量化为刺槐10％－13％，苹果10％－12.5%，侧柏9.5%-11%。     

单株平衡法的建立

单株平衡法的建立孙长忠，黄宝龙（西北林学院林学系杨陵７１２１００）（南京林业大学南京２１００３７）关键词黄土高原，蒸腾量，蒸发量，单株平衡法，单株平衡方程，蒸发偏量在黄土高原地区，水分因素对植物生长及生存具有强烈的限制作用。研究植物生长与环境水分条件...     

Water Use of Tree Lines: Importance of Leaf Area and Micrometeorology in Sub-Humid Kenya

In this research the relative importance of leaf area and microclimatic factors in determining water use of tree lines was examined in sub-humid Western Kenya. Measurements of tree water-use by a heat-balance technique, leaf area, bulk air saturation deficit, daily radiation, and soil water content were done in an experiment with tree lines within crop fields. The tree species were Eucalyptus grandis W. Hill ex Maiden, Grevillea robusta A. Cunn. and Cedrella serrata Royle, grown to produce poles on a phosphorus-fixing Oxisol/Ferralsol with (+P) or without (−P) phosphorus application. Doubling the leaf area of Cedrella and Grevillea doubled water use in a leaf area (LA) range of 1–11 m² per tree. The response of Eucalyptus water use (W) to increases in leaf area was slightly less marked, with W = LAⁿ, n<1. Transpiration rate per unit leaf area (Tr) was the other important determinant of water use, being affected by both tree species and phosphorus fertilization. A doubling of the saturation deficit (SD) halved the water use of all trees except for Cedrella +P, in which water use increased. A direct effect of soil water content on water use was only found in Grevillea -P, with a small increase (60%) as available water increased from 1.4 to 8.9% above wilting point (32%). This low direct response to soil water content is probably due to the extensive tree-root systems and the deep clayey soils supplying sufficient water to meet the evaporative demand. Indirect responses to soil water content via decreases in leaf area occurred in the dry season. The results showed that water use of tree lines was more determined by leaf area and transpiration rate per unit leaf area than by micro meteorological factors. The linear response of tree water use to leaf area, over a wide range leaf areas, is a specific characteristic of tree line configurations and distinguished them from forest stands. In tree lines light interception and canopy conductance increase with leaf area much more than a similar leaf area increase would have caused in a closed forest canopy.     

短柄五加蒸腾作用及其与生理生态因子相关性的初步研究

水分是植物生长和分布的主要限制因子 ,因此对植物蒸腾作用特征及与环境因子关系的探讨一直是植物生理生态学研究的重要内容之一。短柄五加 (Ａｃａｎｔｈｏｐａｎａｘｂｒａｃｈｙｐｕｓ)是分布于我国黄土高原地区的濒危灌木树种 ,具有重要的药用价值。目前 ,已有关于短柄五加无性系种群生长和生殖生物学特征的报道 (田国伟等 ,1 998;王仲礼等 ,1 997;1 998ａ ;1 998ｂ ;1 999;岳春雷等 ,2 0 0 1 ) ,但对其生理生态学的研究尚为空白。本文通过对短柄五加在林下和开阔地两种生境下的蒸腾特征及与环境因子的相关性的比较研究 ,旨在从植物水分…     

中亚热带2种原始兰科植物对变化光环境的响应与适应

金佛山兰为国家二级保护植物 ,个体数极少 ,仅见于南川金佛山及附近的稀疏马尾松林下 ,并只与其亲缘种金兰生长在一起。在温室内 ,模拟研究了金佛山兰和金兰对不同强度变化光环境的响应与适应。结果表明 :①在强光下利用便携式光合仪长时间观测植物的光合速率时 ,由于温度的影响 ,应使用温控系统才能获得更准确结果。②对于PAR 80 0到 2 0 0 μmolphotons·m- 2 s- 1 之间任何强度的光照变化 ,金佛山兰和金兰的净光合速率、蒸腾速率、胞间CO2 浓度、气孔导度和水分利用率均保持相对稳定 ,表现出对大幅度变化光环境较好的适应性 ;对于逐渐遮荫至 10 0 μmolphotons·m- 2 s- 1 左右 ,金佛山兰和金兰均以降低水分利用率、提高光能利用率来适应变化的光环境。③不同强度的间断遮荫对二者光合速率等的影响程度不同。 2种原始兰花的净光合速率随间断遮荫均先迅速降至最低 ,然后逐渐回升 ,在恢复光照后 ,金兰比金佛山兰能更快地恢复光合作用 ;蒸腾速率随间断遮荫迅速降低 ,恢复光照后 ,大多能迅速恢复至遮荫前水平。恢复光照后 ,二者的气孔导度出现不同程度的降低。间断遮荫能导致二者胞间CO2 浓度升高约 2 0 %以上 ,恢复光照后 ,大多能迅速恢复到原有水平。所有这些 ,都反映了金佛山兰和金兰对所在地区变化光     

Interactive responses to water deficits and crop load in olive (Olea europaea L., cv. Morisca). II: Water use, fruit and oil yield

To understand the relations between water use and yield in response to crop load, two experiments were conducted in olive (cv. Morisca), during six consecutive years (2002-2007) in an experimental orchard located in Badajoz, Southwest Spain. Experiment 1, assessed the responses during the early years of the orchard (2002-2004) using four irrigation treatments that applied fractions of the estimated crop evapotranspiration (ET_c) (125%, 100%, 75% and 0%) and three crop load levels (100%, 50% and 0% of fruit removal, termed Off, Medium and On treatments). Experiment 2 assessed the response of more mature trees (2005-2007) to three irrigation treatments (115%, 100%, and 60% of ET_c) and the natural crop load which were Off, On, and Medium in 2005, 2006 and 2007, respectively. Yield was reduced by water deficits and so did the estimated tree transpiration which was linearly related to yield (y = 1.2302x − 21.15, R² = 0.8864), showing the high sensitivity of cultivar Morisca to water deficits. The relations between fruit number and fruit weight showed that high crop loads had lower fruit weights and oil yield, a decrease that was more pronounced as water deficits increased. The yield response to water supply in the control and excess treatments, and the observations on the water relations of these two treatments suggest that the calculations made using the FAO method (Doorenbos and Pruit, 1974) with the crop coefficient proposed by Pastor et al. (1998) and the reduction coefficient (Fereres et al., 1982) to apply 100% of ET_c in the control treatment, underestimated the ET_c of the orchard. The results indicate that, although the absence of fruits lead to reduced water use as compared to situations of medium and high crop loads, canopy size was much more determinant of orchard water requirements than crop load.     

Effect of water deficit in different growth stages on stem sap flux of greenhouse grown pear-jujube tree

The effects of water deficit in different fruit growth stages on the variation of stem sap flux of 6-year old greenhouse-grown pear-jujube trees were investigated. Treatments included sufficient water supply during the whole fruit-growing period (T₁), mild water deficit during the flowering–fruit setting stage (T₂), moderate water deficit during the fruit rapid growth stage (T₃) and severe water deficit during the fruit maturing stage (T₄). Results showed that significant compensation effect on stem sap flux after re-watering was observed in T₂, but not in T₃ and T₄ stages. At the end of rapid growth stage, the diurnal variation of stomatal conductance generally had a similar trend as that of stem sap flux, but with a distinct midday depression from 12:00 to 14:00 p.m. In addition, a linear relationship between the relative available soil water content (RAWC) and the ratio of daily stem sap flux to that of sufficient water treatment was observed (R² = 0.4489).