Defoliation pattern, foraging behaviour and diet selection by lactating dairy cows in response to sward height and herbage allowance of a ryegrass-dominated pasture

The effects of sward surface height (SSH) and daily herbage allowance (HA) on the defoliation pattern and grazing mechanics of early lactation dairy cows grazing on irrigated perennial ryegrass–white clover pasture were studied. The hypothesis tested was that SSH and HA affect intake and diet selection through their effects on the pattern of defoliation which is influenced by the resistance to prehension bites. Factorial combinations of two initial SSH (14 and 28 cm) and two daily HA (35 and 70 kg DM cow^?1 d^?1) were examined in a replicated experiment. The peak longitudinal tensile force required to break the sward portion encompassed in a 100 cm² area [bite fracture force (BFF₁₀₀)] was measured as an index of the resistance to prehension. The volume of herbage defoliated and herbage intake increased with SSH (P < 0·05) and HA (P < 0·01). Corresponding to an increase in HA from 35 to 70 kg DM cow^?1 d^?1, there was a proportional increase in the total defoliation area (TDA) and intake by 0·24 and 0·55 in the short sward compared with 0·16 and 0·32 in the tall sward respectively. The results of this experiment suggest that a consistent spatial pattern of reduction of the canopy exists during defoliation by cows and that the volume of sward canopy defoliated is the major variable affecting herbage intake. The BFF increased down the sward profile at a rate that was higher (P < 0·05) for the taller sward than for the shorter sward. It is proposed that a relatively lower resistance to prehension in the short sward compared with the tall sward explains the greater proportionate increase in TDA and intake corresponding to an increase in HA. The rate at which BFF₁₀₀ increases down the sward profile is suggested as a sward physical variable that can influence the defoliation process. The estimated time and energy costs of prehension bites are discussed in the context that larger bites are handled more efficiently than smaller bites.     

The manipulation of grass swards for summer-calving dairy cows

Two experiments examined the effects of different defoliation treatments in spring on sward morphology and animal performance in mid-season and late season. Three treatments were applied in both experiments: Control (C), sward grazed by cows in spring to 6–8 cm grass height. Grazed Aftermath (GA). sward grazed by cows in spring to 3–4cm and allowed to regrow before being grazed by summer-calving cows, Silage Aftermath (SA), sward not grazed in spring, but a primary cut taken and the sward allowed to regrow before being grazed by summer-calving cows. The aim of treatment GA was to produce a sward with a high tiller density and high intake characteristics to meet the forage intake requirements of continuously grazed summer-calving cows, without resorting to offering forage buffers. Experiment 1 was conducted in 1989 on a sandy loam soil and Experiment 2 in 1990 on a heavy loam soil. In both experiments the GA treatment led to high live tiller density and live: dead tiller ratios compared with the C and SA treatments. Differences in sward morphology were also detected by applying double normal distribution analyses to measurements of grass height. The GA treatment also increased sward herbage mass and, to a limited extent, herbage metabolizable energy and crude protein contents. The results from Experiment 1 suggested that these sward effects lead to increased herbage dry-matter intake (as estimated by the n-alkane technique) and milk yield in cows grazing the GA sward. However, in Experiment 2, where conditions for grass growth in mid-season were more favourable than in Experiment 1, the differences in sward morphology produced in spring were quickly lost in June and July. There were therefore no differences in herbage intake or milk yield in the second experiment. Herbage intakes (kgDMd^?1± s.e.d) estimated in July for cows on treatments C, GA and SA were 11·0, 13·4, 10·1 ± 2·16 for Experiment 1 and 10·7, 11·1, 11·2 ± 2·32 for Experiment 2. Average milk yield (kgd^?1± s.e.d.) for cows on treatments C, GA and SA were 26·1, 28·0, 25·6 ± 0·31 (Experiment 1) and 28·5, 27·3, 28·4 + 0·58 (Experiment 2). The results suggested that acceptable milk yields can be obtained from grazing summer-calving cows, without offering forage buffers, by applying high stocking rates (low grass heights) in spring. However, the benefits of this manipulation could be lost by lax grazing in mid-season.     

Canopy characteristics and tillering dynamics of Marandu palisade grass pastures in the rainy–dry transition season

The objective of this study was to evaluate the effect of pasture management during the rainy–dry transition season in Jaboticabal, Brazil, on the structural and tillering characteristics of Marandu palisade grass. The treatments were as follows: 15 cm sward height in the rainy season and ungrazed in the rainy–dry transition season (15/0); 25 cm sward height during the rainy season and fixed stocking rate of 2·5 AU ha^?1 during the rainy–dry transition season (25/2·5); and 35 cm sward height in the rainy season and fixed stocking rate of 2·5 AU ha^?1 in the rainy–dry transition season (35/2·5). There were more herbage mass, stem and dead herbage mass in treatments 25/2·5 and 35/2·5 than 15/0. Tiller appearance rate was greater in treatment 15/0 and tiller mortality rate was higher in treatment 35/2·5 compared to treatment 15/0. Tiller population stability index was 1 in treatment 15/0 and lower in treatment 35/2·5. Although treatment 15/0 showed favourable structural and tillering characteristics, it also has the lowest herbage mass. Treatment 25/2·5 can be regarded as an adequate management strategy for Marandu palisade grass, since it has similar herbage mass and better tillering characteristics than treatment 35/2·5.     

The effect of sward height and its direction of change on the herbage intake, diet selection and performance of weaned lambs grazing ryegrass swards

Abstract Four sward height treatments were imposed by continuous variable stocking using at least ten Suffolk × Greyface lambs per plot from late July to late August: constant 3·5 cm. constant 6·0 cm. 3·5cm increasing to 6·0 cm and 6·0 cm decreasing to 3·5 cm. The treatments were established on two swards given fertilizer N applications over the season of 97 and 160 kg N ha^?1 respectively. Animal density was greater on the high fertilizer treatment, at the lower sward height and especially on the decreasing height treatments. Liveweight change of lambs was higher (P <0·001) on the 6-cm than on the 3·5-cm treatments (+159 vs-13g d^?1 and was also higher (P <0·001) on the increasing than on the decreasing sward height treatments (+92 vs-26 g d^?1). Herbage organic matter intake (OMI), measured on two occasions in the experiment, was greater (P <0·001) on the 6·0 cm than on the 3·5-cm sward heights whereas values for the increasing sward height treatments were much greater than those for the decreasing sward height treatments. There was little difference in the organic matter digestibility of the diet between treatments. Diets were composed largely of lamina, although there was more pseudostem and dead herbage in the diets of Iambs grazing the decreasing than the increasing sward height treatments at the end of the experiment. Bite mass was closely related to OMI but the treatment and period differences were relatively greater than for OMI. Bite mass was more closely related to the depth of the lamina layer (sward height-pseudostem height) than it was to sward height. There was evidence that pseudostem acted as a barrier to defoliation on these short swards and also that the proportion of youngest leaf in the diet was positively related to sward height and to increases in sward height. Sward height and especially the direction of change in sward height, together with associated stock density, were potent influences on lamb growth rate. This was a consequence of differences in herbage intake, which was strongly influenced by bite mass.     

Does post‐grazing sward height influence sward characteristics,seasonal herbage dry‐matter production and herbage quality?

Extending the grazing season through the production and utilization of high‐quality forage is a key objective in grassland‐based dairy production systems. Grazing swards to a low post‐grazing sward height (PGSH) is a strategy for improving grass utilization. A grazing experiment conducted in Ireland investigated immediate and subsequent effects of PGSH on sward production, utilization and structural characteristics. Swards were grazed to 2·7 cm (severe; S1) or 3·5 cm (moderate; M1) from 10 February to 18 April 2010 (Period 1; P1). From 19 April, each P1 paddock was halved and grazed to either 3·8 cm (S2) or 4·8 cm (M2), until 30 October (Period 2; P2). The first grazing rotation was +7 d on S1 swards compared with M1 swards (45 d), due to greater herbage utilization (+0·22). Herbage production during P1 was not affected by PGSH but a severe PGSH during this period reduced subsequent herbage production: 13·9 (S1) vs. 15·5 t dry matter (DM) ha^?1 (M1) by the end of the study. Leaf proportion was increased (+0·10) on S2 swards compared with M2 swards, but M2 swards produced 1·2 t DM ha^?1 more herbage during P2. Despite the relatively lower level of sward utilization obtained from moderate grazing in P1 (3·5 cm) and P2 (4·5–5·0 cm), such levels of PGSH increased DM production while maintaining sward quality, compared with severe grazing (2·7 cm in P1 and 3·5–4·0 cm in P2).     

Influence of defoliation regime on herbage production and characteristics of intake by dairy cows as affected by grazing intensity

Three contrasting defoliation regimes for dairy cows—four cuts annually, severe rotational grazing and lax rotational grazing integrated with cutting—were compared in terms of herbage production, selection and intake per cow. Lax and severe grazing were compared simultaneously and grazing intensity was characterized by means of changes in herbage mass and sward height during grazing.
Herbage growth and yield were similar under cutting, severe grazing and lax grazing/cutting (120 t organic matter (OM) ha⁻¹). Herbage quality and leaf production were highest with severe grazing, which also had a less marked seasonal pattern of growth. The herbage intake per cow was 111 kg OM d⁻¹ with severe grazing and proportionately (0-20) higher at lax grazing/ cutting. 050 of the herbage yield was harvested by cutting at lax grazing/cutting. Selectivity was described with high certainly by the nutrient content prior to defoliation and by the degree of defoliation. Models to quantify this are presented.
Grazing intensity could be described by the size and degree of utilization of the fouled area, and herbage intake was quantified by means of herbage height before and after grazing. Within the range of 80-240 mm sward height prior to grazing, height measurements led to realistic and more accurate estimates than those obtained by measuring organic matter disappearance.
Herbage utilization was depressed significantly by increasing maturity of herbage due to lower nutritive value, but in particular due to reduced green leaf content and increased reproductive development. If sward height did not exceed 250 mm at any time, good utilization by grazing could be obtained.     

Open pasture,silvopasture and sward herbage maturity effects on nutritive value and fermentation characteristics of cool‐season pasture

In Appalachian USA, silvopasture offers promise of increased farm productivity. A synchronized, temporal understanding of open pasture (OP) and silvopasture (SP) nutritive characteristics is essential for grazing system development. We examined pasture‐type nutritive‐value relationships when herbage was harvested based on morphological maturity rather than calendar date. Neutral detergent fibre and acid detergent fibre (ADF and NDF) content were greater in silvopasture, while organic matter (OM) was lower (P < 0·05). Digestibility of SP herbage dry matter (DM) and OM tended (P = 0·10) to be lower (418 vs. 471 and 437 vs. 491 g kg^?1 respectively). Neutral detergent fibre digestibility was greater (P < 0·05) for OP than SP forage (538 vs. 480 g kg^?1), and ADF tended to be greater (P = 0·10; 551 vs. 501). Open‐pasture forage fermentation effluent exhibited slightly higher microbial richness and Shannon diversity than SP. However, overall community composition of both bacteria and archaea did not differ between pasture types or sampling times. Pasture types show proximate analyses differences generally favourable to OP, although both have similar overall nutritive value. In addition, the SP sward exhibited a 4‐ to 6‐days delay in reaching equivalent maturity. Conversion of farm woodlots to SP would increase overall herbage production and improve pasture management flexibility.     

Effect of grazing severity on perennial ryegrass herbage production and sward structural characteristics throughout an entire grazing season

The objective of this study, which was part of a larger grazing‐systems experiment, was to investigate the cumulative impact of three levels of grazing intensity on sward production, utilization and structural characteristics. Pastures were grazed by rotational stocking with Holstein–Friesian dairy cows from 10 February to 18 November 2009. Target post‐grazing heights were 4·5 to 5 cm (high; H), 4 to 4·5 cm (intermediate; I) and 3·5 to 4 cm (low; L). Detailed sward measurement were undertaken on 0·08 of each farmlet area. There were no significant treatment differences in herbage accumulated or in herbage harvested [mean 11·3 and 11·2 t dry matter (DM) ha^?1 respectively]. Above the 3·5 cm horizon, H, I and L swards had 0·56, 0·62 and 0·67 of DM as leaf and 0·30, 0·23 and 0·21 of DM as stem respectively. As grazing severity increased, tiller density of grass species other than perennial ryegrass (PRG) decreased (from 3,350 to 2,780 and to 1771 tillers m^?2 for H, I and L paddocks respectively) and the rejected area decreased (from 0·27 to 0·20 and to 0·10 for H, I and L paddocks respectively). These results indicate the importance of grazing management practice on sward structure and quality and endorse the concept of increased grazing severity as a strategy to maintain high‐quality grass throughout the grazing season. The findings are presented in the context of the need for intensive dairy production systems to provide greater quantities of high‐quality pasture over an extended grazing season, in response to policy changes with the abolition of EU milk quotas.     

Response of herbage regrowth and water-soluble carbohydrate concentration of ryegrass species to defoliation practices when grown in a Mediterranean environment

Three experiments were conducted to determine the association between leaf number per tiller at defoliation, water‐soluble carbohydrate (WSC) concentration and herbage mass of juvenile ryegrass plants when grown in a Mediterranean environment. Seedlings of ryegrass were grown in nursery pots arranged side‐by‐side and located outside in the open‐air to simulate a mini‐sward in Experiments 1 and 2, and a mixture of annual ryegrass and subterranean clover (Trifolium subterraneum L.) was grown in a small plot field study in Experiment 3. Swards were defoliated mechanically with the onset of defoliation commencing within 28 d of germination. Frequency of defoliation ranged from one to nine leaves per tiller, whilst defoliation height ranged from 30 mm of pseudostem height that removed all leaf laminae in Experiment 1, to 50 mm of pseudostem height with some leaf laminae remaining post‐defoliation in Experiments 2 and 3. A positive relationship between herbage mass of ryegrass, WSC concentration and leaf number per tiller at defoliation was demonstrated in all experiments. In Experiment 1, the herbage mass of leaf, pseudostem and roots of tillers defoliated at one leaf per tiller was reduced to 0·10, 0·09 and 0·06 of those tillers defoliated less frequently at six leaves per tiller. However, the reduction in herbage mass from frequent defoliation was less severe in Experiment 2 and coincided with a 0·20 reduction in WSC concentration of pseudostem compared with 0·80 measured during Experiment 1. In Experiment 3, the highest harvested herbage mass of ryegrass occurred when defoliation was nine leaves per tiller. Although the harvested herbage from this sward contained senescent herbage, the in vitro dry‐matter digestibility of the harvested herbage did not differ significantly compared with the remaining treatments that had been defoliated more frequently. Leaf numbers of newly germinated ryegrass tillers in a Mediterranean environment were positively associated with WSC concentration of pseudostem and herbage mass. A minimum period of two to three leaf appearances was required to restore WSC concentrations to levels measured prior to defoliation thereby avoiding a significant reduction in herbage mass. However, maximum herbage mass of a mixed sward containing ryegrass and subterranean clover was achieved when defoliation was delayed to nine leaves per tiller.     

Effect of replacing time available for grazing with time available for eating maize silage and soyabean meal on milk yield and feeding behaviour in dairy cows

Low rates of herbage dry matter (DM) intake impose limits on total daily DM intake in grazing dairy cows. The objective of this study was to increase total daily DM intake and milk production by restricting daily time available for grazing (TAG) and replacing it with time available for eating a maize silage/soyabean meal (TAMS) diet indoors. The treatments (TAG + TAMS) were 20 + 0, 19 + 1, 10 + 10 and 5 + 15 h. Measurements were made of milk production, intake and feeding behaviour. The interactions of TAG + TAMS treatments with sward height (SH) and concentrate level (CL) were also examined. Two experiments, each lasting 42 days, were carried out in spring ( Experiment 1 ) and autumn ( Experiment 2 ) using forty‐eight and twenty‐four Holstein‐Friesian cows respectively. Treatments were arranged in a factorial design with TAG + TAMS treatments, SH ( Experiment 1 only) and CL as the independent variables and a TAG + TAMS of 20 h. Reducing TAG and increasing TAMS significantly reduced estimated herbage DM intake and significantly increased maize silage/soyabean meal intake in both experiments, but there were no significant main effects of TAG + TAMS treatments on milk yield (mean, 27·4 and 25·5 kg d^?1 for Experiments 1 and 2 respectively), and yield of milk constituents. Increasing SH ( Experiment 1 ) and CL ( Experiments 1 and 2 ) significantly increased milk yield. In Experiment 1 , there was a significant interaction between TAG + TAMS treatments and SH with the taller sward height of 8–10 cm and the 20 + 0 treatment having the highest milk yield (29·7 kg d^?1) and the 5 + 15 treatment the lowest (27·2 kg d^?1), whereas at the lower sward height of 4–6 cm, milk yield was lowest on the 20 + 0 treatment (25·5 kg d^?1) with the other three treatments being higher (mean, 26·9 kg d^?1). Replacing TAG with TAMS significantly increased liveweight gain in Experiment 1 but not in Experiment 2 . Estimated rates of intake of herbage were lower in the autumn experiment ( Experiment 2 , 9·6 g DM min ^?1) than in the spring experiment ( Experiment 1 , 29·4 g DM min ^?1) but rates of intake of maize silage were higher in the autumn (112·4 g DM min^?1) than in the spring (72·5 g DM min^?1). In conclusion, in spring the response to replacing TAG with TAMS was dependent on sward conditions with the highest milk fat plus protein yield being on the 20 + 0 treatment at the high sward height and on the 19 + 1 treatment at the low sward height. The high liveweight gain of the 5 + 15 treatment could be an important means of restoring body condition in grazing lactating cows. In autumn, intakes of herbage were low in spite of its high estimated nutritive value with all treatments having a similar level of performance.     

Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

Continuous and rotational grazing of dairy cows – the interactions of grazing system with level of milk yield, sward height and concentrate level

An experiment was conducted to test the hypothesis that for cows with high levels of milk yield, rotational grazing produces higher milk yields than continuous grazing. The comparison of grazing systems was made at two levels of milk yield (initially 20·3 and 32·5 kg d^?1), and interactions with sward height and concentrate level were also examined. The study used 48 multiparous Holstein Friesian cows over a period of 62 d. Mean milk yield, its persistency and composition, live weight, body condition score and liveweight gain were not significantly affected by grazing system at either level of milk yield. There were no significant interactions between grazing system and sward height or concentrate level for any milk production measurement. Mean estimated herbage and total dry matter (DM) intake (P < 0·01), grazing time (P < 0·05) and ruminating time (P < 0·01) were significantly greater on the continuous grazing system. The cows in the higher milk yield group and those grazed at the higher sward height had a significantly (P < 0·05) higher estimated daily herbage DM intake and rate of herbage intake on the continuous grazing system than those on the rotational grazing system. There was no evidence to support the hypothesis that rotational grazing systems support higher levels of milk production than continuous grazing for cows of high milk yield. The shorter grazing time on the rotational grazing system indicated that cows may anticipate the timing of the daily movement of the electric fence, and this reduces their time spent grazing residual herbage.     

The defoliation dynamics of a stockpiled native grassland pasture follow similar patterns between supplemented and unsupplemented beef calves

It is unclear to what extent and on which variables does supplementing beef cattle on native grasslands affect sward structure, specifically on the dynamics of its grazing horizons. Three hypotheses were tested: (i) during a grazing down process under similar forage allowance, supplemented animals take longer to finish each grazing stratum, than their unsupplemented counterparts, (ii) in both cases, the upper stratum will be heavily depleted before the subsequent strata are grazed, (iii) some species and/or forage fractions are consumed faster than others, regardless of the animals being supplemented (corn dried distillers grains with solubles, DDGS, at 0.7% of their body weight, BW, on a dry matter, DM, basis) or not. Three blocks of stockpiled native grasslands were used and split into two treatments plots (n = 6), on which either supplemented (S) or control (C) heifers of 10.6 ± 0.6 months of age and an initial BW of 143 ± 9 kg, were used. A 2.5 × 0.5 m observation grid was installed on the sward, generating 384 observation points on each plot. On these observation plots, sward height (SH) and visually assessed green forage mass percentage (%G) were registered every other day for 12 consecutive days. No differences were found between the horizontal grazing dynamics between supplemented and control animals in terms of how they switched from the upper grazing horizon to the successive ones. In both cases, when the upper grazing horizon was heavily depleted, the subsequent horizon was being depleted by its half. Differences of preference for C3 species over C4 was observed for both treatments, but this effect was more meaningful for control animals. Grazing time never fully compensated for the decline in intake rate during depletion throughout the grazing horizons. Pasture intake declined when the animals transition from grazing the top grazing horizons to the lower horizons, irrespective of the level of supplementation. Managing the sward structure in terms of sward height will be beneficial to maximizing individual animal performance, for both C and S animals. Native grasslands paddocks with a greater C3 gasses predominance will always be preferred to C4 dominated paddocks, regardless of an eventual supplementation practice.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

GrazeIn: a model of herbage intake and milk production for grazing dairy cows. 2. Prediction of intake under rotational and continuously stocked grazing management

Decision support tools to help dairy farmers gain confidence in grazing management need to be able to predict performance of grazing animals with easy‐to‐obtain variables on farm. This paper, the second of a series of three, describes the GrazeIn model predicting herbage intake for grazing dairy cows. The model of voluntary intake described in the first paper is adapted to grazing situations taking account of sward characteristics and grazing management, which can potentially affect intake compared to indoor feeding. Rotational and continuously stocked grazing systems are considered separately. Specific effects of grazing management on intake were quantified from an extensive literature review, including the effect of daily herbage allowance and pre‐grazing herbage mass in rotational grazing systems, sward surface height in continuously stocked grazing systems, and daily time at pasture in both grazing systems. The model, based on iterative procedures, estimates many interactions between cows, supplements, sward characteristics and grazing management. The sensitivity of the prediction of herbage intake to sward and management characteristics, as well as the robustness of the simulations and an external validation of the GrazeIn model with an independent data set, is presented in a third paper.     

Frequency of defoliation of native and naturalized species of the Flooding Pampas (Argentina)

This study describes the effect of herbage mass of species on defoliation patterns in a multispecies temperate–subhumid natural grassland of the Flooding Pampas at low, medium and high sward herbage mass [761  ±  8·7, 989  ±  137 and 1591  ±  44·0 kg green dry matter (DM) ha^–1 respectively] maintained by continuous grazing of cattle at variable stocking rates. Four native and naturalized warm-season species were studied: the grasses Cynodon dactylon , Leersia hexandra and Paspalum dilatatum and the legume Lotus tenuis . Sward state of the four species was described in terms of sward surface height, herbage biomass and the number of individual grass tillers and legume stems per unit area. The defoliation pattern was monitored twice-weekly during an experimental period of 3 weeks on permanently marked individual plants. Proportions of individual plants defoliated and the ranking of species selected differed among sward treatments and species. The results showed that the density of the species was a major factor determining the pattern of species defoliation and suggest that, in continuously grazed mixtures, the pattern of defoliation might be more dependent on the abundance of a species in the plant community than on species preferences. The results show the importance of considering this structural characteristic of swards in understanding the process of species selection by large herbivores in multispecies vegetation.     

Pattern of herbage intake rate and bite dimensions of rotationally grazed dairy cows as sward height declines

To allow improved prediction of daily herbage intake of dairy cows in rotational grazing systems, intake behaviour was assessed throughout the day in 24‐h paddocks. Herbage intake in 16 lactating Holstein–Friesian cows was assessed using the short‐term (1‐h) weight gain method at four predetermined natural meal times throughout the day (early morning, T1; late morning, T2; mid‐afternoon, T3; and early evening, T4). The study comprised two 4‐day experiments, each with a cross‐over design of four blocks. In both experiments, cows grazed a 24‐h paddock daily, and the effect of the immediately previous grazing experience on intake behaviour was investigated throughout the day, taking account of daily fluctuations in the short‐term physiological condition of the cows. Experiment 1 was carried out to investigate overall grazing behaviour during meals as a sward is progressively depleted during the day, with intake being assessed within the paddock and, hence, on a depleted sward. Experiment 2 similarly investigated the effect of sward depletion and physiological condition throughout the day on intake, but cows were removed to fresh, undefoliated swards during intake measurement periods; thus, intake rate was not influenced by differences in sward condition. Intake behaviour from both experiments was compared to establish the effect on herbage intake of changes in sward state and non‐sward factors. In Experiment 1, sward surface height, available herbage mass, proportion of leaf and green leaf mass declined as the day progressed. Bite mass declined with sward depletion, and mean intake rate was 1·64 kg dry matter (DM) h^–1, which was significantly lower at T3 (P < 0·01) than during other meals. In Experiment 2, plot sward conditions did not change throughout the day, and intake behaviour also remained constant, with a mean intake rate of 2·11 kg DM h^–1. Mean bite depth as a proportion of pregrazing extended tiller height was constant throughout the day (mean 0·32). The results show that, although cows grazed throughout the day on progressively depleted swards, indicative of rotationally grazed paddocks (Experiment 1), bite mass declined linearly and intake behaviour was variable. However, where intake was assessed on high‐quality, undefoliated swards (Experiment 2), intake behaviour was similar regardless of the time of day and the immediately previous experience. There was some indication of an interaction between the effects of the sward and the physiological condition of the animal on herbage intake.     

A model describing the utilization of herbage by cattle in a rotational grazing system

A model that describes the utilization of herbage by cattle in a rotational grazing system is presented. The model considers swards as being structured into two phases: a long phase associated with old dung patches, and a short phase. Both phases are treated as consisting of a series of horizontal strata corresponding in depth to a bite depth. The sward is divided into feeding stations consisting of either the long or the short phase. In each, only the surface stratum is available for grazing at each time step. At any time step, the individuals of a herd of cattle, distributed at random, encounter the entire range of strata. The rate of intake of each member of the herd depends on the intake properties of the stratum that it has encountered. The number of cattle that encounter each stratum type is variable so that the mean intake per member of the herd is the weighted mean. The core feature of the model is the simulation of the change over time in the frequency distribution of exposed stratum types and the distribution of grazing across this range of strata. The members of the herd are assumed to select a feeding station based on preference for leafiness of the encountered stata and the phase. The decision to graze or not is based on the comparison between the current vs. the previous feeding station. Model parameter values were based on published data. The proportion of leaf and bulk density of a phase or the strata were determined from an analysis of a sample of sward profiles. Using bite dimension, bite weight, biting rate, search time, feeding station area and selective behaviour, it was possible to simulate sward depletion that is very similar to the observed data from grazed paddocks in experiments in south‐east Ireland. The model of herbage utilization adequately described the changes in intake and sward structure during grazing and it was concluded that it was suitable for use as part of a simulation of a grazing system.     

Allowance-intake relations of cattle grazing vegetative tall fescue†

Herbage allowance is one of the important pasture factors in the determination of intake by grazing livestock. Ingestive behaviour of 12 adult Angus cows (Bos taurus) was measured over a range of allowances (0·25 to 0·72 kg dry matter (DM) per 100 kg live weight (LW) for a 1-h period) of vegetative tall fescue (Festuca arundinacea Schreb.). A balanced change-over design was used to estimate direct, residual and permanent effects of herbage allowance on rate of DM intake, rate of biting and herbage DM intake per bite. In Experiment 1, herbage DM intake per meal increased linearly from 0·68 to 1·72 kg (100 kg LW)^?1 as DM allowance increased from 0·25 to 0·72 kg (100 kg LW)^?1 h^?1. Cows grazed at ·30 kg (100 kg LW)^?1 h^?1 and stopped grazing when the sward was reduced to a height about 10 to 12 cm above the soil surface, approximately defined by the tops of pseudostems. In Experiment 2, herbage DM intake rates of 0·29, 0·47 and 0·42 kg (100 kg LW)^?1 h^?1 were recorded as cows grazed allowances of 0·43, 0·70 and 0·90 kg (100 kg LW)^?1 h^?1 for most of the 1-h grazing period. Limiting herbage DM allowances in Experiment 2 were associated with small reductions in rate of biting and herbage DM intake per bite as allowance declined. Sward DM density (>5 cm) was an important variable in the determination of herbage DM intake rates at lower herbage allowances.     

Maintaining post‐grazing sward height of Panicum maximum (cv. Mombaça) at 50 cm led to higher animal performance compared with post‐grazing height of 30 cm

In swards of tall, tufted, tropical grasses like Mombaça guinea grass (Panicum maximum (Jacq.)), post‐grazing heights promote changes in sward structure, which influence animal performance. This study evaluated changes in sward structure in response to grazing management at two post‐grazing heights (30 and 50 cm), associated with 90 cm pre‐grazing height. Each treatment was allocated to experimental unit (1.5 ha) in three replicated blocks. Pastures were evaluated pre‐ and post‐grazing to estimate herbage mass, percentages of leaf (LP), stem (SP) and dead material (DP), and nutritive value (VN). Stocking rate was adjusted twice a week “using the put‐and‐take approach,” and animals were weighed every 28 days. Forage accumulation rate was greater for pasture managed at 30 cm (64.8 vs. 55.1 kg ha^?1 day^?1) than 50 cm. A greater number of days were required after grazing 30‐cm residual pasture to achieve the pre‐grazing target height, resulting in 25% decrease in number of grazing cycles compared with pastures managed at 50 cm. Regardless of post‐grazing height targets, SP and DP decreased, whereas LP and NV increased from soil level to top of canopy. Stocking rate was greater in pastures managed to 30 cm than in those managed to 50 cm residual height (4.7 vs. 3.4 AU ha^?1). However, average daily gain was greater for 50 cm than for 30 cm post‐grazing height (795 vs. 590 g steer^?1 day^?1), resulting in a greater animal production per area (917 vs. 794 kg/ha of live weight). Thus, Mombaça guinea grass subject to intermittent grazing should be managed at 50 cm residual height.