Herbage losses from tiller pulling in a continuously grazed perennial ryegrass sward

Tiller pulling was studied in a perennial ryegrass sward that was continuously grazed by cattle. The treatments included severe (sward height after grazing 25 mm), medium (50 mm) or lenient (75 mm) grazing from turn-out in April to 1 June, followed by grazing to 50 mm in the remainder of the season. Tiller pulling was confined to the midsummer-autumn period of the grazing season. The losses were most severe in swards that had been leniently grazed to a mean height of 75 mm during the spring and least severe in swards grazed to a height of 25 mm. The lenient grazing treatment allowed both the true stem development and aerial tillering whereas in the more tightly grazed swards true stem development was significantly less and aerial tillering was virtually absent. The pulled organic matter in the lenient treatment was equivalent to about 5·3% of the total harvested yield. In the severely grazed swards, pulling losses were equivalent to about 1·7% of the total harvested yield. A high rate of turnover of the pulled herbage was found in all the treatments with between 69 and 78% of the freshly pulled herbage disappearing within a week of being pulled. Tiller pulling was found to have no effect on either the subsequent autumn-winter tiller density or yields of cuts taken in the following year.     

Herbage intake and milk production by grazing dairy cows 2. The effects of level of winter feeding and daily herbage allowance

Two experiments are described in which two levels of winter feeding and three levels of herbage allowance during the grazing season were imposed upon March/April calving British Friesian dairy cows. The winter treatments resulted in differences in live weight and milk yield at turnout of 35 and 53 kg and 3·4 and 3·2 kg d^-1 for the two trials. Subsequently, when grazed at generous herbage allowances, the cows were able to compensate for much of this difference but when herbage was restricted the milk yield differences were accentuated. Groups of cows from each winter treatment were offered 25, 50 or 75 (Experiment 1) and 30, 50 or 70 (Experiment 2) g herbage DM per kg LW daily during the grazing season. Daily herbage intakes on the three allowances in each trial were 14·1, 13·3, 10·7 and 12·5, 12·1, 11·5 kg OM and milk yields were 16·0, 15·3, 12·5 and 15·2, 14·3, 11·8 kg SCM respectively. Both intake and milk production were depressed once the cows were forced to consume more than 50% of herbage on offer or to graze the sward down to a mean height of less than 8–10 cm. Grazing behaviour observations indicated that under rotational managements the cows did not compensate for restrictions in available herbage by grazing longer. Highest levels of milk production per unit area were observed in both trials when production per cow was depressed by 20–25%.     

Effect of grazing severity on perennial ryegrass herbage production and sward structural characteristics throughout an entire grazing season

The objective of this study, which was part of a larger grazing‐systems experiment, was to investigate the cumulative impact of three levels of grazing intensity on sward production, utilization and structural characteristics. Pastures were grazed by rotational stocking with Holstein–Friesian dairy cows from 10 February to 18 November 2009. Target post‐grazing heights were 4·5 to 5 cm (high; H), 4 to 4·5 cm (intermediate; I) and 3·5 to 4 cm (low; L). Detailed sward measurement were undertaken on 0·08 of each farmlet area. There were no significant treatment differences in herbage accumulated or in herbage harvested [mean 11·3 and 11·2 t dry matter (DM) ha^?1 respectively]. Above the 3·5 cm horizon, H, I and L swards had 0·56, 0·62 and 0·67 of DM as leaf and 0·30, 0·23 and 0·21 of DM as stem respectively. As grazing severity increased, tiller density of grass species other than perennial ryegrass (PRG) decreased (from 3,350 to 2,780 and to 1771 tillers m^?2 for H, I and L paddocks respectively) and the rejected area decreased (from 0·27 to 0·20 and to 0·10 for H, I and L paddocks respectively). These results indicate the importance of grazing management practice on sward structure and quality and endorse the concept of increased grazing severity as a strategy to maintain high‐quality grass throughout the grazing season. The findings are presented in the context of the need for intensive dairy production systems to provide greater quantities of high‐quality pasture over an extended grazing season, in response to policy changes with the abolition of EU milk quotas.     

Lactation performance of spring-calving dairy cows grazing mixed perennial ryegrass/white clover swards of differing composition and height

An experiment was designed to examine the changes in clover content of three mixed perennial ryegrass/white cover swards of differing initial clover contents subjected to different grazing height management regimes and their effect on lactation performance of 48 Friesian dairy cows and heifers. Two paddocks were established for each treatment and grazed on alternate days. Treatments T17 and Tl3 consisted of swards with initial clover contents of 0·17 and 0·13 of the dry matter (DM) mass, respectively, grazed to maintain compressed sward heights of 6 cm throughout the season. A third treatment, SI5, consisted of a sward with an initial clover content of 0·15 grazed to maintain a compressed sward height of 4·5 cm for the first 78 days of the grazing season (period 1). Throughout period 1, half the animals on each treatment each received 4 kg of a concentrate supplement daily, while the others remained unsupplemented. From days 79 to 90, the cattle on treatment S15 grazed a similar sward, while the compressed sward height of the S15 paddocks was allowed to increase to 6 cm before re introduction of the animals. The three swards were then grazed for a further 47 days (period 2) before the animals were housed and milk yield recorded for a further 63 days (period 3). While sward T17 showed little change in clover content over the first 29 days of grazing, remaining at just below 0·18 of DM mass, swards T13 and S15 showed a marked decline in clover content to 0·05 and 0·07 of DM mass respectively. However, by the end of period 1 the clover content of all three swards had increased markedly (0·25, 0·15 and 0·15 of DM mass respectively). By the end of period 2, clover proportions were slightly higher than initial values (0·19. 0·15 and 0·15 of DM mass for treatments T17, T13 and S15, respectively). Owing to the relatively small differences in clover content of swards TI7 and T13, there were no significant effects of these two treatments on milk yield or composition in any period. Supplementation had no effect on milk composition and had little effect on milk yield, except when sward height was maintained at 4·5 cm. There was no carryover effect of supplementation on milk yield or composition in periods 2 or 3.     

Does post‐grazing sward height influence sward characteristics,seasonal herbage dry‐matter production and herbage quality?

Extending the grazing season through the production and utilization of high‐quality forage is a key objective in grassland‐based dairy production systems. Grazing swards to a low post‐grazing sward height (PGSH) is a strategy for improving grass utilization. A grazing experiment conducted in Ireland investigated immediate and subsequent effects of PGSH on sward production, utilization and structural characteristics. Swards were grazed to 2·7 cm (severe; S1) or 3·5 cm (moderate; M1) from 10 February to 18 April 2010 (Period 1; P1). From 19 April, each P1 paddock was halved and grazed to either 3·8 cm (S2) or 4·8 cm (M2), until 30 October (Period 2; P2). The first grazing rotation was +7 d on S1 swards compared with M1 swards (45 d), due to greater herbage utilization (+0·22). Herbage production during P1 was not affected by PGSH but a severe PGSH during this period reduced subsequent herbage production: 13·9 (S1) vs. 15·5 t dry matter (DM) ha^?1 (M1) by the end of the study. Leaf proportion was increased (+0·10) on S2 swards compared with M2 swards, but M2 swards produced 1·2 t DM ha^?1 more herbage during P2. Despite the relatively lower level of sward utilization obtained from moderate grazing in P1 (3·5 cm) and P2 (4·5–5·0 cm), such levels of PGSH increased DM production while maintaining sward quality, compared with severe grazing (2·7 cm in P1 and 3·5–4·0 cm in P2).     

The manipulation of grass swards for summer-calving dairy cows

Two experiments examined the effects of different defoliation treatments in spring on sward morphology and animal performance in mid-season and late season. Three treatments were applied in both experiments: Control (C), sward grazed by cows in spring to 6–8 cm grass height. Grazed Aftermath (GA). sward grazed by cows in spring to 3–4cm and allowed to regrow before being grazed by summer-calving cows, Silage Aftermath (SA), sward not grazed in spring, but a primary cut taken and the sward allowed to regrow before being grazed by summer-calving cows. The aim of treatment GA was to produce a sward with a high tiller density and high intake characteristics to meet the forage intake requirements of continuously grazed summer-calving cows, without resorting to offering forage buffers. Experiment 1 was conducted in 1989 on a sandy loam soil and Experiment 2 in 1990 on a heavy loam soil. In both experiments the GA treatment led to high live tiller density and live: dead tiller ratios compared with the C and SA treatments. Differences in sward morphology were also detected by applying double normal distribution analyses to measurements of grass height. The GA treatment also increased sward herbage mass and, to a limited extent, herbage metabolizable energy and crude protein contents. The results from Experiment 1 suggested that these sward effects lead to increased herbage dry-matter intake (as estimated by the n-alkane technique) and milk yield in cows grazing the GA sward. However, in Experiment 2, where conditions for grass growth in mid-season were more favourable than in Experiment 1, the differences in sward morphology produced in spring were quickly lost in June and July. There were therefore no differences in herbage intake or milk yield in the second experiment. Herbage intakes (kgDMd^?1± s.e.d) estimated in July for cows on treatments C, GA and SA were 11·0, 13·4, 10·1 ± 2·16 for Experiment 1 and 10·7, 11·1, 11·2 ± 2·32 for Experiment 2. Average milk yield (kgd^?1± s.e.d.) for cows on treatments C, GA and SA were 26·1, 28·0, 25·6 ± 0·31 (Experiment 1) and 28·5, 27·3, 28·4 + 0·58 (Experiment 2). The results suggested that acceptable milk yields can be obtained from grazing summer-calving cows, without offering forage buffers, by applying high stocking rates (low grass heights) in spring. However, the benefits of this manipulation could be lost by lax grazing in mid-season.     

Spring and autumn animal treading effects on pre‐grazing herbage mass and tiller density on two contrasting pasture types in Ireland

A perennial ryegrass (Lolium perenne L.)‐dominated sward on a well‐drained soil (Experiment 1) and a creeping bent (Agrostis stolonifera L.)‐dominated sward on a poorly drained soil (Experiment 2) were subjected to four treading treatments: control (C, no damage), light damage (L), moderate damage (M) or severe damage (S) to quantify the effects on herbage dry‐matter (DM) production and tiller density. In Experiment 1, treading damage was imposed in spring. In Experiment 2, one‐third of the site was damaged in autumn, one‐third in spring and one‐third in both spring and autumn. Both sites were rotationally grazed after treading treatments. Pre‐grazing herbage mass was measured eight times in Experiment 1 and seven times in Experiment 2 on each plot, and tiller density was assessed four times in each experiment. In Experiment 1, pre‐grazing herbage mass was reduced by 30% in S plots at the first harvest after damage, but cumulative pre‐grazing herbage DM production was not different between treatments (12·7 t DM ha^?1). In Experiment 2, annual cumulative pre‐grazing herbage mass was reduced by between 14 and 49%, depending on intensity of treading damage event and season when damage occurred. Tiller density was not affected by treatment in either experiment. A perennial ryegrass‐dominated sward on a well‐drained soil was resilient to heavy treading damage. A creeping bent‐dominated sward on poorly drained soil requires a more careful grazing management approach to avoid major losses in cumulative pre‐grazing herbage mass production during wet weather grazing events.     

The effect of sward height and its direction of change on the herbage intake, diet selection and performance of weaned lambs grazing ryegrass swards

Abstract Four sward height treatments were imposed by continuous variable stocking using at least ten Suffolk × Greyface lambs per plot from late July to late August: constant 3·5 cm. constant 6·0 cm. 3·5cm increasing to 6·0 cm and 6·0 cm decreasing to 3·5 cm. The treatments were established on two swards given fertilizer N applications over the season of 97 and 160 kg N ha^?1 respectively. Animal density was greater on the high fertilizer treatment, at the lower sward height and especially on the decreasing height treatments. Liveweight change of lambs was higher (P <0·001) on the 6-cm than on the 3·5-cm treatments (+159 vs-13g d^?1 and was also higher (P <0·001) on the increasing than on the decreasing sward height treatments (+92 vs-26 g d^?1). Herbage organic matter intake (OMI), measured on two occasions in the experiment, was greater (P <0·001) on the 6·0 cm than on the 3·5-cm sward heights whereas values for the increasing sward height treatments were much greater than those for the decreasing sward height treatments. There was little difference in the organic matter digestibility of the diet between treatments. Diets were composed largely of lamina, although there was more pseudostem and dead herbage in the diets of Iambs grazing the decreasing than the increasing sward height treatments at the end of the experiment. Bite mass was closely related to OMI but the treatment and period differences were relatively greater than for OMI. Bite mass was more closely related to the depth of the lamina layer (sward height-pseudostem height) than it was to sward height. There was evidence that pseudostem acted as a barrier to defoliation on these short swards and also that the proportion of youngest leaf in the diet was positively related to sward height and to increases in sward height. Sward height and especially the direction of change in sward height, together with associated stock density, were potent influences on lamb growth rate. This was a consequence of differences in herbage intake, which was strongly influenced by bite mass.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

Effect of replacing time available for grazing with time available for eating maize silage and soyabean meal on milk yield and feeding behaviour in dairy cows

Low rates of herbage dry matter (DM) intake impose limits on total daily DM intake in grazing dairy cows. The objective of this study was to increase total daily DM intake and milk production by restricting daily time available for grazing (TAG) and replacing it with time available for eating a maize silage/soyabean meal (TAMS) diet indoors. The treatments (TAG + TAMS) were 20 + 0, 19 + 1, 10 + 10 and 5 + 15 h. Measurements were made of milk production, intake and feeding behaviour. The interactions of TAG + TAMS treatments with sward height (SH) and concentrate level (CL) were also examined. Two experiments, each lasting 42 days, were carried out in spring ( Experiment 1 ) and autumn ( Experiment 2 ) using forty‐eight and twenty‐four Holstein‐Friesian cows respectively. Treatments were arranged in a factorial design with TAG + TAMS treatments, SH ( Experiment 1 only) and CL as the independent variables and a TAG + TAMS of 20 h. Reducing TAG and increasing TAMS significantly reduced estimated herbage DM intake and significantly increased maize silage/soyabean meal intake in both experiments, but there were no significant main effects of TAG + TAMS treatments on milk yield (mean, 27·4 and 25·5 kg d^?1 for Experiments 1 and 2 respectively), and yield of milk constituents. Increasing SH ( Experiment 1 ) and CL ( Experiments 1 and 2 ) significantly increased milk yield. In Experiment 1 , there was a significant interaction between TAG + TAMS treatments and SH with the taller sward height of 8–10 cm and the 20 + 0 treatment having the highest milk yield (29·7 kg d^?1) and the 5 + 15 treatment the lowest (27·2 kg d^?1), whereas at the lower sward height of 4–6 cm, milk yield was lowest on the 20 + 0 treatment (25·5 kg d^?1) with the other three treatments being higher (mean, 26·9 kg d^?1). Replacing TAG with TAMS significantly increased liveweight gain in Experiment 1 but not in Experiment 2 . Estimated rates of intake of herbage were lower in the autumn experiment ( Experiment 2 , 9·6 g DM min ^?1) than in the spring experiment ( Experiment 1 , 29·4 g DM min ^?1) but rates of intake of maize silage were higher in the autumn (112·4 g DM min^?1) than in the spring (72·5 g DM min^?1). In conclusion, in spring the response to replacing TAG with TAMS was dependent on sward conditions with the highest milk fat plus protein yield being on the 20 + 0 treatment at the high sward height and on the 19 + 1 treatment at the low sward height. The high liveweight gain of the 5 + 15 treatment could be an important means of restoring body condition in grazing lactating cows. In autumn, intakes of herbage were low in spite of its high estimated nutritive value with all treatments having a similar level of performance.     

The performance of summer-calving cows grazing perennial ryegrass swards

The objectives of this experiment were to study the effects of different grazing managements in spring on herbage intake and performance of summer-calving dairy cows and to examine the effects of regrowth in early June on herbage intake and cow performance. Four spring-grazing treatments were applied to predominantly perennial ryegrass swards: Control (C), sward grazed by cows to 6–8 cm sward surface height (SSH); CG16, sward grazed by cows to 3–4 cm SSH in May and allowed to regrow to a target SSH of 16cm in early June; CG8, sward grazed by cows to 3–4 cm SSH in May and allowed to regrow to 8cm in early June; and SG8, sward grazed by sheep to 2–3 cm SSH in May and allowed to regrow to 8 cm in early June, All swards were continuously stocked by summer-calving (May and July) primiparous and multiparous cows from 16 June to 7 September, to a target SSH of 8–10cm. Spring treatments bad marked effects on herbage intakes and milk production. Estimated in July by n alkane analysis, the mean herbage intake ± s.e.d. of cows on each treatment were 1·8, 1·4, 1·4 and 3·0 ± 0·31 kg dry matter (DM) 100 kg live weight (LW)^?1 d^?1 (P < 0·01) for treatments C, CG16, CG8 and SG8 respectively. Measured in August, intakes were 1·8, 20, 2·1 and 2·4 ± O·33kg DM 100kg LW^?1 d^?1 respectively. Severe spring grazing led to increased milk yield and reduced milk fat content from summer-calving cows fed 5·2 kg d^?1 of a proprietary concentrate. Average milk yields for the eleven experimental cows on each treatment were 24·3, 23·4, 26·2 and 29·0 ± 1·20 kgd^?1 (P < 0·01) for C, CG16, CG8 and SG8, and average milk fat contents were 45·4. 42·4, 43·9 and 40·9 ± 1·02gkg^?1 (P<0·05) respectively. The results suggest that severe grazing of swards in early season could improve herbage intake and milk yield of summer-calving cows in mid- and late season. The most favourable spring treatment in this respect was severe grazing by sheep. However, this advantage could be negated in midseason by lax grazing at that time.     

Some effects of applying undiluted silage effluent to grassland

Two small-plot experiments were carried out to assess the influence on herbage dry matter (DM) production, chemical composition and soil fertility status of applying undiluted silage effluent at a range of application rates and intervals after a silage cut. In the first experiment, in 1990, silage effluent was applied at 25, 50, 100, 150 and 200 m³ ha^?1 1, 8, 15 and 22 d after a silage cut in August. In the second experiment, in 1991, silage effluent was applied at 7, 14, 21, 28, 35, 42, 49, 75, 100, 125 and 150 m³ ha^?1 1, 4, 8, 15, 22 and 29 d after silage cuts were taken from different sites in May, July and August. An untreated control and an Inorganic fertilizer treatment were incorporated in both experiments. The immediate effects of the treatments on herbage yield, chemical composition and soil nutrient status were assessed 6–8 weeks after the initial application; residual effects on herbage and soil fertility were measured at a subsequent harvest. Compared with the untreated control, herbage yield increases were obtained with increasing rates of effluent application. Although there was evidence that higher yields could be obtained from earlier applications, up to 50 m³ ha^?1 of effluent could be applied up to 15 d after taking a silage cut with little damage to the sward. Delaying the timing of application, and increasing the application rate, increased the proportion of the sward which was damaged; this reached a maximum of 0·84 when the highest application rates were applied 29 d after a silage cut. The increase in the proportion of dead herbage in the sward, associated with increasing rate of effluent application, reduced the quality of the herbage harvested in Experiment I. In Experiment 2 the N, P and, in particular, the K content of the herbage increased with increasing rate of effluent application, whereas the effect on Mg content was variable with contents generally being less than 2·0 g kg^?1 DM. Apparent recovery of nutrients applied in the effluent was both low and variable ranging from 0·58 to ?0·03 for N, 0·10 to ?0·005 for P, 0·34 to ?0·02 for K and 0·21 to ?0·002 for Mg over both experiments. Effluent had little effect on soil pH, whereas P and, in particular, K contents increased with increasing rate of effluent application. There was evidence that effluent had a beneficial effect on both herbage yield and chemical composition at the residual cut, the extent depending upon rate and time of effluent application.     

Evaluating seasonal variation in mineral concentration of cool‐season pasture herbage

Pasture herbage is a major source of minerals for livestock in pasture‐based production systems. Herbage mineral concentrations vary throughout the growing season, whereas mineral supplementation to livestock is often constant. The study objectives were to analyse the seasonal variation in herbage mineral concentrations in tall fescue [Schedonorus phoenix (Scop.) Holub]‐based pasture with regard to beef cattle mineral requirements and to create a statistical model to predict variation in herbage mineral concentrations across the growing season. Pasture herbage was analysed from 12 grazing systems in Virginia to determine its mineral concentration from April to October of 2008–2012. The pasture herbage, grown without fertilization, contained adequate macronutrient concentrations to meet the requirements of dry beef cows through the growing season and the requirements of lactating beef cows in April. Phosphorus supplementation appeared to be unnecessary for dry beef cows given adequate concentrations in pasture herbage. A model using month of harvest, soil moisture and relative humidity explained 75% of the variation in an aggregated mineral factor. The 90% prediction intervals indicated that N, P, K, S and Cu concentrations could be predicted within 1·35, 0·08, 0·80 and 0·07% and 3·83 mg kg^?1 respectively. Prediction of herbage mineral concentrations could help to improve livestock health, reduce costs to producers and limit nutrient losses to the environment.     

Net production of herbage from a continuously stocked Lolium perenne-dominated sward in late autumn

Preliminary studies were carried out on the effect of stocking rate during late autumn on a continuously stocked Lolium perenne -dominated sward at an upland site in central Scotland. Measurements were made of L. perenne tiller population density on 29 September and 2 November and of L. perenne net production, mean sward height and total herbage mass in early and late October and early and late November. Stocking rates were 12 ewes per ha during October and 8 and 16 ewes per ha during November. Sward height and herbage mass declined with time and more rapidly at the higher stocking rate. L. perenne growth per tiller and per unit area was influenced by time but not by stocking rate and was closely related to the 5·5°C soil temperature at 10 cm depth. Tiller senescence was greatly reduced at the higher stocking rate and/or the consequentially lower sward height and herbage mass. Tiller net production was therefore sustained at a positive level on the higher stocked sward throughout November while on the lower stocked sward it fell below zero early in November.     

Effect of continuous stocking by sheep at four sward heights on herbage mass, herbage quality and tissue turnover on grass/clover and nitrogen-fertilized grass swards

The effects of continuous stocking by sheep at sward surface heights (SSH) of 3, 5, 7 and 9 cm in grass/clover (GC) and nitrogen-fertilized grass (GN) swards were examined in relation to herbage mass and quality, clover content, tiller density and rates of herbage production and senescence in two periods in each of three grazing seasons (1987-89). The GN swards received a total of 300 kg N ha^?1 each year in six equal dressings from March; GC swards received a single dressing of 50 kg N ha^?1 in March each year. Herbage mass measured from ground level increased linearly with SSH with overall mean herbage masses of 0·89, 1·38, 1·78 and 2·12 t OM ha^?1 (s.e.m.0·024, P < 0·001) at SSH of 3, 5, 7 and 9 cm respectively. GN and GC swards had mean herbage masses of 1·58 and 1·51 t OM ha^?1 (s.e.m. 0·051, NS) respectively. Mean N content of herbage on GN swards was greater than that on GC swards and declined with increasing SSH. Crude, fibre (CF) content of herbage was similar for both sward types and increased with increasing SSH. Clover content of GC swards remained low throughout the experiment, ranging from 0·002 to 0·074 of herbage mass. However, from tissue turnover rates it was estimated that its contribution to herbage production was in the range of 0·049–0·219 of net herbage growth. Total growth increased with increasing SSH in both sward types, with maximum growth rates in GN swards of 143 and 130 kg DM ha^?1 d^?1 and in GC swards of 88·2 and 85·4 kg DM ha^?1 d^?1 in Periods 1 (up to early July) and 2 (after July) respectively. Senescence rates ranged between 13·3 and 50·1 kg DM ha^?1 d^?1 and tended to be higher in Period 2 than in Period 1. Net production increased with increasing SSH in Period 1, while in Period 2 net production declined at SSH above 6·5 cm. The increased net herbage production in taller swards was not associated with greater utilized metabolizable energy production at sward heights above 5 cm.     

Performance of cultivated perennial grass mixtures under different grazing intensities in the alpine region of the Qinghai-Tibetan Plateau

In 1998, seeds of four cultivated native perennial grasses, Bromus inermis (B), Clinelymus nutans (C), Elymus nutans (E) and Agropyron cristatum (A), were sown as mixtures with different seed rates in three combinations (B + C, B + E + A and B + E + C + A) in a field study in the north‐eastern part of the Qinghai‐Tibetan Plateau, China. A grazing experiment was conducted in 2000 to investigate the performance of these grass mixtures at leaf, plant and sward scales under different grazing intensities (GI). Four GIs, expressed as the proportion of herbage consumed by Tibetan lambs in relation to the available biomass (IP), were applied in the summer: no grazing, and 0·30, 0·50 and 0·70 of IP. Tiller numbers of the grasses increased and leaf photosynthetic rate, sward leaf area index (LAI) and herbage mass declined with increase in GI. No effect of GI on specific leaf area was observed. Interactions between GI and grass mixtures on the dynamics of species composition, swards’ LAI and herbage mass were found. Large fluctuations in species composition with different GIs showed the poor species compatibility and sward persistence of these grass mixtures under high GI by sheep.     

The effects of a flexible grazing management strategy and leader/follower grazing on the milk production of grazing dairy cows and on sward characteristics

An experiment was carried out during 1984 to examine the effects of three alternative grazing strategies for January/February calved British Friesian dairy eows on sward and animal production. Cows were rotationally grazed across 1 d paddocks without concentrate supplementation from 30 April to 1 October. A flexible grazing (EG) treatment involved manipulating residual herbage height, as assessed by a rising-plate sward stick, with cows initially grazing to 80 mm, reducing to 60 mm when milk yield declined below 20 kg d^?1 and finally to 50 mm when milk yield declined below 15 kgd^?1. Control (C) cows grazed to a constant residual herbage height of 60 mm throughout the season (a 60-inm rising-plate sward stick height is equivalent to a sward surface height of approximately 80 mm). On a further treatment a leader/follower (LF) approach was used, with cows paired for calving date and parity and within pairs allocated to either a high-(leader) or a low-yielding (follower) group, according to milk yield at turnout, with the leader group grazing 1 d ahead of the follower group. Overall stocking rates on C and LE treatments were identical but herbage allowances differed as a result of treatment effects. Animal performance data for the FG, C and LE treatments, respectively, were: milk yield (kg d^?1) 14·5, 14·7 and 16·0 (s.e. 0·59); milk fat yield (g d^?1) 577, 571 and 637 (s.e. 29); milk protein yield (g d^?1) 528, 527 and 576 (s.e. 19); and liveweight gain (kg d^?1) 0·09, 0·20 and 0·14 (s.e. 0·04). Overall, there was no benefit in animal production following lax grazing in spring even with high-yielding cows, and this approach resulted in the accumulation of stem and senescent material in the sward in mid-season. However, preferential treatment of high-yielding cows by grazing as a leader group in a leader-follower system resulted in higher milk production, particularly in late season, with an overall improvement in milk yield for the LF treatment of 9% relative to treatment C.     

Response of herbage regrowth and water-soluble carbohydrate concentration of ryegrass species to defoliation practices when grown in a Mediterranean environment

Three experiments were conducted to determine the association between leaf number per tiller at defoliation, water‐soluble carbohydrate (WSC) concentration and herbage mass of juvenile ryegrass plants when grown in a Mediterranean environment. Seedlings of ryegrass were grown in nursery pots arranged side‐by‐side and located outside in the open‐air to simulate a mini‐sward in Experiments 1 and 2, and a mixture of annual ryegrass and subterranean clover (Trifolium subterraneum L.) was grown in a small plot field study in Experiment 3. Swards were defoliated mechanically with the onset of defoliation commencing within 28 d of germination. Frequency of defoliation ranged from one to nine leaves per tiller, whilst defoliation height ranged from 30 mm of pseudostem height that removed all leaf laminae in Experiment 1, to 50 mm of pseudostem height with some leaf laminae remaining post‐defoliation in Experiments 2 and 3. A positive relationship between herbage mass of ryegrass, WSC concentration and leaf number per tiller at defoliation was demonstrated in all experiments. In Experiment 1, the herbage mass of leaf, pseudostem and roots of tillers defoliated at one leaf per tiller was reduced to 0·10, 0·09 and 0·06 of those tillers defoliated less frequently at six leaves per tiller. However, the reduction in herbage mass from frequent defoliation was less severe in Experiment 2 and coincided with a 0·20 reduction in WSC concentration of pseudostem compared with 0·80 measured during Experiment 1. In Experiment 3, the highest harvested herbage mass of ryegrass occurred when defoliation was nine leaves per tiller. Although the harvested herbage from this sward contained senescent herbage, the in vitro dry‐matter digestibility of the harvested herbage did not differ significantly compared with the remaining treatments that had been defoliated more frequently. Leaf numbers of newly germinated ryegrass tillers in a Mediterranean environment were positively associated with WSC concentration of pseudostem and herbage mass. A minimum period of two to three leaf appearances was required to restore WSC concentrations to levels measured prior to defoliation thereby avoiding a significant reduction in herbage mass. However, maximum herbage mass of a mixed sward containing ryegrass and subterranean clover was achieved when defoliation was delayed to nine leaves per tiller.     

Concentrate supplementation, and the herbage intake and milk production of heifers grazing Cenchrus ciliaris

Friesian heifers grazing Cenchrus ciliaris cv. Biloela were supplemented with 0, 3 or 6 kg concentrates daily during weeks 10–34 (±1·7) of lactation during either the rainy or the dry season. The overall responses to concentrate were identical between seasons at 0·27 kg extra milk and solids corrected milk per kg. Supplementation increased total feed intake and modified the grazing behaviour of cows. For each kg concentrate organic matter eaten, herbage organic matter intake was reduced by 0·64 and 0·42 kg in the rainy and dry seasons respectively and the time spent grazing by 11 min. Higher intakes in the dry season were the result of an increased rate of biting and were reflected in liveweight change but not milk yield.     

The relationship between level of sward height reduction in a rotationally grazed sward and short-term intake rates of dairy cows

The objective of this study was to investigate the relationship between level of sward height reduction (SHR) and short-term intake of herbage by lactating dairy cows offered swards differing in initial bulk density (BD). Three experiments were carried out in which cows were presented with swards representing different levels of SHR (nominally described as ungrazed, low, moderate and high). Experiments 1 and 2 differed with respect to initial sward BD [ungrazed sward 1·7 vs. 2·5 kg dry matter (DM) m^?3 respectively]. Experiment 3 investigated the interaction between BD and SHR. In each experiment, sixteen Holstein/Friesian cows (fitted with excreta collection bags) grazed for a 1-h period in 200-m² plots that had been grazed on the previous day to predetermined sward surface heights (SSH) (= levels of SHR). Herbage intake rates were assessed from changes in live weight, with a correction for insensible weight loss (IWL). Biting rates were recorded from visual observation. High levels of SHR were associated with a significant reduction in SSH, herbage mass and leaf fraction, and a significant increase in sward BD, stem and dead fraction, and DM concentration. Herbage intake, expressed either as DM intake per bite or DM intake per hour, declined as level of SHR increased from low to high. The level of SHR generally had no effect on biting rate. Intake rates varied from 1·9 to 4·4 kg DM h^?1, whereas DM intake bite^?1 ranged from 0·5 to 1·3 g. Pooled regression analysis identified SSH (P < 0·001; r² = 0·94) as the principal determinant of DM intake bite^?1. The regression equation was not significantly improved by the addition of terms for leaf fraction, BD, or herbage mass. In Experiment 3, a significant interaction between level of SHR and sward BD was observed. It is concluded that the principal factor controlling intake (g DM bite^?1 or kg DM h^?1), as swards are progressively grazed down, is SSH, but at a high level of SHR, sward BD also influences intake bite^?1.