基于MaxEnt模型的陕西省菊芋种植潜在适宜区分析

利用陕西省29个菊芋空间分布数据和26个环境因子,综合陕西省边际土地分布,运用最大熵模型(MaxEnt)模拟历史时期及未来气候变化下陕西省菊芋潜在适宜区分布,并将结果叠加陕西省现有边际土地进行分析。结果表明：(1)MaxEnt模型模拟陕西省菊芋种植潜在适宜区结果精度较高,曲线下面积AUC为0.914。(2)基于历史时期环境数据,陕西省菊芋潜在分布区主要分布在榆林定边县、延安志丹县、安康汉滨区、商洛商州区和汉中西乡县,边际土地总适宜区面积约为75992km²,占陕西省总面积的36.9%,其中低、中、高适宜性面积分别约为63169、11617和1205km²,主要边际土地利用类型为中覆盖度草地。(3)影响菊芋生长的5个重要环境因子为土壤深度(142.4～155.4cm)、最暖月最高气温(29.4～31.6℃)、高程(210～593m)、坡向(70.7°～239.5°)和坡度(0～10.5°)。(4)未来4种气候情景模式下陕西省菊芋种植中、高适宜区面积总体明显增加,2021-2040年分别增加14.39%～46.46%和46.90%～95.39%,...     

基于生态约束-发展潜力的城镇建设用地开发适宜性分区

[目的]从生态和经济的角度划分城镇建设用地开发适宜性类型区,为江苏省射阳县土地利用空间布局提供参考。[方法]依据最小累积阻力值分析射阳县区域生态约束强度,划分强度等级;运用多因素多因子综合判别模型,从开发强度和空间距离角度构建评价模型,划分区域发展潜力等级;最终在生态约束等级和发展潜力等级的基础上,进行全域城镇建设用地开发适宜性分区。[结果](1)生态约束高区域与生态约束低的区域相差较大,总体上以生态保护为主。从空间分布上,不同等级强度生态约束力表现出了阶梯式的分布,由内向外辐射,强度逐步提高。(2)开发潜力与生态约束分级结果对比,矛盾突出,经济需求旺盛,生态保护用地形势严峻。(3)适宜保护区、较适宜保护区、较适宜建设区、适宜建设区4类,所占比率分别为11.04%,24.78%,34.59%,29.59%。[结论]开发适宜性分区表现出"核心—边缘"的城乡二元结构特征,同时合德镇及周围的适宜建设区较为集聚,外围适宜建设区逐渐减少。     

牡丹江市土地利用适宜性评价

[目的]评价牡丹江市退耕还林地区土地利用效益,对土地利用结构进行优化,为该区进一步土地开发利用及结构调整提供科学依据与支持。[方法]选取牡丹江市的坡度、高程、土壤类型、水资源系数、降水量和≥10℃积温6个因素作为评价因子,通过模型计算及GIS空间分析,得出研究区内宜农用地、宜林用地适宜性等级。[结果]处于适宜级以上的农业用地总面积约9 300km2,占研究区面积的22.9%,主要分布于坡度小于5°、海拔500m以下的区域;处于适宜级以上的林业用地总面积约28 000km2,占研究区总面积的69.0%,主要分布于海拔高、坡度大、多水土流失的山区丘陵地带。[结论]土地利用现状分析结果表明,在农林交错地带存在过度退耕、造成耕地浪费的现象。     

基于功能用地适宜性的农业科技园区规划方案评价

农业科技园区用地适宜性评价和规划方案评估是规划区用地功能配置的前提。该文以南京农业大学白马教学科研基地为例,应用GIS地形数据和RS遥感数据,在已有用地适宜性模型的基础上,将水生态安全因子融入农业科技园区建设用地适宜性评价指标体系,结合高程、坡度、噪声和土地利用等评价因子,构建功能用地适宜性空间定量模型,阐述了功能用地适宜性评价在规划方案评估中的应用与技术路线,分别对生活功能、公共服务、农业实践、展示功能和教学实践等功能用地适宜性进行定量评价,最后对6种不同规划方案综合用地适宜性进行总体评估,并充分结合空间布局中规划理性要求,确定方案6为最佳规划方案。分析和评价结果显示,结合水生态安全因子对功能用地适宜性进行评价,能够较好地对不同规划方案综合评估,从而选取最佳规划方案,最后为用地功能布局进行辅助决策,因此评价结果具有一定的科学性和实用性。该文将区位选择与农业科技园区规划中的功能布局进行关联,结果表明该文提出的方法是科学可行的,为农业科技园区规划方案评估探索了一种途径,也为城乡用地统筹规划管理提供了科学依据和决策支持手段。     

甘肃省冬油菜种植适宜性及影响因子评价

全球气候变暖背景下,随着气候变化、油菜栽培技术的进步和油菜新品种的育成推广,我国传统油菜生产格局和种植结构发生明显变化,甘肃省冬油菜的种植地带也随之向北扩展,种植的海拔高度抬升,种植面积和产量显著增加。研究甘肃省冬油菜种植适宜性,对改进甘肃省油菜生产布局,合理调整农作物种植结构具有重要意义。本研究获取多年田间试验数据,初步筛选9个影响冬油菜种植分布的潜在气候因子,采用基于DEM的小网格法建立9个影响冬油菜种植分布的潜在气象因子空间分布图层,在此基础上利用最大熵模型和GIS空间分布技术,构建冬油菜潜在分布模型,分析影响甘肃省冬油菜种植分布的主要生态因子及其特征,评价甘肃省冬油菜种植适宜性。研究结果表明:甘肃省冬油菜种植分布概率为0~0.89,按其分布概率可将甘肃省冬油菜种植区域划分为不适宜种植区域、次适宜种植区域、适宜种植区域和最适宜种植区域,可以种植冬油菜的区域占甘肃省总面积的60%,大于实际冬油菜种植面积,冬油菜在甘肃省种植具有很大的发展潜力。按照9个潜在气象因子对冬油菜种植分布贡献率的大小,确定影响甘肃省冬油菜种植分布的6个主导气象因子及其阈值为:负积温≥-800℃,最冷月最低温度≥-20℃,最冷月平均温度≥-15℃,年平均温度≥4℃,极端低温≥-28℃,50 mm≤生育期降水量≤200 mm。并以此指导甘肃省冬油菜种植布局,调整甘肃省农作物种植结构,引导农民优化种植模式,提高经济效益。     

五常市北部地区农村居民点用地适宜性评价与整治分区规划

农村居民点用地适宜性评价是实现农村土地合理布局、挖掘土地整理潜力的重要依据。以五常市北部地区为例,从自然、经济社会和生态三方面构建典型粮食产区农村居民点用地适宜性评价指标体系,结合地区内部特点量化指标因子;依据适宜性评价分级和区域模式差异,综合考虑人口城镇化影响实现居民点的整治分区规划。结果表明:北部12个乡镇中农村居民点用地适宜性区间为0~3.996 8,不存在最高适宜的居民点用地,域内以高适宜和中适宜用地为主,低适宜和不适宜的居民点用地相对比例之和5.02%,地区仍有部分居民点需重点整治;兴隆乡、营城子乡、民乐乡等部分地区应引导村内人口外迁,八家子乡、常堡乡、牛家镇等应积极迁村并点,兴隆乡部分地区、八家子乡和卫国乡实施内部改造,保留发展型均有分布。研究结果可为该地区居民点整理规划提供科学依据。     

影响鱼类分布的生态水文因子建模及分析

人类活动干扰下河流自然水势改变,导致鱼类栖息地环境变化。为准确分析影响鱼类分布的生态水文环境,该文以美国阿肯色河流域的国家级濒危小型鱼类:产漂流性卵鲤鱼(Arkansas River Shiner,Notropis girardi,ARS)为例,根据其繁殖期特有的水文条件需求,选取流域40个水文站点的日径流数据,分别计算历史时期(1950-1962年)和当前时期(1990-2010年)的3个繁殖期生态水文因子:高流量,较高流量历时及断流历时。每个生态水文因子分别结合其他25个环境因子,基于最大熵理论(MaxEnt)构建物种分布模型。运用经典受试者工作曲线(receiver operating characteristic,ROC)及Parolo等提出的模型转移前后预测结果的Spearman’s rho相关系数,评价模型的自适应性(model fit)和转移能力(transferability)。3个模型在两个时期的ROC曲线与横坐标围成的面积均大于0.95,模型转移前后预测结果的Spearman’s rho相关系数均大于0.6。研究分析表明,模型具有较好的自适应性及转移性。3个生态水文因子中,高流量对模型的贡献率最大,对该种鱼类在两个时期的潜在分布影响最显著。不同的历史时期,物种的潜在适生概率随同一生态水文因子的变化呈现出了不一致的变化趋势。3个模型均预测出ARS的潜在生境有相同的空间变化趋势。该研究结果能够为中国保护河流生态系统与濒危鱼类提供借鉴经验。     

基于适宜性与发展水平评价的村庄用地类型划分

科学划分村庄用地类型并提出差异化的整治策略,是编制村庄发展规划和建设用地减量规划的基础。该文以北京市密云区为例,基于土地利用、兴趣点和社会统计等数据,从自然条件和用地形态2方面构建村庄用地适宜性评价模型,从区位条件、基础设施、生态条件和资源产业4方面构建村庄发展状况评价模型,系统分析村庄用地适宜性与村庄发展状况及其二者的组合状态,进而划分村庄用地类型。结果表明:1)密云区村庄用地适宜性呈现明显的从西南到东北降低的分布趋势;2)村庄发展指数较高的村庄主要分布在西南区域的环密云城区一带,其次是中部围绕密云水库区域,北部区域村庄发展水平较低;3)基于二者的评价结果将全区村庄用地划分为9种组合,进而合并为4种村庄用地类型。该研究可为村庄用地的高效、有序整治利用提供依据。     

影响鱼类分布的生态水文因子建模及分析（英文）

人类活动干扰下河流自然水势改变,导致鱼类栖息地环境变化。为准确分析影响鱼类分布的生态水文环境,该文以美国阿肯色河流域的国家级濒危小型鱼类:产漂流性卵鲤鱼(Arkansas River Shiner,Notropis girardi,ARS)为例,根据其繁殖期特有的水文条件需求,选取流域40个水文站点的日径流数据,分别计算历史时期(1950-1962年)和当前时期(1990-2010年)的3个繁殖期生态水文因子:高流量,较高流量历时及断流历时。每个生态水文因子分别结合其他25个环境因子,基于最大熵理论(Max Ent)构建物种分布模型。运用经典受试者工作曲线(receiver operating characteristic,ROC)及Parolo等提出的模型转移前后预测结果的Spearman’s rho相关系数,评价模型的自适应性(model fit)和转移能力(transferability)。3个模型在两个时期的ROC曲线与横坐标围成的面积均大于0.95,模型转移前后预测结果的Spearman’s rho相关系数均大于0.6。研究分析表明,模型具有较好的自适应性及转移性。3个生态水文因子中,高流量对模型的贡献率最大,对该种鱼类在两个时期的潜在分布影响最显著。不同的历史时期,物种的潜在适生概率随同一生态水文因子的变化呈现出了不一致的变化趋势。3个模型均预测出ARS的潜在生境有相同的空间变化趋势。该研究结果能够为中国保护河流生态系统与濒危鱼类提供借鉴经验。     

基于马尔科夫模型的大凌河流域土地利用/覆被变化趋势研究

本文通过对大凌河流域2010年、2013年和2016年三期遥感影像进行分类,得到流域土地利用分布现状。对分类结果进行叠加分析可以获得大凌河流域2010~2013年的地类转移概率矩阵,再运用马尔科夫原理预测出2016年流域土地利用分布情况,将其与实际情况对比分析,证明马尔科夫模型适用于大凌河流域未来土地利用/覆被变化趋势的预测。进而本文分别对大凌河流域2019年、2022年和2025年的土地利用情况进行预测,发现未来近10年流域建设用地面积持续增长,变化率为38.96%;农林用地面积从2016年开始减少,总体下降了0.77%;水域面积稳定增长,到2022年之后趋于稳定值,变化率为17.21%;旱地及其他未利用地面积继续缩减,到2019年开始趋于稳定,整体下降了1.71%。研究结果明晰了大凌河流域农林用地和建设用地之间的矛盾,可以为流域水土资源管理有关部门提供科学依据。     

低氮和干旱胁迫对富士和秦冠生长及氮素利用的影响

【目的】以富士（Fuji）、 秦冠（Qinguan）嫁接在平邑甜茶（Malus hupehensis Rehd.）上的当年生盆栽苗为试验材料,采用砂培方法,研究了缺氮胁迫和干旱对富士和秦冠生长情况、 光合参数、 植株各部位氮磷钾含量及氮素利用效率的影响,分析比较了低氮干旱条件下富士和秦冠生长及氮素利用的差异,以期为果树生产高效肥水利用提供理论指导。【方法】试验共设四个处理: 正常氮正常水（ZZ）、 低氮正常水（DZ）、 正常氮干旱（ZG）、 低氮干旱（DG）。氮素和水分均设置两个水平,分别为正常氮（6 mmol/L NO-3-N）、 低氮（0.3 mmol/LNO-3-N）、 正常供水（保持盆中砂子相对含水量为饱和含水量的80%~85%）、 干旱处理（保持盆中砂子相对含水量为饱和含水量的60%~65%）。【结果】富士和秦冠的生物量（茎和叶）、 株高茎粗等生长指标以及光合速率、 气孔导度、 蒸腾速率均为正常氮正常水（ZZ）＞低氮正常水（DZ）＞正常氮干旱（ZG）＞低氮干旱（DG）,并且相对应处理下秦冠的以上指标均高于富士;正常供水下,缺氮处理使富士、 秦冠的根冠比比正常氮处理均有所增加,富士提高了2.05%,秦冠提高了22.40%。富士和秦冠的氮、 磷、 钾含量均表现出正常氮正常水（ZZ）＞低氮正常水（DZ）＞正常氮干旱（ZG）＞低氮干旱（DG）; 氮、 钾元素含量在植株各部位的分布顺序依次是叶＞根＞茎,磷元素则是根＞叶＞茎;光合氮素利用效率（PNUE）和氮素利用效率表现为秦冠处理之间差异极显著,富士处理之间差异不显著;秦冠的PNUE和NUE明显高于富士,在低氮正常水（DZ）处理下,秦冠氮肥利用率比富士高42.07%,在低氮干旱（DG）处理下高64.14%;低氮胁迫下富士和秦冠的NUE显著提高,并且秦冠提高的幅度高于富士。【结论】施用氮肥能够显著提高富士与秦冠的干物质量,同等水肥条件下,秦冠生长优于富士;水分亏缺会减少叶片对氮的吸收,干旱条件下适度增施氮肥,可提高果树的抗旱能力;低氮干旱胁迫下秦冠的生长指标、 光合指标及氮素利用效率指标均优于富士,表现出较强的抗低氮干旱胁迫的能力。     

Biodiversity and ecogeography of wild Lactuca spp. in some European countries

Staff members of the Department of Botany of Palacký University in Olomouc and Gene Bank Department – Workplace Olomouc, Research Institute of Crop Production in Prague, Czech Republic, conducted an expedition in seven European countries (Austria, Czech Republic, France, Germany, Italy, the Netherlands, Switzerland) in August/September 1999 to collect wild Lactuca spp. germplasm and study its geographic distribution, ecology and biodiversity. During the mission, more than 600 locations were visited resulting in the collection of 602 seed samples (accessions) of wild Lactuca species and 13 seed samples of related genera (Chondrilla and Mycelis). Lactuca serriola f. serriola, L. serriola f. integrifolia, L. saligna and L. viminea subsp. chondrilliflora were prevalent in southern Europe (Italy, France), however, only L. serriola was common in central and western Europe (Austria, Czech Republic, Germany, Netherlands, Switzerland). The greatest diversity of Lactuca species was found in France, where also the most seed samples (165) were collected. The most characteristic habitats with a high density of Lactuca spp. populations were observed along roads and highways, grassy ditches, ruderal communities, and dust-heaps. Natural infections by powdery mildew (Erysiphe cichoracearum) and downy mildew (Bremia lactucae) on some wild Lactuca spp. were observed. Recent observations concerning the geographic distribution, population structure, habitats, and natural occurrence of diseases of Lactuca spp. are discussed. This assemblage of genetic resources of Lactuca spp. can serve as the basis of future studies of species diversification, spatial population structure, plant microevolution, domestication processes, and genetic variability of host-parasite interactions.     

Pasture Vegetation Elemental Analysis by Laser-Induced Breakdown Spectroscopy

Laser-induced breakdown spectroscopy (LIBS) is a new technique for the analysis of plant material. This study investigates the application of LIBS to pasture-based plant samples. The LIBS measurements were obtained from pelletized pasture samples (100 samples) that had been also analyzed by inductively coupled plasma–optical emission spectroscopy (ICP-OES) following microwave digestion for calibration and comparison purposes. Comparisons for elements sodium (Na), potassium (K), magnesium (Mg), calcium (Ca), manganese (Mn), iron (Fe), copper (Cu), zinc (Zn), boron (B), phosphorus (P), and sulfur (S) showed that LIBS could be used for almost all the standard profile total elements with concentrations down to low mg/kg levels (observed error of Na: 0.024 percent, K: 0.18 percent, Mg: 0.016 percent, Ca: 0.073 percent, P: 0.017 percent, Mn: 31 mg/kg, Fe: 150 mg/kg, Zn: 6.6 mg/kg, and B: 1.1 mg/kg). Elemental analysis at less than mg/kg levels was not possible using LIBS. The elements S and Cu were particularly difficult to analyze with reliability using LIBS at the concentration levels found in the plant samples. Replacing microwave digestion and subsequent ICP analysis with a direct analysis of dried plant samples using LIBS has the potential to improve the productivity and reduce the cost of testing.     

Properties of anaerobically digested and composted municipal solid waste assessed by linking soil mesofauna dynamics and nitrogen modelling

We have studied the effect of anaerobically digested (ADMSW) and composted municipal solid waste (CMSW) on mineralization and foodweb dynamics to verify the hypothesis that ADMSW would immobilize N right after addition to soil in contrast to addition with CMSW. Another hypothesis was that the mesofauna (enchytraeids and microarthropods) would stimulate N release from the decomposer community. We measured excretion of -N and urea-N from the mesofauna and hypothesized that enchytraeids would release urea. ADMSW and CMSW were amended to pots with sandy loam and barley. The pots were divided into treatments with or without mesofauna. Mesofauna, plant N and biomass, soil N and ergosterol (fungal biomass) were measured over a 113-day period of four equidistant samplings. Soil respiration, N mineralization and N release by the mesofauna were modelled from concurrent studies. ADMSW- and CMSW-treated soils initially (<20 days) immobilized N. The amendments did not increase plant growth substantially, and this was probably due to N-limitation in the early stages of plant growth. Enchytraeid abundance was about three times higher in ADMSW- than CMSW-treated soils, indicating that ADMSW contained more labile compounds, bacteria, and microfauna. The mesofauna did not affect N-content, but the cumulated -N excreted by the mesofauna was estimated to be substantial and about one-fifth of total plant N in ADMSW. An explanation for this discrepancy might be that in the absence of the mesofauna, other members of the detrivore and microbivore community performed the mesofauna’s function. Neither enchytraeids nor microarthropods excreted urea.     

Faculty positions: Center for Agricultural Resources Research,Chinese Academy of Sciences

正The Center for Agricultural Resources Research(CARR),the Institute of Genetics and Developmental Biology(IGDB),Chinese Academy of Sciences,invites applicants for several research group leader positions.CARR is one of the research organizations in Chinese Academy of Sciences(CAS).We seek nominations and applications from individuals who have expertise and a record of accomplishment in research areas related to ecology,agro-hydrology,     

Faculty positions: Center for Agricultural Resources Research,Chinese Academy of Sciences

正The Center for Agricultural Resources Research(CARR),the Institute of Genetics and Developmental Biology(IGDB),Chinese Academy of Sciences,invites applicants for several research group leader positions.CARR is one of the research organizations in Chinese Academy of Sciences(CAS).We seek nominations and applications from individuals who have expertise and a record of accomplishment in research areas related to ecology,agro-hydrology,     

Faculty positions:Center for Agricultural Resources Research,Chinese Academy of Sciences

正The Center for Agricultural Resources Research(CARR),the Institute of Genetics and Developmental Biology(IGDB),Chinese Academy of Sciences,invites applicants for several research group leader positions.CARR is one of the research organizations in Chinese Academy of Sciences(CAS).We seek nominations and applications from individuals who have expertise and a record of accomplishment in research areas related to ecology,agro-hydrology,     

Assessing the effectiveness of conservation measures: Resolving the “wicked” problem of the Steller sea lion

“Wicked” problems are those that are complex and that change when solutions are applied. Many conflicts in conservation fall into this category. The study approached the problem of how to constrain the apparent wickedness of a problem in the conservation management of a species by using simple empirical indicators to carry out iterative assessment of the risk to a population and to document how this risk evolves in relation to the addition of new data and the implementation of management actions. Effects of high levels of uncertainty within data and also concerning population structure were examined through stochastic simulation and by exploration of scenarios. Historical trends in the example used, the Steller sea lion, showed rapid declines in abundance in some regions during the 1980s. The current total population is 130,000-150,000 Steller sea lions through Alaska and British Columbia and this number has been stable since about 1990 in spite of regional differences in population dynamics. Regional differences in the sequence of changes in the number of pups and non-pups, suggested that an internal re-distribution of juveniles could have happened between 1980 and 1990. Current productivity also appears close to the long-term mean. Stochastic population projection using various scenarios showed that, based upon this history, the risk of extinction for the population has declined and is below reasonable thresholds for considering the population to be endangered. The trends in risk suggest that management actions taken since 1990 have probably been effective. Consequently, the conservation management objectives for the Steller sea lion are probably being met. The approach provides a mechanism, based upon experience and scenario analysis, for exploring future policy options and may help to constrain the debate amongst stakeholders about the cost-benefit trade-offs associated with different options.     

Soil-profile distribution of primary and secondary plant-available nutrients under conventional and no tillage

Nutrient distributions under no tillage (NT) compared with conventional disk-and-bed tillage (CT) management in the warm, humid region of the southeastern USA need to be assessed so that future placement, quantity, and type of fertilizers can be altered, if necessary, to efficiently match crop demands. We determined soil-profile distributions of pH, N, P, S, K, Ca, Mg, Na, Zn, Fe, Mn, and Cu to a depth of 0.9 m at the end of 8.5 years of continuous CT and NT management on a Weswood silty clay loam (fine, mixed, thermic Fluventic Ustochrept) in southcentral Texas. Most dramatic changes occurred within the 0–0.05 m depth, where soil under NT had lower pH, Fe, and Cu than under CT, but greater P, K, Zn, and Mn. Greater P and K under NT than under CT also occurred below the till-zone (0.15–0.3 m). At a depth of 0–0.3 m, soil under NT contained greater amounts of extractable P, K, Zn, Fe, Mn, and Cu than under CT. Nitrogen fertilization had little effect on nutrient distributions, except resulting in greater extractable K at 0–0.05 m and greater nitrate at 0–0.15 m. Few changes in soil-profile distributions were observed for extractable S, Ca, Mg, and Na. Long-term continuous use of NT on this fine-textured, high-fertility (except for N) soil had no apparent adverse effects on nutrient distributions relative to CT, but enhanced conservation and availability of P, K, Zn, Fe, Mn, and Cu near the soil surface where crop roots proliferate.     

Macroinvertebrates in North American tallgrass prairie soils: effects of fire, mowing, and fertilization on density and biomass

The responses of tallgrass prairie plant communities and ecosystem processes to fire and grazing are well characterized. However, responses of invertebrate consumer groups, and particularly soil-dwelling organisms, to these disturbances are not well known. At Konza Prairie Biological Station, we sampled soil macroinvertebrates in 1994 and 1999 as part of a long-term experiment designed to examine the effects and interactions of annual fire, mowing, and fertilization (N and P) on prairie soil communities and processes. For nearly all taxa, in both years, responses were characterized by significant treatment interactions, but some general patterns were evident. Introduced European earthworms (Aporrectodea spp. and Octolasion spp.) were most abundant in plots where fire was excluded, and the proportion of the total earthworm community consisting of introduced earthworms was greater in unburned, unmowed, and fertilized plots. Nymphs of two Cicada genera were collected (Cicadetta spp. and Tibicen spp.). Cicadetta nymphs were more abundant in burned plots, but mowing reduced their abundance. Tibicen nymphs were collected almost exclusively from unburned plots. Treatment effects on herbivorous beetle larvae (Scarabaeidae, Elateridae, and Curculionidae) were variable, but nutrient additions (N or P) usually resulted in greater densities, whereas mowing usually resulted in lower densities. Our results suggest that departures from historical disturbance regimes (i.e. frequent fire and grazing) may render soils more susceptible to increased numbers of European earthworms, and that interactions between fire, aboveground biomass removal, and vegetation responses affect the structure and composition of invertebrate communities in tallgrass prairie soils.