疏叶骆驼刺幼苗根系生态学特性对水分处理的响应

通过人工壕沟挖掘法,对疏叶骆驼刺幼苗在不同水分条件下的根系生态学特性的季节变化进行研究.结果表明:①在土壤水分较好的环境中,疏叶骆驼刺幼苗大量拓展水平根,并产生分蘖植株竞争光照资源;而在土壤水分相对缺乏时垂直根系发达,向深层土壤拓展资源空间.根系形态的可塑性是疏叶骆驼刺幼苗获取水分适应干旱环境的重要策略之一.②根冠比随土壤水分的减少而增加,且在生长季后期这种趋势更加明显,增加根冠比是疏叶骆驼刺幼苗适应干旱的策略之一.③幼苗根系的扎根深度和垂直生长速度随着土壤水分的减少而增加.④根系生物量、根系表面积随土壤深度的增加而减小,并在垂直剖面上呈倒金字塔状分布.根系生物量形成过程符合Logistic"慢-快-慢"的S型生长曲线,总生物量随着土壤干旱程度的增加而降低.     

Optimal root system strategies for desert phreatophytic seedlings in the search for groundwater

Desert phreatophytes are greatly dependent on groundwater, but how their root systems adapt to different groundwater depths is poorly understood. In the present study, shoot and root growths of Alhagi sparsifolia Shap. seedlings were studied across a gradient of groundwater depths. Leaves, stems and roots of different orders were measured after 120 days of different groundwater treatments. Results indicated that the depth of soil wetting front and the vertical distribution of soil water contents were highly controlled by groundwater depths. The shoot growth and biomass of A. sparsifolia decreased, but the root growth and rooting depth increased under deeper groundwater conditions. The higher ratios of root biomass, root/shoot and root length/leaf area under deeper groundwater conditions implied that seedlings of A. sparsifolia economized carbon cost on their shoot growths. The roots of A. sparsifolia distributed evenly around the soil wetting fronts under deeper groundwater conditions. Root diameters and root lengths of all orders were correlated with soil water availabilities both within and among treatments. Seedlings of A. sparsifolia produced finer first- and second-order roots but larger third- and fourth-order roots in dry soils. The results demonstrated that the root systems of desert phreatophytes can be optimized to acquire groundwater resources and maximize seedling growth by balancing the costs of carbon gain.     

Root characteristics of Alhagi sparsifolia seedlings in response to water supplement in an arid region,northwestern China

The effect of variation in water supply on woody seedling growth in arid environments remain poorly known.The subshrub Alhagi sparsifolia Shap.(Leguminosae),distributed in the southern fringe of the Taklimakan Desert,Xinjiang,northwestern China,has evolved deep roots and is exclusively dependent on groundwater,and performs a crucial role for the local ecological safety.In the Cele oasis,we studied the responses of A.sparsifolia seedling roots to water supplement at 10 and 14 weeks under three irrigation treatments(none water supply of 0 m3/m2(NW),middle water supply of 0.1 m3/m2(MW),and high water supply of 0.2 m3/m2(HW)).The results showed that the variations of soil water content(SWC) significantly influenced the root growth of A.sparsifolia seedlings.The leaf area,basal diameter and crown diameter were significantly higher in the HW treatment than in the other treatments.The biomass,root surface area(RSA),root depth and relative growth rate(RGR) of A.sparsifolia roots were all significantly higher in the NW treatment than in the HW and MW treatments at 10 weeks.However,these root parameters were significantly lower in the NW treatment than in the other treatments at 14 weeks.When SWC continued to decline as the experiment went on(until less than 8% gravimetric SWC),the seedlings still showed drought tolerance through morphological and physiological responses,but root growth suffered serious water stress compared to better water supply treatments.According to our study,keeping a minimum gravimetric SWC of 8% might be important for the growth and establishment of A.sparsifolia during the early growth stage.These results will not only enrich our knowledge of the responses of woody seedlings to various water availabilities,but also provide a new insight to successfully establish and manage A.sparsifolia in arid environments,further supporting the sustainable development of oases.     

科尔沁差巴嘎蒿根系分布规律与土壤水分关系的研究

以生长在科尔沁大青沟两侧台面上的差巴嘎蒿为研究对象,采用整株挖掘的方法,研究差巴嘎蒿根系分布的规律。结果表明:差巴嘎蒿根系垂直分布深度为0～90㎝,其中0～30㎝的土层中根系分布最多,占根系总长度的85.74%,水平方向上表现为近密远疏的趋势;整体上根系呈主根型分布,形态上呈"伞"型分布。差巴嘎蒿根系分布与土壤水分密切相关。随着深度的增加,土壤含水量逐渐增大,但根系长度、生物量及密度均逐渐降低,两者呈负相关关系,其中根长平均变化率-12.61%、根生物量平均变化率-29.07%,均大于土壤含水量平均变化率7.92%,但比根长却与土壤含水量呈正相关关系,且比根长平均变化率7.58%与土壤含水量平均变化率7.92%相差不大。     

毛乌素沙地不同水分梯度根系垂直分布与土壤水分关系的研究

通过对毛乌素沙地不同水分梯度根系垂直分布的研究表明:不同水分梯度根系分布随土壤深度的增加呈指数形式下降,不同梯度由于土壤水分、植被根系类型的差异,不同水分梯度植被根系的空间分布也有差异,不同梯度根量随土壤垂直深度的模拟方程分别为:y=9.5736e-0.1341x,R2=0.8859(I);y=16.246e-0.2037x,R2=0.9301(II);y=32.001e-0.1904x,R2=0.9544(III);y=28.336e-0.1993x,R2=0.9484(IV)。不同水分梯度土壤含水率变化程度不同,同一水分梯度各层土壤含水率变化幅度亦有差异,0.1m土壤含水率变异系数最大,随着土壤深度增加变异系数减小,根据土壤含水率变异系数分析我们将各层土壤水分垂直变化划分为活跃层(0-0.1m)、次活跃层(0.2-0.6m)、相对稳定层(1m)(I、II、III);而对IV梯度划为活跃层(0-0.1m、1m)、相对稳定层(0.3m)。根系生物量垂直分布与其对应土壤含水率有明显相反的关系,土壤含水率的变化与根系生物量的变化趋于相反,当土壤含水率增大时相应区域根系生物量减小;反之则增加。随水分梯度的增加各梯度最高水分利用层逐渐向表层发展,从第I梯度0.4m的6.84到第IV梯度0.1m的14.33。     

干旱胁迫下AMF对多枝柽柳幼苗和疏叶骆驼刺根系生长和氮素吸收分配的影响

丛枝菌根真菌(Arbuscular Mycorrhizal Fungi,AMF)对植物抗旱、养分吸收等有重要作用,但在特定环境胁迫下不同生活型植物对AMF的响应存在差异。本文以塔里木河下游荒漠河岸林的优势灌木多枝柽柳(Tamarix ra?mosissima)和常见半灌木疏叶骆驼刺(Alhagi sparsifolia)为研究对象,分析了干旱胁迫处理下(对照组土壤相对含水量为70%±5%、实验组土壤相对含水量为20%±5%)接种AMF(对照组不接菌M-、实验组接菌M+)对多枝柽柳与疏叶骆驼刺混合种植(对照组单一种植)根系生长状况和氮素吸收分配的影响。结果表明:(1)植物遭受干旱胁迫时,多枝柽柳幼苗和疏叶骆驼刺菌根侵染率均降低了,混合种植显著增加了多枝柽柳幼苗的菌根侵染率(P<0.05);(2)干旱胁迫下,混合种植M+处理显著增加了多枝柽柳幼苗的地上、地下生物量;(3)干旱胁迫下,AMF使不同种植模式下两种植物的细根根长和细根表面积均显著增加,使疏叶骆驼刺的比根长显著减小,且混合种植M+处理显著减小了多枝柽柳幼苗的细根比根长;(4)相比单一种植,干旱胁迫下AMF显著增加了混合种植多枝柽柳幼苗的氮摄取量和地上部分氮分配比率。因此,AMF对于干旱胁迫下与疏叶骆驼刺混生的多枝柽柳幼苗的生长和氮素吸收具有明显的补偿作用,能够帮助塔里木河下游多枝柽柳幼苗较好地度过生长脆弱期。     

刺槐和油松根系密度分布特征研究

通过分层挖掘法,对20龄刺槐、油松林根系密度的空间分布进行了研究.结果表明:油松主根和副主根粗壮发达,水平根系形成构架根;刺槐水平根极为发达,无明显主根.刺槐和油松根系重量、长度和体积密度的垂直分布趋势基本相同,随土层深度增加而降低.0～60 cm土层的根长和根重密度占总根量密度的70%以上,粗根(直径＞3 mm)在上层所占比例比下层略大,细根(直径＜1 mm)和较细根(直径1～3 mm)在各土层分布较为接近,总根量中粗根占较大比重.刺槐和油松林下土壤干密度随深度加深而增大,油松林下土壤含水量高于刺槐林,土壤干密度低于刺槐林.与刺槐林地相比,油松林地的土壤物理性质得到了明显改善,增强了土壤的抗旱、保墒能力.     

利用HYDRUS-2D模拟膜下滴灌玉米农田深层土壤水分动态与根系吸水

河套灌区农田地下水埋深普遍较浅且年内波动较大,明确不同膜下滴灌条件下深层土壤水分对根区的补给作用及作物根系吸水的响应差异有利于膜下滴灌技术的完善和推广。本研究开展了连续2 a(2017—2018年)的春玉米田间试验,设置3个膜下滴灌灌溉水平,分别控制土壤基质势下限为-10 kPa(S1)、-30 kPa(S3)和-50 kPa(S5)。利用HYDRUS-2D模型模拟0～120 cm深度土壤含水量、根层下边界(100 cm深度处)水分通量和作物根系吸水速率。结果表明,经过率定后的HYDRUS-2D模型对0～120 cm深度土壤含水量模拟结果的根均方差(RMSE)和决定系数(R~2)分别为0.039～0.042 cm~3·cm~(-3)和0.78～0.73,模拟结果可靠。膜下滴灌农田100 cm和120 cm深度处土壤含水量较高且处理间差异不大,说明不同滴灌条件对于100 cm以下深层土壤含水量影响较小;但不同处理显著影响根区下边界的水分通量和根系吸水速率。基质势下限控制水平越低,深层土壤水分对于根区的补给量(毛管上升)越大,S1、S3、S5生育期内累积补给量在31.9～49.6 mm之间。S5处理根系吸水速率较低,根系吸水受到显著抑制,从而造成作物生长指标和产量显著低于S1和S3处理(P0.05);而S1和S3之间籽粒产量差异不显著。综上,在本研究所设置的3个滴灌处理中,S3生育期内灌溉定额为240～300 mm,既较S1显著减少灌水量、提高水分利用效率,又具有较好的根系活力,有效利用深层土壤水分,因此建议该地区春玉米膜下滴灌的灌水下限为-30 kPa。     

植物根系水力提升作用研究进展综述

植物根系水力提升作用是植物深层根系吸收深层土壤水分,再通过浅层根系释放到较干燥浅层土壤中的过程。水力提升作用所释放的水分可缓解干燥土层水分匮乏,对改善自身及邻近植物的水分亏缺、促进植物养分吸收、促进土壤养分分解和矿化过程具有重要意义。随着对植物生态系统水分平衡问题的关注和研究手段的发展,水力提升作用日益成为干旱、半干旱区植物生态学与水文学共同关注的重要研究内容之一。通过回顾和分析水力提升作用的研究历史、影响因素、提升数量、研究方法、生态作用等,特别阐述了水力提升作用在半干旱区人工固沙植被恢复中的潜在应用价值,改善人工固沙植被区土壤水分条件,提高固沙植物群落的稳定性。     

浑善达克沙地榆树根系分布特征及生物量研究

本文采用挖掘法对浑善达克沙地榆树的根系进行了研究。研究表明 ,草被层的根系主要分布在土壤表层 ,榆树根系则占据了 2 0 cm以下的土层空间。根系的垂直分布在冠缘内主要集中在 2 m深范围内 ,冠缘外则集中分布在 1 m深的土层中 ,不同的根系剖面形成不同的根栖层 ;根系的水平分布呈近密远疏的趋势 ,其半径为 1 1 m左右。根量的垂直分布随土层深度增加呈先增加后降低的趋势 ;其水平分布随着距根基距离的增大而减少 ,以 3m内最为密集 ,冠缘内根量为冠缘外根量的 1 2倍。     

Root growth and spatio-temporal distribution of three common annual halophytes in a saline desert, northern Xinjiang

Root growth and spatial and temporal distribution in the 0-100 cm soil profiles of three common annual halophytes Salsola subcrassa, Suaeda acuminate and Petrosimonia sibirica distributed in a saline desert in northern Xinjiang, China were studied in 2009 and 2010. The results showed that the root systems of the three halophytes were of the taproot type, vertically distributed in the 90-cm soil profile, and were deepest in late July. Their taproots reached maximum depth rapidly, early in the growth period, but with rare lateral roots. They were then dug out in an orderly way, from bottom to top, exhibiting vertical development first and then horizontal development. The distribution of specific root length, which reflects the characteristics of the feeder root, was gradually increased from top to bottom, whereas root weight displayed an opposite distribution pattern. The root length distribution of the three halophytes was concentrated (62% to 76%) in the middle soil profile (20-60 cm), with less distribution in the surface (0-20 cm) and bottom (60-90 cm) soil profiles. The results indicated that the roots of the three annual halophytes grew rapidly into the deeper soil layer after germination, which ensured the plant survival and uptake of water and nutrition, and thus built up a strong tolerance to an arid, high-salt environment.     

Warming effects on plant biomass allocation and correlations with the soil environment in an alpine meadow,China

Alpine meadow ecosystem is fragile and highly sensitive to climate change.An understanding of the allocation of above-and below-ground plant biomass and correlations with environmental factors in alpine meadow ecosystem can result in better protection and effective utilization of alpine meadow vegetation.We chose an alpine meadow in the Qinghai-Tibetan Plateau of China as the study area and designed experimental warming plots using a randomized block experimental design.We used single-tube infrared radiators as warming devices,established the warming treatments,and measured plant above- (AGB) and below-ground biomass (BGB) during the growing seasons (May to September) in 2012 and 2013.We determined the allocation of biomass and the relationship between biomass and soil environment under the warming treatment.Biomass indices including above-ground biomass,below-ground biomass and the ratio of root to shoot (R/S) ,and soil factors including soil moisture and soil temperature at different depths were measured.The results showed that (1) BGB of the alpine meadow had the most significant allometric correlation with its AGB (y=298.7x~ (0.44) ,P0.001) ,but the relationship decreased under warming treatment and the determination coefficient of the functional equation was 0.102 which was less than that of 0.188 of the unwarming treatment (control) ; (2) BGB increased,especially in the deeper soil layers under warming treatment (P0.05) .At 0–10 cm soil depth,the percentages of BGB under warming treatment were smaller than those of the control treatment with the decreases being 8.52% and 8.23% in 2012 and 2013,respectively.However,the BGB increased 2.13% and 2.06% in 2012 and 2013,respectively,at 10–50 cm soil depths; (3) BGB had significant positive correlations with soil moisture at 100 cm depth and with soil temperature at 20–100 cm depths (P0.05) ,but the mean correlation coefficient of soil temperature was 0.354,greater than the 0.245 of soil moisture.R/S ratio had a significant negative correlation with soil temperature at 20 cm depth (P0.05) .The warmer soil temperatures in shallow layers increased the biomass allocation to above-ground plant parts,which leading to the increase in AGB;whereas the enhanced thawing of frozen soil in deep layers causing by warming treatment produced more moisture that affected plant biomass allocation.     

Untangling the influence of soil moisture on root pullout property of alfafa plant

Root pullout property of plants was of key importance to the soil reinforcement and the improvement of slope stability. To investigate the influence of soil moisture on root pullout resistance and failure modes in soil reinforcement process, we conducted pullout tests on alfalfa (Medicago sativa L.) roots at five levels (40, 30, 20, 10 and 6 kPa) of soil matric suction, corresponding to respectively 7.84%, 9.66%, 13.02%, 19.35% and 27.06% gravimetric soil moisture contents. Results showed that the maximal root pullout force of M. sativa decreased in a power function with increasing soil moisture content from 7.84% to 27.06%. Root slippage rate increased and breakage rate decreased with increasing soil moisture content. At 9.66% soil moisture content, root slippage rate and breakage rate was 56.41% and 43.58%, respectively. The threshold value of soil moisture content was about 9.00% for alfalfa roots in the loess soil. The maximal pullout force of M. sativa increased with root diameter in a power function. The threshold value of root diameter was 1.15 mm, because root slipping force was greater than root breaking force when diameter >1.15 mm, while diameter ≤1.15 mm, root slipping force tended to be less than root breaking force. No significant difference in pullout forces was observed between slipping roots and breaking roots when they had similar diameters. More easily obtained root tensile force (strength) is suggested to be used in root reinforcement models under the condition that the effect of root diameter is excluded as the pullout force of breaking roots measured in pullout tests is similar to the root tensile force obtained by tensile tests.     

晋西黄土区林草复合系统刺槐根系分布特征

为了研究晋西黄土区林草复合系统树木根系分布特征,采用钻土芯法对刺槐×天然草复合系统0-100 cm土层中刺槐根系生物量、根长和根表面积进行了测定和分析。研究结果表明:同一坡向上刺槐根系生物量、根长和根表面积分布具有相似的特征,在垂直方向上,根系生物量、根长和根表面积均随土层深度的增加呈指数减少,且集中在0～60 cm的土层中;在水平方向上,离树行越近,刺槐根系生物量、根长和根表面积均越大。不同坡向刺槐根系差异显著,相同径级、相同位置阳坡刺槐根系生物量、根长和根表面积均小于阴坡,但根系消弱系数表明阳坡刺槐细根深层土壤生物量分布比例较阴坡多。     

局部高浓度硝酸盐供应对玉米根系 形态及氮累积的影响

以郑单958和鲁单981为研究对象，进行水培分根试验，在正常供水和水分胁迫条件下，分别以均匀低浓度硝酸盐处理主根和种子根(LPR-LSR)、局部高浓度硝酸盐处理主根（HPR-LSR）和局部高浓度硝酸盐处理种子根（LPR-HSR），测定分析根系形态、生物量以及氮含量。结果表明：与氮低效鲁单981相比，氮高效郑单958具有较高的主根根长、根表面积、根系生物量、地上部生物量和氮累积量。水分胁迫条件下，郑单958和鲁单981的主根的根长、根表面积、根体积、地上部生物量和氮累积量总体上均低于正常水分条件。玉米主根和种子根对局部高浓度硝酸盐的反应存在差异。与均匀低浓度硝酸盐处理相比，局部高浓度硝酸盐处理促进正常水分条件下主根和种子根根系的生长，尤其是根长和根系表面积；在正常水分条件下，主根根长和根系表面积增加幅度范围为6.8%~27.3%和1.9%~21.9%，除HPR-LSR处理条件下的郑单958外，种子根根长和根系表面积增加幅度范围为30.4%~92.7%和10.5%~135.1%；在水分胁迫条件下，主根根长和表面积增加幅度范围为24.6%~152.9%和62.1%~229.9%，然而种子根根长降低了10.0%~29.9%，表明水分胁迫会影响种子根对高浓度硝酸盐的响应。除水分胁迫条件下LPR-HSR处理外，局部高浓度处理可同时增加两侧根系的生物量和氮累积量。无论是正常供水还是水分胁迫，与LPR-LSR处理相比，局部高浓度硝酸盐供应均能够增加地上部生物量以及氮累积量，在LPR-HSR处理条件下，增加幅度范围分别在35.0% ~107.9%和162.9%~291.1%，在HPR-LSR处理条件下分别为56.7%~109.4%和204.1%~377.0%，HPR-LSR处理条件下增加幅度较大，表明在氮素非均匀分布环境中，当主根处于高浓度硝酸盐区域时将会更显著促进生物量的增加和氮累积。     

3种因素对油松移植苗根系活力的影响

采用盆栽的方法,以3种不同的光照条件、土壤含水量及苗木含水量,对4 a生油松（Pinus tabulaeformis）移植苗进行处理,测定3种因素交互作用下油松移植苗的根系活力。结果表明：光照强度的降低、苗木含水率的减少及土壤含水量的增加,促使油松移植苗根系活力不断降低;油松移植苗的苗木含水量为63%或土壤含水量为30%时,自然光照下根系活力均显著大于遮光条件下根系活力;在自然光强、63%的苗木含水量及30%土壤含水量其中一种条件影响下,其他两个因素的变化均可引起根系活力的显著变化;在自然光强与53%自然光强条件下,或在土壤含水量为30%和50%条件下,苗木失水均导致根系活力降低。     

高寒沙区中间锦鸡儿人工林细根动态及其周转

通过对青海省治沙试验站3 a、5 a、10 a、16 a和30 a生的中间锦鸡儿(Caragana intermedia Kuang et H. C.Fu)人工林细根生物量的测定分析,了解其细根生物量的垂直分布和动态变化特征以及周转速率。结果表明:①中间锦鸡儿人工林约41%的活细根和39%的死细根分布在0~20 cm的土层,38%的活细根和死细根分布在20~40 cm的土层,21%的活细根和23%的死细根分布在40~60 cm的土层。②细根生物量表现出明显的季节动态性,活细根生物量峰值出现在6月和8月或者6月和9月,8月或9月达到最大值;死细根生物量峰值出现8月和9月,也是8月或9月达到最大值。③不同林龄人工林细根周转速率分别为:0. 53、0. 66、0. 56、0. 73、0. 78次·a^-1。④细根生物量和周转率均随林龄增大而增加,随土层加深而减小。     

中国西北干旱区生态地下水埋深适宜深度的确定

干旱区影响天然植被生长的土壤水分和盐分,都和地下水埋深密切相关。地下水埋藏浅,在强烈的蒸发作用下,溶解于地下水中的盐分沿毛管上升水流于地表聚集,使土壤发生盐渍化,产生盐胁迫。地下水埋深过低,毛管上升水流不能到达植物根系层,植物生长受到水分胁迫,发生荒漠化。因此,确定既不会使土壤发生盐渍化和荒漠化的地下水埋深十分重要。本文根据地下水、土壤水及植物生长状况综合研究,把适宜的地下水埋深界定为在温带荒漠区1.5—4m,暖温带荒漠区2.0~4.0(4.5)m,即在潜水的强烈蒸发深度以下和蒸发极限深度之上的区间,为防治土地盐渍化、荒漠化和估算生态用水提供了科学依据。     

覆盖处理苹果细根分布与土壤物理性状响应关系研究

以12年生红富士苹果树为试材,研究黄土高原旱塬区不同覆盖措施(覆膜、覆草、覆沙)对苹果细根(直径≤2 mm)及土壤性状的影响。结果表明:地表覆盖可有效降低土壤容重,增大土壤含水量及孔隙度,增加细根数量,增大根系吸收水肥效率。覆膜处理细根水平分布范围与清耕(CK)相似,在距干0~90 cm范围内,垂直密集分布最大深度由CK的60 cm提升至40 cm,36.05%的细根分布在0~20 cm土壤表层。与CK相比,覆草、覆沙处理的细根水平分布范围由90 cm扩展至120 cm,垂直均匀分布整个土壤剖面,利于树体对深层土壤水肥的吸收利用。土壤物理性状与细根根长、表面积、根长密度呈极显著相关,覆膜处理中根径与比根长也与土壤物理性状表现出相关性。综合分析根系分布与土壤物理性状,覆草处理是陇东旱塬区苹果园较为适宜的地表覆盖模式。     

浑善达克沙地不同植被下的土壤水分状况

浑善达克沙地的中东段阳坡近裸的流动风沙土 0 - 1 0cm土层整个生长季内平均含水量较低、波动也比较大 ,而在 1 0cm以下则基本稳定在 3%左右 ,尤其是深层。丘间低地草本群落下的土壤湿度随深度增加变化较大 ,2 0cm以上土层含量较高 ,40 - 5 0cm土层的含水量最低 ,波动也剧烈 ,90cm以下达裸沙含水量水平。阴坡灌丛下 ,最低土壤含水量出现在 70 -90cm土层 ,深层土壤含水量长期稳定在 1 .5 %左右。这说明植物的生长使根层土壤含水量下降 ,而且不同植物利用水的土层及利用土壤水的量不同。这可能是引起流动风沙土向固定风沙土转变过程中地表植被演替及决定地区顶级群落的主要原因。在干旱半干旱地区 ,植被影响着降水在土层中的分布及地表的蒸散条件 ,使土壤有效水向浅层分配。而降水在土壤不同深度的分配及入渗深度 ,决定着地表植被的生活型 ,从而影响地表植被的演替方向及顶级类型。需水量较大的灌丛在沙丘阴坡能形成稳定的群落 ,是由于沙丘为植物避免干旱胁迫 ,延长在干旱季节的生存时间创造了条件 ,因此 ,沙丘是沙地的重要环境资源 ,尤其是高大沙丘 ,在这类地区的防风固沙中具有重要的作用。