森林土壤酶的研究进展

土壤酶在土壤生态系统的物质循环和能量流动方面扮演重要的角色。目前 ,在几乎所有的森林生态系统研究中 ,土壤酶活性的监测似乎成为必不可少的研究内容。森林凋落物分解过程中的酶活性动态 ,植被特征与土壤酶活性的关系 ,土壤微生物与土壤酶的关系 ,植物 -土壤界面的土壤酶 ,森林土壤质量评价指标的土壤酶及人类活动干扰对森林土壤酶活性的影响等是当前森林土壤酶学的研究重点。由于土壤酶的功能和生态重要性 ,森林土壤酶研究可能包括 :(1 )土壤酶系统分异 ;(2 )作为森林土壤质量综合评价指标的土壤酶活性 ;(3)植被动态与土壤酶的关系 ;(4 )退化森林生态系统的土壤酶活性特征 ;(5 )人工林土壤酶活性特征 ;(6 )人类活动对森林土壤酶系统的影响。本文从土壤酶系统分异和生态系统的角度对土壤酶在森林生态系统中的作用和地位进行了综述 ,这对于加深理解森林生态系统中的物质循环、土壤酶的生态重要性以及森林生态系统退化机理有重要作用     

森林土壤微生物分布及其功能特征研究进展

土壤微生物是森林生态系统中促进养分循环、能量流动的重要推动力。根据目前对森林土壤微生物的研究现状,文中对森林土壤微生物的分布特征（地理区域、季节变化和垂直分布）、功能（促进碳氮循环）以及环境影响因素（植被类型、气候条件、土壤性质）和人为影响因素进行归纳和总结,对将来研究方向提出研究展望,以期为今后的土壤微生物生态学研究和森林经营实践提供参考。     

林火对森林土壤有机碳影响的研究进展

指出了森林生态系统土壤有机碳是全球碳循环的重要组成,在全球生态系统碳循环和全球变化的研究中占据着重要的地位。火因子是干扰森林生态系统发展的重要因子之一,对森林生态系统中土壤与大气的碳素交换过程有着重要的影响。因此,加强火干扰对森林生态系统土壤有机碳的研究,了解林火对森林生态系统土壤有机碳循环过程的具体影响,对发现火干扰下森林土壤中碳循环的机理,特别是土壤有机碳的动态变化有着重要的作用。综述了林火对森林土壤有机碳的研究进展,讨论了测定土壤有机碳的几种不同方法,比较了其适用性,探讨了在不同林火强度和不同土壤深度条件下土壤有机碳的变化情况。对实际工作中遇到的问题以及今后森林土壤有机碳研究进行了展望。     

森林生态系统固碳过程对氮沉降增加的响应机制研究进展

氮沉降增加对森林生态系统固碳过程具有重要影响。文中介绍了森林生态系统固碳过程对氮沉降增加响应机制的研究进展,概述了氮沉降增加对植被固碳过程的影响以及土壤碳输入和碳排放对氮沉降增加的响应机理,探讨未来该领域的研究重点,指出在全球氮沉降增加背景下森林植被和土壤固碳分配、森林生态系统固碳过程植被和土壤界面碳、氮循环过程的耦合机理将是未来森林碳循环研究的热点。     

采伐对森林土壤碳库影响的不确定性

森林土壤有机碳(SOC)是全球碳循环的重要组成部分,然而,多样的森林类型和不同森林经营措施的干扰,使得森林土壤碳库维持机制以及碳固存过程的研究和森林土壤碳库的估算存在较大的变异。作为主要的森林经营措施之一,采伐对森林土壤碳储量以及碳过程均产生直接或间接地影响。为深刻理解森林土壤碳库对于采伐干扰的响应,本文综述了近十几年来不同采伐方式下森林土壤碳储量及其主要碳排放过程——土壤呼吸的研究现状,综合分析了采伐方式、森林类型、采伐剩余物管理以及微生物因子等对土壤碳库的影响及其不确定性,并在此基础上阐述了研究中尚未解决的主要问题:1)生物因子作为CO₂产生的主体,在应对干扰时结构、功能的变化直接影响着土壤碳排放以及碳固定,但它们具体作用机制以及过程并不清楚,需展开进一步的调查;2)不同森林采伐方式对不同地区和不同类型森林土壤的影响的复杂性,亟须在进一步加强实验研究的基础上,发展森林土壤碳循环的过程或机理模型,为森林生态系统完整的碳循环过程表达及碳计量提供技术支撑,以期为我国森林经营以及碳汇等方面研究提供参考。     

火干扰对土壤呼吸的影响及测定方法研究进展

林火是森林生态系统的重要干扰因子,是导致土壤碳储量发生变化的重要因子。火干扰所排放的含碳气体对物质循环、能量流动和信息传递具有重要影响,进而影响森林生态系统的碳平衡以及气候变化。土壤作为巨大的陆地生态系统储碳库,其与大气碳交换的方式主要是通过土壤呼吸,由于土壤的碳储量巨大,土壤碳库的微小变化都会引起大气CO2浓度的巨大改变,从而影响森林生态系统碳平衡。阐述土壤呼吸的各种测量方法,并对比各种测定方法;火干扰对森林生态系统土壤呼吸速率的影响及其测定方法;火干扰对土壤呼吸环境的影响。最后提出今后在火干扰对土壤呼吸研究中应关注的问题,同时对火干扰与森林生态系统土壤呼吸的研究方向进行展望。     

林火干扰对土壤氮组分及微生物氮循环影响研究进展

林火干扰作为森林生态系统中能量传递和养分循环的重要因子,对于研究火烧迹地植被更新与快速恢复可提供一定的数据理论支撑,而林火干扰对土壤微生境及微生物氮循环的影响机制一直是广大学者研究的热点。文中从火烧强度、恢复时间、火烧木管理方式等3个方面总结国内外林火干扰对土壤氮组分及氮循环的影响研究进展,揭示了林火干扰和全球气候变化对土壤氮循环的短期及长期影响机制,探讨不同生物技术在林火干扰下土壤微生物氮循环基因丰度变化中的应用;提出未来林火干扰对森林生态系统土壤氮循环影响的研究展望:1）全面比较研究不同林火干扰模式对土壤氮组分、微生物氮循环的影响;2）研究评价不同林火干扰模式下土壤氮库的稳定性;3）加强高通量测序、定量PCR技术、宏基因组学、稳定同位素探测等技术在林火干扰与气候变化对森林生态系统影响研究中的应用。研究结果对火烧迹地植被更新、土壤氮库的重建与功能发挥具有重要意义。     

森林土壤呼吸对降雨脉冲的响应研究进展

森林生态系统的土壤碳储量是陆地生态系统碳库的重要组成部分。土壤呼吸则是土壤碳库参与陆地碳循环的主要方式,其受到诸多生物和非生物因子的综合调控。偶发性降雨会引起森林土壤湿度的瞬间增加,进而导致土壤呼吸速率的快速提高。文中介绍了森林土壤呼吸及其组分,综述了降雨脉冲效应的特征和诱导机制（包括物理替代机制、微生物代谢机制、土壤养分限制机制及光化学调控机制等）,分析了当前降雨脉冲研究存在的不足并展望了未来的研究重点,认为森林生态系统是未来开展降雨脉冲效应研究的重要方向,探究森林土壤呼吸不同组分对降雨脉冲的响应差异与机制也是潜在的重要研究方向。     

我国森林土壤微生物多样性研究进展

土壤微生物是森林生态系统必不可少的重要组成部分,承担着森林生态系统的物质转化和能量流动,是林地中最为活跃的部分。大量研究表明,森林土壤微生物多样性存在着明显的时空变异特征,受植被类型、气候条件、土壤肥力及海拔高度等因子的影响。本文从物种、功能、遗传和生态系统多样性方面综述了我国森林土壤微生物多样性研究现状及影响因素,分析了其研究方面存在的主要问题,并展望了今后的研究重点。     

森林土壤碳储量研究综述

通过查阅大量森林生态系统土壤碳储量的文献资料,综述了森林土壤碳储量的研究进展,概述了自然和人为因素对森林土壤碳储量的影响,并分析了今后森林土壤碳的研究趋势。森林土壤碳储量的分布受地形、气候、植被类型、土壤性质等自然及人为因素的综合影响,具有显著的空间异质性特征。人为因素对森林土壤碳储量的影响具有两面性,合理的人为调控能促进森林土壤碳的积累,过度的干扰则会显著降低土壤碳储量。构建全国森林土壤生态监测网络对森林土壤碳进行长期定位观测,并加强新技术应用,建立森林土壤碳库系统是未来森林土壤碳研究的重要方法。     

Stand restoration burning in oak–pine forests in the southern Appalachians: effects on aboveground biomass and carbon and nitrogen cycling

Understory prescribed burning is being suggested as a viable management tool for restoring degraded oak–pine forest communities in the southern Appalachians yet information is lacking on how this will affect ecosystem processes. Our objectives in this study were to evaluate the watershed scale effects of understory burning on total aboveground biomass, and the carbon and nitrogen pools in coarse woody debris (CWD), forest floor and soils. We also evaluated the effects of burning on three key biogeochemical fluxes; litterfall, soil CO₂ flux and soil net nitrogen mineralization. We found burning significantly reduced understory biomass as well as the carbon and nitrogen pools in CWD, small wood and litter. There was no significant loss of carbon and nitrogen from the fermentation, humus and soil layer probably as the result of low fire intensity. Burning resulted in a total net loss of 55 kg ha⁻¹ nitrogen from the wood and litter layers, which should be easily replaced by future atmospheric deposition. We found a small reduction in soil CO₂ flux immediately following the burn but litterfall and net nitrogen mineralization were not significantly different from controls throughout the growing season following the burn. Overall, the effects of burning on the ecosystem processes we measured were small, suggesting that prescribed burning may be an effective management tool for restoring oak–pine ecosystems in the southern Appalachians.     

自然因子对中国森林土壤碳储量的影响分析

文中分析了森林植被、土壤属性、立地条件、气候条件以及凋落物和根系输入等自然因素对中国森林土壤碳储量的影响。森林植物种类组成决定了进入土壤的植物残体量和分解速率,导致土壤有机碳的含量及分布有很大差异。随着林龄的增加,土壤碳储量会呈现增加或产生波动2种情况。土壤理化性质影响土壤有机碳的含量,而土壤碳储量又影响着土壤结构、根系深度、土层特性、有效水分保持能力、土壤生物多样性等; 海拔、坡度、坡向、坡位等立地条件对森林土壤有机碳储量的影响各不相同; 温度、水分、CO₂浓度等气候因子在森林土壤有机碳的蓄积过程中起着至关重要的作用; 凋落物和根系对土壤的输入也可以改变土壤碳库。     

Country-level carbon balance of forest soils: a country-specific model based on case studies in Hungary

International agreements require countries to annually report on greenhouse gas emissions and removals. For the land-use sector, this includes estimating stock changes in various carbon pools. For carbon pools like mineral forest soil where a country-level statistical inventory based on measurements is very difficult, models are usually applied together with data from case studies. In this paper, we present a country-specific model together with case studies that aim at capturing major soil processes due to forestry activity. These processes include “hot moments”, e.g., disturbances that occur rarely but might result in relatively high emissions. The model only aims at developing a conservative estimate, rather than a central one, of net country-level carbon stock change with emissions overestimated and removals underestimated. The model is partially parameterised using paired sampling of soil organic carbon in the uppermost 30-cm layer, applying standard methods including those suggested by IPCC, in afforestations on former croplands and in artificial regenerations. Results show that soils of afforested croplands act as a sink, and carbon stock after regeneration might decrease due to disturbance by forest operations, but might also increase due to transfer of carbon from dead roots to soil depending on disturbance levels. The estimation at the country level, which involves additional considerations and data from the literature, suggests that overall, forest soils are a net sink in Hungary, but also that artificially limiting soil organic carbon changes estimation to the uppermost 30-cm layer as applied in the IPCC methodology might lead to artefacts.     

氮沉降对森林土壤碳动态过程的影响

本文综述了国内外关于森林土壤碳动态过程对氮沉降响应机制的研究进展,概述了大气氮沉降对土壤碳释放及其影响因子的作用机制,从土壤生物学特性、凋落物动态、土壤碳释放等方面揭示大气氮沉降对土壤碳平衡过程影响机制和机理,探讨了森林土壤碳动态过程对氮沉降响应的不确定性因素,并指出未来该领域研究重点。     

康禾自然保护区常绿阔叶林土壤质地与土壤碳氮含量的关系研究

在广东省东源县康禾自然保护区常绿阔叶林设置4 hm2样地,按＜0.01 mm土粒的含量从小到大将土壤质地分为3个类型,依次为：轻壤土、中壤土、重壤土,通过实地采样统计分析了不同土壤质地表层土壤有机碳含量、全氮含量、碳氮比（C/N）的大小及其关系。结果表明：土壤表层有机碳含量及全氮含量大小依次为：重壤土>中壤土>轻壤土。样地内土壤表层有机碳含量和全氮含量的最大值分别为31.343 g?kg-1、1.882 g?kg-1,最小值分别为8.411 g?kg-1、0.621 g?kg-1。本研究表明保护区土壤中碳氮含量在空间上存在异质性且中壤土最能代表整个样地土壤表层碳氮含量的整体情况。此外,不同土壤质地有机碳含量及碳氮比（C/N）存在显著差异（P＜0.05）,而全氮含量差异不显著。土壤有机碳含量与土壤全氮含量之间呈极显著的正相关关系（P＜0.001）,相关系数为r=0.833 9,说明不同土壤质地土壤有机碳含量和全氮含量有极高的相关性。     

Investigating biochemical processes to assess deadwood decay of beech and silver fir in Mediterranean mountain forests

Context

We investigated the relationships between decay classes, morphological characteristics and chemical compounds in Abies alba Mill. and Fagus sylvatica L. stumps in two forest stands in the central Apennines (Italy). The analysis of deadwood decomposition is important in estimating carbon sequestration potential and carbon residence time in forest ecosystems. In addition, deadwood decomposition affects nutrient cycling and microhabitat distribution.

Aims

The overall aim of this study was to investigate the decomposition processes in a mountainous Mediterranean ecosystem, and specifically to assess changes in chemical variables (lignin, cellulose, carbon and nitrogen content) in deadwood during the decomposition process, in relation to decay classes and to the species.

Methods

Cross-sections were collected from stumps. Samples were assigned to visually discernible decay classes, and cellulose, lignin and carbon (C) and nitrogen (N) content in relation to total dry mass were analysed during the decay process.

Results

Results showed how C/N ratio decreased over decades due to an increase in nitrogen content. On the contrary, carbon concentration in stumps remained substantially stable. Lignin degraded slowly in comparison with cellulose and differences between species were observed.

Conclusion

The hypothesis that the conifer species would decompose faster than deciduous species did not prove correct. The slower decay of lignin compared to cellulose suggested that lignin could be an important long-term source of soil organic carbon, and that this process could be greatly affected by forest species mixtures. Finally, decomposition processes in the investigated montane?CMediterranean forests were definitely faster than in the colder climates of northern Europe.     

Carbon stocks in coffee agroforests and mixed dry tropical forests in the western highlands of Guatemala

Tree removal in Latin American coffee agroforestry systems has been widespread due to complex and interacting factors that include fluctuating international markets, government-supported agricultural policies, and climate change. Despite shade tree removal and land conversion risks, there is currently no widespread policy incentive encouraging the maintenance of shade trees for the benefit of carbon sequestration. In facilitation of such incentives, an understanding of the capacity of coffee agroforests to store carbon relative to tropical forests must be developed. Drawing on ecological inventories conducted in 2007 and 2010 in the Lake Atitlán region of Guatemala, this research examines the carbon pools of smallholder coffee agroforests (CAFs) as they compare to a mixed dry forest (MDF) system. Data from 61 plots, covering a total area of 2.24 ha, was used to assess the aboveground, coarse root, and soil carbon reservoirs of the two land-use systems. Results of this research demonstrate the total carbon stocks of CAFs to range from 74.0 to 259.0 Megagrams (Mg)?C ha^?1 with a mean of 127.6?±?6.6 (SE)?Mg?C ha^?1. The average carbon stocks of CAFs was significantly lower than estimated for the MDF (198.7?±?32.1?Mg?C?ha^?1); however, individual tree and soil pools were not significantly different suggesting that agroforest shade trees play an important role in facilitating carbon sequestration and soil conservation. This research demonstrates the need for conservation-based initiatives which recognize the carbon sequestration benefits of coffee agroforests alongside natural forest systems.     

Biogeochemical cycling and chemical fluxes in a managed northern forested wetland, Michigan, USA

Forest harvesting and subsequent regeneration treatments may cause changes in soil and solution chemistry that adversely affect forest productivity and environmental quality. The objective of this study was to assess soil carbon (C), nitrogen (N), and base cation pools and fluxes, and to construct a hydrogen ion (H⁺) mass balance to identify major processes controlling acidity production and consumption 14 years following whole-tree harvesting and regeneration in a northern forested wetland with underlying mineral soils derived from calcareous glacial drift. Results for soil elemental and nutrient pools in the harvested/regenerated stand were compared to an adjacent non-harvested stand and a riparian zone. The riparian zone had the highest soil total C, total N, and exchangeable calcium (Ca) and magnesium (Mg) pools; however, no difference in exchangeable potassium (K) was evident among stand types. Moreover, no differences between the harvested/regenerated and uncut stands were evident in any of the soil chemical pools.Net export of base cations was minimal and the H⁺ mass balance indicated that net cation exchange was not a significant process in H⁺ production or consumption in either the uncut or harvested/regenerated stands. The most striking differences in the H⁺ mass balance were (1) eight times the H⁺ consumption from sulfate (SO₄²⁻) reduction in the harvested/regenerated stand compared to that in the uncut condition and (2) nearly twice the H⁺ production due to N immobilization in the harvested/regenerated stand. However, both stand types were net H⁺ sinks and increases in H⁺ export due to whole-tree harvesting were not evident.The riparian zone was a net exporter of base cations. This finding was attributed to a combination of base cation exchange and carbonate mineral weathering; data suggested the importance of the latter. More research, however, is required to isolate the contributions of cation exchange and carbonate weathering on base cation export from the riparian zone. Stream chemistry was consistent with that of the riparian zone, indicating a strong linkage between the riparian zone stream chemistry, and whole-tree harvesting had no intermediate term (i.e., 14 years) effects on stream acidification in this managed northern wetland ecosystem.     

Soil enzyme activities as potential indicators of soluble organic nitrogen pools in forest ecosystems of Northeast China

Context

Soluble organic nitrogen is considered to reflect the effect of forest types on soil nitrogen status. As a major process affecting the soil-soluble organic nitrogen pool, degradation of insoluble organic nitrogen in the production of soluble organic nitrogen is mediated by a suite of soil enzymes.

Aims

This study aims to examine soil-soluble organic nitrogen pools and their relationships with the activities of soil enzymes in natural secondary forest stands and adjacent larch plantation stands.

Methods

Four pairs of larch plantation stands and secondary forest stands were randomly selected from a mountainous area, and the top 15?cm of the mineral soils were sampled from each field.

Results

The soil-soluble organic nitrogen concentrations were up to 2-fold greater in the secondary forest stands than in the larch plantation stands, whereas the ratio of soluble organic nitrogen/total nitrogen was comparable between the two forest types. The concentrations of soluble organic nitrogen were positively correlated with approximately 2-fold differences in urease and protease activities, a 1.2-fold difference in N-acetyl-??-glucosaminidase and a 1.7-fold difference in l-asparaginase between the two forest types.

Conclusions

Our results suggest that relationships between soil-soluble organic nitrogen and enzyme activities are independent on sampling time, and that the soil enzyme activities can be used as potential indicators of soil soluble organic nitrogen pools in the temperate forest ecosystem.