Seed rain of ponderosa pine beneath partial overstories

An analysis was conducted of the 1999 ponderosa pine (Pinus ponderosa var. ponderosa Dougl. ex Laws.) seed crop at two stands of differing overstory density in each of two sites that differed in productivity (forest community series) on the east slope of the Cascade Range in central Oregon, USA. A total 2,166 viable seeds were collected between 2 September 1999 and 6 June 2000. Total seed quantities varied more by site than by tree density, with the xeric site producing six-fold greater seed yields. Within each site, the stand of higher density produced more viable seed. Per-tree cone yields were also greater at the site of lesser productivity. At three of the four stands, tree fecundity (as measured by cone counts) was positively related to diameter at breast height and height, but not to live crown ratio. A difference in temporal seedfall patterns among the two sites occurred late and was relatively minor: most seedfall occurred within the first month after it started regardless of absolute seed production. Seedfall distribution varied spatially within stands, but no seed trap (of 55) received less than an equivalent 28 thousand seeds per hectare. Seedfall was not perceived to be a limiting factor on natural regeneration at any of the four partial-overstory stands in this study.     

栎属树种生长模型研究进展

栎属树种是亚热带常绿阔叶林和温带落叶阔叶林的主要建群树种之一,分布范围极为广泛,在国内外被广泛应用于城市园林绿化、水源涵养林、水土保持林,也是重要的珍贵用材树种,同时其果实、栓皮等具有重要的工业和药用价值.研究栎林的生长过程,对其经营抚育决策具有重要的参考意义.文中对近年来国内外关于栎属树种的生长过程,特别是栎属树种生长模型的研究进行了综述,包括栎属树种全林分生长模型、单木生长模型、径阶分布模型等;阐述了栎属树种生长模型的研究现状及发展趋势,以期为栎林的经营抚育决策提供参考.     

The effects of selective logging on stand structure and the regeneration of subboreal forests in Hokkaido, northern Japan

We investigated the effects of selective logging on stand structure and regeneration in selectively logged subboreal forests in Taisetsuzan National Park in Hokkaido in northern Japan. The basal area decreased and the size structure of trees altered in the stands studied due to repeated, intense selective logging, in which larger trees were cut down as a priority. Sapling density in the stands was much lower than that in primary forests. In the simple and multiple regression analyses that were used to estimate the effects of selective logging on sapling density, sapling density had a significant positive correlation with tree density and had little correlation with the density of logged stumps or the height ofSasa (dwarf bamboo) growing on the forest floor. These results suggest that the establishment sites around canopy trees influenced the establishment of saplings, rather than the gaps caused by selective logging. However, both the coefficient of determination and the standardized partial regression coefficient of multiple regression analysis were higher for the stand with a dense cover ofSasa than for the stand with a sparse cover ofSasa. Thus, the success of regenerating forests with selective logging depends on both the site of advanced regeneration and the light conditions that regulate growth.     

Insect natural enemies of <Emphasis Type="Italic">Ips typographus</Emphasis> (L.) (Coleoptera,Scolytinae) in managed and unmanaged stands of mixed lowland forest in Poland

We tested the effects of forest management type (managed vs. unmanaged) on the abundances of Ips typographus (L.) (Col., Curculionidae, Scolytinae) and related parasitoids and predators reared from spruce bolts exposed in Białowieża, Poland. Habitat availability for these species differed between forest types mainly as a result of regular sanitary fellings occurring in managed forests. Populations of the common polyphagous insect predators Paromalus paralellepipedus (Herbst) and Plegaderus vulneratus (Panz.) (Col., Histeridae) or parasitoids Rhopalicus tutela (Walk.), Dinotiscus eupterus (Walk.) and Roptrocerus xylophagorum Ratz. (Hym., Pteromalidae) related to I. typographus did not differ between managed and unmanaged forests. Only Thanasimus spp. (Col., Cleridae) was significantly more abundant in managed forest, possibly because it was favored by the more open, sunny stands. However, the experiment was performed in stands where spruce is only a minor component and during the non-epidemic phase of the bark beetle population, so further experiments may be necessary to determine if the effects of sanitary fellings are greater in different forest types and at different stages of the population cycle.     

The penetration rate of laser pulses transmitted from a small-footprint airborne LiDAR: a case study in closed canopy, middle-aged pure sugi (Cryptomeria japonica D. Don) and hinoki cypress (Chamaecyparis obtusa Sieb. et Zucc.) stands in Japan

Penetration rates of laser pulses transmitted from a small-footprint airborne light detection and ranging (LiDAR) system in closed-canopy, middle-aged (40–50 years old) hinoki cypress (Chamaecyparis obtusa Sieb. et Zucc.) and sugi (Cryptomeria japonica D. Don) plantations that had similar levels of canopy openness were studied. The number of transmitted pulses was 107 427 points/ha for the hinoki cypress stand and 122 883 points/ha for the sugi stand. The penetration rates of the first (P_f) and second pulses (P_s) that reached the ground were 0.1% and 1.0% in the hinoki cypress stand, and 2.6% and 5.5% in the sugi stand, respectively. Penetration rates for the total number of pulses (P_f+s) that reached the ground within each stand were 1.1% and 8.1%, respectively, and values for P_f+s, P_f, and P_s for sugi and hinoki cypress stands were significantly different (P < 0.001). The results of this study indicated that the generation of accurate digital terrain models in dense hinoki cypress stands with complex topographies using only the data from laser pulses with poor laser penetration rates is likely to be difficult.     

Growth and nut production of black walnut in relation to site, tree type and stand conditions in south-central United States

Black walnut (Juglans nigra) is a prime tree species for agroforestry practices in the United States providing highly prized wood and nuts for human consumption and wildlife. In 54 black walnut stands in south central United States, the site index (i.e., mean dominant height [DH], at age 25 years) ranged between 5.2 and 21.4 m, and was independent of stand density. There were no differences in height and stem diameter (DBH) growth rates between stands with improved varieties and native stock. Most stands were in a “free growth” stage because of either early age or wide spacing. Mean annual increments in DBH and height were positively related both for improved varieties and native stock. Understory competition had a substantial detrimental effect on DH. In a 26 year-old stand, trees growing within Kentucky blue grass (Poa pratensis) had a site index 5 m greater than trees growing within tall fescue (Festuca arundinacea). Predicted nut yields in 2002 varied between 0 and 1370 kg of hulled nuts per ha. Improved varieties had, in general, higher nut yields than predicted from a nut yield-DBH equation developed for individual trees. Nut yields were highly variable both within and among stands, and were related to DBH in native stock but not in improved varieties.     

Individual-based measurement and analysis of root system development: case studies for <Emphasis Type="Italic">Larix gmelinii</Emphasis> trees growing on the permafrost region in Siberia

We present results of individual-based root system measurement and analysis applied for Larix gmelinii trees growing on the continuous permafrost region of central Siberia. The data of root excavation taken from the three stands were used for the analyses; young (26 years old), mature (105 years old), and uneven-aged over-mature stand (220 years old). In this article, we highlight two topics: (1) factors affecting spatio-temporal pattern of root system development, and (2) interactions between aboveground (i.e., crown) and belowground (i.e., root) competition. For the first topic, the detailed observation of lateral roots was applied to one sample tree of the overmature stand. The tree constructed a superficial (<30 cm in depth) and rather asymmetric root system, and each lateral root expanded mainly into elevated mounds rather than depressed troughs. This indicated that spatial development of an individual root system was largely affected by microtopography (i.e., earth hummocks). For these lateral roots, elongation growth curves were reconstructed using annual-ring data, and annual growth rates and patterns were compared among them. The comparison suggested that temporal root system development is associated with differences in carbon allocation among the lateral roots. For the second topic, we examined relationships between individual crown projection area (CA) and horizontal rooting area (RA) for the sample trees of each stand. RA was almost equal to CA in the young stand, while RA was much larger (three or four times) than CA in the mature and overmature stands. Two measures of stand-level space occupation, crown area index (aboveground: CAI; sum of CAs per unit land area) and rooting area index (belowground: RAI; sum of RAs), were estimated in each stand. The estimates of RAI (1.3–1.8 m² m₋₂) exceeded unity in all stands. In contrast, CAI exceeded unity (1.3 m² m₋₂) only in the young stand, and was much smaller (<0.3 m² m₋₂) in the two older stands. These between-stand differences in RAI–CAI relationships suggest that intertree competition for both aboveground and belowground spaces occurred in the young stand, but only belowground competition still occurred in the two older stands. Based on this finding, we hypothesized that competition below the ground may become predominant as a stand ages in L. gmelinii forests. Methodological limitations of our analysis are also discussed, especially for the analysis using the two indices of space occupation (CAI, RAI).     

Effect of thinning on allometry and needle-age distribution of trees in natural Abies stands of northern Japan

We tested the effects of thinning on allometry and needle-age distribution in natural stands of Abies sachalinensis Masters by comparing a thinned stand to an unthinned, control stand. Specifically, we attempted to clarify how allometry was altered after a thinning. We assumed that the needle-age distribution of trees in the thinned stand would show a younger composition than in the control stand, given the effect of improved light conditions on needle dynamics following a thinning. These investigations were conducted in dense Abies stands located in central Hokkaido, northern Japan, 19 years post-thinning. In the thinned stand, the ratio of individual needle mass to stem mass increased significantly, as compared to the control. A difference in the H–DBH relationship between the stands was probably related to this tendency. Mean needle age of trees differed significantly between the two stands, and the thinned stand showed a younger needle age than the control. Within each stand, dominant trees showed older mean needle age than codominant or suppressed trees. These tendencies may have been caused by differential needle dynamics affected by light conditions in the stands, and by different crown positions among the trees within a stand. In summary, trees in the thinned stand showed increased growth rates after thinning, which were caused by increased needle mass, younger composition of needles, and improved light conditions.     

Relationships between mass mortality of two oak species (Quercus mongolica Turcz. var.grosseserrata Rehd. et Wils. andQ. serrata Thunb.) and infestation by and reproduction ofPlatypus quercivorus (Murayama) (coleoptera: Platypodidae)

This study investigated the number of adults ofPlatypus quercivorus (Murayama) captured on host oak (Quercus spp.) trees, the attack density (the number of entry holes per 100 cm²), and the number of emerging adults to clarify the relationships between the beetle’s colonization on trees in oak stands and tree mortality. The initial attack ofP. quercivorus, which was the most intense attack, was observed on most living trees simultaneously. Although some attacked trees died within the year of the attack or in the next year, a high density of attack did not usually cause the death of host trees. Surviving trees suffered low levels of consecutive attack after the initial attack. BecauseP. quercivorus successfully produced broods only in the dead trees, the population of the insect seems to be maintained only in stands where oak mortality occurs. However, the adults that landed on most of the surviving trees appeared unable to reproduce probably due to degradation of host quality. Thus, oak mortality probably ceases within 3 or 4 years after the start of infestation in a stand, with subsequent reductions in population density of the borer.     

Regeneration growth in different light environments of mixed species, multiaged, mountainous forests of Romania

Growth of regenerating trees in different light environments was studied for the mountainous, mixed-species forests in the Carpathian Mountains of Romania. The primary species in these mixtures were silver fir (Abies alba Mill.), European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst). Seedlings/saplings of these species were selected and measured in different stands from two different geographical locations. Regenerating trees were measured for height and diameter growth during the summer of 2002. For each seedling/sapling, percentage of above canopy light (PACL) and stand basal area (BA) were used to assess available and occupied growing space respectively. Regeneration growth was compared against these two variables and regression relationships were developed. Using these models, we predicted the dynamics of regeneration as both growth and species composition. Our results showed that in low-light environments (PACL<20–35%; BA>30 m²/ha), shade tolerant fir and beech clearly outcompeted the spruce. Therefore, in dense stands, spruce could be eliminated by the shade tolerant species. For intermediate levels of cover (PACL=35–70%; BA=15–35 m²/ha) the spruce grew at comparable rates as the beech and fir. All three species showed similar growth rates in open conditions (PACL>80–90%; BA<15–20 m²/ha) with the spruce having a tendency to outgrow the others. However, in terms of establishment, such conditions favor spruce and inhibit fir and beech.     

Prescribed burning and hexazinone herbicide as release treatments in a sapling hardwood-loblolly pine stand

To compare release treatments, a randomized complete block study was established in a 7-year-old hardwood-loblolly pine (Pinus taeda L.) stand in central Louisiana established using chopping and burning. There were 5 blocks of 3 treatments each: (1) check, (2) hexazinone applied once, and (3) prescribed backfiring applied twice. The first burn in December 1985 (7 years after site preparation) had a fire intensity of 90 kJ/s/m. The hexazinone herbicide was applied in April 1986 (the 8th year after site preparation) with a metered spotgun applicator at a rate of 3.0 kg active ingredient/ha. The second burn in March 1989 (the 11th year after site preparation) had a fire intensity of 106 kJ/s/m.The two prescribed burns increased the number of stems less than 1.5 m tall from 1,380 to 2,960/ha red maple (Acer rubrum L.), blackgum (Nyssa sylvatica var. bifora), and sweetgum (Liquidambar styraciflua L.). Burning reduced the number of loblolly pines less than 2.0 m tall, which resulted in a significant increase in average loblolly pine height and diameter. Loblolly pine comprised 62 and 59% of the basal area on the check and burn treatments, respectively, 11 years after site preparation. Hexazinone reduced the number of blackgum, sweetgum, and oak (Quercus spp.) from 6,100 to 4,560 stems/ha and resulted in significantly less hardwood tree basal area than found on the check or burn treatments. Therefore, the herbicide treatment resulted in principally a loblolly pine stand (over 80% of the total tree basal area was pine) four years after hexazinone application.     

睡莲在园林中的应用及配置手法

睡莲 Nymphaea spp. 是园林水景营造中十分优秀的植物 料。介绍了水景园、睡莲专类园、盆栽 3 种睡莲的园林应用形式及各形式的应用要点,结合三潭印月的睡莲应用实例探讨其园林配置手法强调了在水面布局上需要留白以便形成倒影;阐述了不同类型睡莲的搭配,推荐了基底与重点种植、单纯片区种植 2 种种植形式,以及睡莲片区之间以不同花期、色系、开花时间的品种进行搭配的方式;还阐述了睡莲与其他植物的搭配,列举了常用搭配植物及其配置手法。最后对睡莲在园林中进一步推广提出 建议和思路。     

Composition and structure of natural mixed-oak stands in northern and central Portugal

Stand composition and structure of natural mixed-oak stands of common-oak (Quercus robur L.) and pyrenean-oak (Quercus pyrenaica Willd.) were studied. Diverse compositional and structural elements in early and late successional stand stages were analysed. The study was conducted in north and central Portugal where different natural mixed oak forests types are located. The following mixed-oak forest types involving common-oak and pyrenean-oak were studied: common-oak & other hardwoods; common-oak & cork-oak (Quercus suber L.); ash (Fraxinus angustifolia Vahl) & pyrenean-oak; and pyrenean-oak & madrone (Arbutus unedo L.). Measurements were made in early and late successional stand stages on the different mixed oak forest types. Different stand characteristics and indices were used to describe and compare stand structure and composition. The study showed changes in species diversity and stand structure. Most tree species in mature stands are present in early stages but with higher abundance. Shannon diversity index may change between 0.798 and 1.915. Significant differences on species diversity and abundance were found depending on the forest type and successional stage. Mature mixed-oak forests have high species diversity with an abundance of small to medium tree size species. Species distribution and diameter differentiation indices range from 0.30 to 0.70 and 0.52 to 0.82, respectively, revealing significant structural complexity. The average number of standing and downed dead trees was 265 and 83 trees ha⁻¹ for early and late stage, respectively, with 6.9 and 65.4 m³ ha⁻¹. Higher values of stand diversity index were 41 and 53 in more complex and developed forests. Later stand stages have complex structure, with a wider range of tree diameter distribution and higher degree of irregularity.     

Long-term effects of single prescribed fires on hardwood regeneration in oak shelterwood stands

One of the arguments against using prescribed fire to regenerate oak (Quercus spp.) forests is that the improvement in species composition of the hardwood regeneration pool is temporary and multiple burns are necessary to achieve and maintain oak dominance. To explore this concern, I re-inventoried a prescribed fire study conducted in the mid-1990s to determine the longevity of the effects of a single prescribed fire on hardwood regeneration. The initial study was conducted in three oak shelterwood stands in central Virginia, USA. In 1994, each stand was divided into four treatments (spring, summer, and winter burns and a control) and the hardwood regeneration was inventoried before the fires. During the burns, fire intensity was measured and categorized in each regeneration sampling plot. Second-year postfire data showed marked differences in species mortality rates, depending on season-of-burn and fire intensity: oak and hickory (Carya spp.) regeneration dominated areas burned by medium- to high-intensity fire during the spring and summer while yellow-poplar (Liriodendron tulipifera) and red maple (Acer rubrum) seedlings dominated unburned areas and all areas treated with low-intensity fire regardless of season-of-burn. The treatments were re-inventoried in 2006 and 2007 to determine whether these fire effects were still present. The new data show that the species distributions by season-of-burn and fire intensity found in 1996 still existed 11 years after the treatments. The fact that fire effects in oak shelterwood stands can last at least a decade has important management implications for resource professionals interested in sustaining oak forests in the eastern United States.     

Spatial structure of deciduous forest stands with contrasting human influence in northwest Spain

Five contrasting deciduous forest stands were studied to characterize the spatial structural variability in human-influenced forests. These stands are representative of cultural forest types widely represented in western Europe: one plantation, two coppices, one wood-pasture forest and one high forest stand. All stems with DBH > 5 cm were measured and mapped, and stem DBH distributions, spatial structure of DBH, spatial point patterns and spatial associations were analysed. Spatial autocorrelation for DBH was calculated with Moran’s I correlograms and semivariograms. Complete spatial randomness hypothesis for spatial point patterns, and both independence and random labelling hypotheses for spatial associations were analysed using Ripley’s K function. The results showed that tree sizes were conditioned by particular former management systems, which determined unimodal symmetric, positively skewed or compound DBH distributions. Spatial structure was more complex when human influence became reduced. Coppice stands showed clumped spatial patterns and independence among size classes, as a consequence of sexual and vegetative establishment of new stems in open areas. The largest clumping intensity was observed in the wood-pasture with an intermediate disturbance frequency and low inter-tree competition. The high forest stand displayed spatial traits consistent with the gap-dynamics paradigm, such as clumping of smaller trees, random arrangement of larger trees, negative association between juveniles and adults, and high structural heterogeneity. It can be expected that after cessation of human interference, coppices and wood-pastures would evolve to a more heterogeneous structure, probably with a higher habitat and species diversity.     

Canopy openings may prevent fir broom rust (<Emphasis Type="Italic">Melampsorella caryophyllacearum</Emphasis>) infections

Melampsorella caryophyllacearum is a heteroecious fungus that produces disease on silver fir resulting in galls that depreciate the value of trees. Currently, little is known about its epidemiology. The purpose of this study was to identify those site or management factors associated with fir broom rust infection. A 29-plot systematic sampling was carried out throughout all silver fir forests in the Spanish Pyrenees. In every sampling plot, we examined each tree for the presence of M. caryophyllacearum galls or witch’s brooms. We differentiated their position as either on the main stem or on the branches of trees. We identified the most susceptible stands by relating a compendium of tree and stand variables with the presence and severity of symptoms. Disease symptoms were observed in 11.8% (95% CI: 5.8–17.9%) of trees. Fir broom rust-infected trees tended to be those of larger diameters. The rust infection was observed in 55.2% (95% CI: 35.7–73.6%) of the stands. We identified a wide range of severities with 5 of 29 stands having more than 30% of trees infected. Fir broom rust appeared to be a widespread disease, although we observed important differences in terms of severity. The presence and severity of the rust in stands was associated with understories composed of more shade tolerant plant species and with longer periods without thinnings. Canopy opening should be experimentally tested as a possible control method.     

Bark beetle community structure under four ponderosa pine forest stand conditions in northern Arizona

We studied the bark beetle guild (Coleoptera: Scolytidae) in the ponderosa pine forests of northern Arizona to explore if the species assemblages and relative abundance differ between managed and unmanaged stands. Four stand conditions were assessed: (1) unmanaged stands with high tree density, (2) thinned stands, (3) thinned and burned (with prescribed fire) stands and (4) stands that had been burned by stand replacing wildfires. The study was conducted in the ponderosa pine forests of the Coconino Plateau, northern Arizona. For several decades this area has been relatively free of bark beetle outbreaks despite the current overstocked condition of many stands. We found that a similar species assemblage composed of Dendroctonus frontalis, D. brevicomis, D. valens, D. approximatus, D. ponderosae, and Ips pini occurred across all four stand conditions over 3 years of study. The population levels of all these species were endemic across all stand conditions. The non-aggressive D. approximatus and D. valens were indicator species for thinned and unmanaged stands, respectively, but this was not consistent among years. The ambrosia beetle Gnathotrichus sp. and the bark beetle predator Enoclerus sp. consistently indicated stands burned by wildfire. In addition to our field experiment, we analyzed the historical pattern of attacks of bark beetles in our area of study. Our findings suggest that the pattern of attack of D. brevicomis (the only Dendroctonus species for which attacks have been reported) and Ips spp. has been through scattered small infestations in groups of 1–10 trees. Whereas small infestations by Ips spp. are increasing, those for D. brevicomis are decreasing. Although we agree that the high density stands in northern Arizona are in an “unhealthy” condition, our results do not show that they were supporting large bark beetle outbreaks. Our results challenge the theoretical assumptions about the relationship between stand structure, tree resistance and bark beetle performance.     

Changes in the spatial structure of oak carbon-based secondary compounds after pine harvesting

In natural plant populations, leaf polyphenols show high intraspecific variation that occurs both temporally and spatially. Leaf phenolics may be induced by diverse ecological factors such as light, nitrogen availability or herbivory attack. Both light and nitrogen availability can show spatial structure in forested stands, meaning that they each have a high degree of autocorrelation, which can determine the appearance of spatial structure in leaf polyphenols. However, the availability of these resources may be drastically changed by forest disturbance, and little is known about the effect of forest disturbance on the spatial pattern and scale of leaf secondary compounds. We hypothesise that the spatial structure of leaf polyphenols in understory vegetation will disappear due to forest harvesting, because these compounds depend on light availability, yet it will remain unaltered for those compounds that either depend on the availability of other resources or are under major genetic control. The study was performed in young pedunculate oak (Quercus robur) populations growing either under a pine canopy (Pinus pinaster) stand or in a pine harvested stand in NW Spain. The spatial structures of green and senescent leaf polyphenols, tannins, non-tannin polyphenols and nitrogen were analysed in both stands using geostatistical analysis. The spatial structures observed for green and senescent leaf polyphenols and tannins in the forested stand disappeared in the harvested stand. However, non-tannin polyphenols, as well as nitrogen, showed spatial structure in both stands. Understanding these changes may be important for the successful recovery of native oak populations growing under pine forests in NW Spain, one of the priorities of the local government. Our results showed that changes in the concentration of leaf secondary compounds after disturbance may be accompanied by differences in their spatial properties, which may have important consequences for ecosystem function.