高黎贡山生物多样性保护与社区林业发展的研究

高黎贡山自然保护区森林资源丰富,动植物种类繁多,其物种多样性的丰富程度为国内外罕见。由于周边社区集体森林破坏严重,使保护区受到周围村社的"蚕食",给高黎贡山的森林资源管理和生物多样性保护工作带来了巨大的压力。在对村社集体林地的管理与高黎贡山生物多样性保护相关性论述的基础上,提出加强对乡村集体森林的管理,发展社区林业,实行"以林养林",即以发展集体林来保护国有林、保护区,使社区林业的发展既为当地农户提供日常生产、生活和经济发展所需要的非木材林产品、薪材及木材,又为保护区构筑一道以集体林为主的绿色屏障,以实现自然保护区森林资源管理、生物多样性保护和周围农村经济的协调发展。     

The potential value of agroforestry to dung beetle diversity in the wet tropical forests of the Pacific lowlands of Colombia

In tropical regions where agricultural activities are limited, agroforestry is an alternative for both economic development and for the management and conservation of biodiversity. The potential role of different types of land use as reservoirs of dung beetle diversity in the wet tropical forest of the Pacific lowlands of Colombia is evaluated in three agroforestry systems that differ in canopy cover and the sowing density of Borojoa patinoi crops associated with timber forest. Although total species richness was similar among land use types, differences related to the decrease in the abundance and biomass of the species were remarkable, and reflected in turn by the diversity and structure of the guilds. The general pattern observed was one in which the structure of the dung beetle assemblage of B. patinoi growing below a diversified and permanent tree cover was similar to that of the primary and secondary forest. Beetle diversity in management systems with less tree cover or a high sowing density of B. patinoi was lower and very similar to that of abandoned agricultural fields. This suggests that B. patinoi agroforestry systems can be viewed as valuable instruments for biodiversity management and conservation in the wet tropical forests of the Pacific lowlands and not just as substitutes for forest, though we must be aware that structural changes in the beetle community may in turn affect the ecological processes regulated by these insects in the agrosystems under study.     

关于发展人工林与建立人工林业问题探讨

全世界森林面积为 34 5 4× 1 0 8hm2 ,其中 3%是人工林。人工林是重要的森林生态系统 ,在木材生产、环境保护和景观建设中具有重要功能和作用。人工林在生产功能方面的补偿作用 ,使天然林可以更好地发挥生态环境价值、森林娱乐和生物多样性保护功能。人工林可持续经营的前提是遗传材料的正确选择与营林实践艺术的结合以及保证这种结合的林业政策。愈来愈多地使用外来树种营建人工林是全球趋势。人工林对于环境和生物多样性的影响 ,可以通过合理的规划和经营措施得到解决。林木育种与遗传改良和经营制度的技术创新是不断提高人工林生态系统生产力的动力。人工林的可持续经营是一个十分复杂的问题 ,中国人工林实现可持续经营面临技术和体制两方面的挑战 ,但是 ,根本出路在于林业经济体制的改革 ,象农业那样 ,在市场经济环境中实行人工林产业化经营 ,建立人工林业 (Forestplantationindustry)。     

Conservation of forest biodiversity and ecosystem properties in a pastoral landscape of the Ecuadorian Andes

High Andean cloud forests are home to a diversity of unique wildlife and are important providers of ecosystem services to people in the Andean regions. The extent of these cloud forests has been widely reduced through conversion to pasture for livestock, which threatens the forests’ ability to support biodiversity and provide ecosystem services. This paper explores whether impacts on woody plant biodiversity and four ecosystem properties (woody plant species richness, juvenile timber tree abundance, soil organic matter content and soil moisture) from converting forest to pasture can be mitigated if some woody forest vegetation is maintained within pastures. Woody vegetation in pastures was found to conserve those woody plant species that are more tolerant to exposure and grazing, but conservation of the high montane cloud forest community required areas of forest from which livestock were restricted. The sampled sites clustered according to woody plant species cover; these clusters represented a gradient from pasture with patches of shrubs to mature forest. Clusters differed in both woody plant species richness and number of juvenile timber trees whereas soil organic matter and soil moisture were observed to be similar among all clusters. This suggests that the different habitats may have some equivalent ecosystem properties. We conclude that the presence of woody vegetation in pastures may reduce some of the impacts of converting forest to pasture, but should not be considered a substitute for protecting large areas of forest, which are essential for maintaining woody plant species diversity in high Andean cloud forest.     

国内外混交林研究进展

在应对气候变化与实现碳中和目标以及社会发展对森林生态系统要求更加多样化的时代背景下,混交林的重要性在世界各国已达成共识。通常认为,相对于纯林,混交林能够增加生物多样性、提高生产力、增加碳储量,具有更高的生态韧性（抵抗力与恢复力）。但混交林是否必然优于纯林,且混交林与生物多样性、生产力、碳汇、生态韧性以及混交效应的关系一直是国内外学者探讨与研究的问题。混交林绝不是简单的树种组合与排列,其结构与生态过程相对复杂。文中通过文献梳理,总结分析混交林与生物多样性、生产力、碳储量、生态韧性以及混交林经营管理5个方面的国内外研究成果,发现关于混交林与生态功能和生产力的关系研究结果多样,总的结论是,若要得到混交林的积极效应,不能单纯增加树种多样性,更重要的是要考虑树种特性、生态互补、林分结构以及立地和环境的影响。我国混交林研究主要是通过造林实验对比分析生长效果,选择较优混交造林技术。文中最后提出我国混交林研究发展建议,旨在促进对混交林的科学和全面认识。     

Effects of Different Management Practices on Stand Composition and Species Diversity in Subtropical Forests in Nepal: Implications of Community Participation in Biodiversity Conservation

In this article, we compared the structure, composition, and diversity of trees, shrubs and saplings, seedlings and herbaceous species of community- and government-managed forests in the lowlands of eastern Nepal. Results suggest that among the trees, the community forest was dominated by a single species, Shorea robusta. However, Shorea robusta and Terminalia myriocarpa were codominant in the government forest. Tree density and basal area were higher in the government forest, but shrub/sapling density and basal area were higher in the community forest, suggesting a positive effect of community management on tree regeneration. Overstory species assemblages showed an obvious compositional difference between the forests, but understory species assemblages were less obvious. Plot-level tree and shrub/sapling species richness was higher in the government forest than the community forest. However, seedling-herbaceous species richness was higher in the community forest. The dominance of Shorea robusta trees in the community forest suggests that people involved in managing forests may be more interested in a limited number of economically valuable species while removing less important trees. Such preferential management practices may increase resource heterogeneity within a forest and maintain species diversity in the understory. Thus, community participation in forest management should be encouraged, with guided management techniques and exercises, to achieve maximum forest recovery, provide sustainable ecosystem services, and maintain forest diversity.     

An Interdisciplinary Research Initiative Towards Sustainable Management of Forest Resources in Lowland Rain Forests of Sri Lanka

Ever-wet forests of SW Sri Lanka are biodiversity hot spots with a wealth of potentially useful and currently used species. Traditional peoples surrounding these forests depend on them for a variety of focds, medicinals and the like as well as for timber. Some of these forests are being preserved as storehouses of diversity that include rare endemics and wild relatives of useful plants. Other natural forests and plantations must be managed for multiple uses including timber and non-timber products used by the forest dependent villagers. Understanding the ecological, social and economic interdependence of forests and rural communities is vital for the future forestry sector development in Sri Lanka. As many of the plants used locally are characteristic of early stages of gap regeneration, timber and non-timber products are compatible uses, given sufficiently judicious management. Anthropological and biological research is now in progress to plan for multiple use management of Sri Lankan ever-wet forests.     

Response of fungal and plant communities to management-induced overstorey changes in montane forests of the Western Carpathians

The effect of forest management on biodiversity is a crucial issue for sustainable forestry and nature conservation. However, the ways in which management affects macrofungal and plant communities and diversity of mountain temperate forests still remain poorly understood. We performed a random sampling stratified by stand age and stand type on the sites of temperate montane fir–beech forests. Diversity of macrofungi and the vascular plant understorey in beech- and spruce-dominated managed stands was investigated and compared to primeval forests located in the Po?ana Biosphere Reserve, Western Carpathians. Both the vascular plant and the macrofungal communities were altered by management, and the response of the macrofungal species (especially wood-inhabiting fungi) was more pronounced in terms of species composition change. Species turnover evaluation seems to be an important tool of forest natural status assessment, because alpha diversity did not change as much as species composition. Certain species of Carpathian primeval forests were confirmed as good indicators for natural forest change; others were proposed. Species pool and mean number of species per plot were the highest in unmanaged fir–beech forests, and species diversity significantly decreased in spruce plantations. The number of species decreased significantly due to the change of canopy tree species composition only in the macrofungal communities. As an outcome for forest management, we recommend keeping mixed forests involving all natural tree species and providing at least a minimal amount of dead wood necessary for wood-inhabiting organisms and leaving some area of unmanaged natural forests within complexes of managed stands.     

云南松天然林及人工林群落结构和物种多样性比较

通过对云南省滇中地区云南松天然林和人工林的群落结构调查及物种多样性研究,结果表明:干扰较少的云南松天然林具有较为稳定、合理的群落结构,其乔木层优势种群具有较强的自然更新能力和长远的演替潜力,并且物种多样性和生境多样性。干扰较少的云南松天然林的这些特性均优于干扰较大的云南松天然林,干扰较大的云南松天然林又优于人工林。随着森林起源过程中干扰(人为干扰和自然干扰)程度的逐渐增加,云南松群落的稳定性、生物多样性、协调平衡能力及自然修复能力相应下降;而入侵种紫茎泽兰的分布范围和数量则相应上升,提示其或可作为森林群落生物多样性、干扰程度和森林健康评价的指示生物。     

Homestead forest resources and their role in household economy: A case study in the villages of Gazipur Sadar Upazila of central Bangladesh

This study examines the species composition, diversity and economic importance of homestead forests in the household economy of central Bangladesh. The study documents 57 homestead forest species and their main usage in 90 households across three villages of Gazipur Sadar Upazila. Homestead forests in Bangladesh comprise a mixture of fruit, timber and bamboo species. While superficially homestead forestry appears unimportant in rural livelihoods, in reality the contribution is huge, both as a source of food security and for other necessary household materials. A clear understanding of the physical characteristics and economic role of homestead forests in rural livelihoods is vital for ensuring sustainable resource management. Income and production of homestead forestry on a per hectare basis are found to vary widely between landholding size classes. Significant relationships are identified between forest performance (production and income) and species richness and education level. Homestead forestry appears to be a potential subsistence income generating land-use practice in the study area. The economic scope of homestead forestry can be further enhanced provided the appropriate species composition of the forest is achieved and the education of forest owners is ensured through targeted management and policy interventions.     

Influence of tree species on understory vegetation diversity and mechanisms involved—A critical review for temperate and boreal forests

Tree species composition is a primary attribute of forest ecosystems, and is often manipulated by silvicultural practices. Forest management to diversify tree species is now being promoted to favor biodiversity. To assess the soundness of this policy we reviewed and analyzed the literature on the relationship between tree species composition and floristic diversity, including the mechanisms involved therein. Coniferous forests generally provide less diversified vascular understories than broadleaved forests. At the tree species scale, there are not enough reports to draw firm conclusions on the effect of any particular species. Mixing of deciduous and coniferous tree species generally affects understory diversity, but in almost all cases maximum diversity is observed in one of the pure stands, not in mixed stands. Understory vegetation is influenced by overstory composition and structure through modifications of resource availability (light, water and soil nutrients) and other effects, such as physical characteristics of the litter layer. Overstory light transmittance and diverse properties of forest litter are factors that have been most fully studied to date, but other factors such as throughfall water quantity and chemistry may also play a role. While the relative importance of mechanisms that account for the effect of overstory on understory biodiversity has often been discussed, these mechanisms have rarely been the subject of formal experiments. Overall, varying management practices and site attributes make it difficult to generalize results. They combine with the effects of tree species in influencing understory vegetation diversity, but they have been rarely considered. Future research is needed to gain a better understanding of the relationship between overstory and understory diversity and establish general laws.     

川中丘陵区防护林群落多样性研究

本文对川中丘陵地区主要防护林群落的物种多样性进行研究,结果表明：①该流域共有维管束植物55科,82属,88种,其科、属、种数与四川省乃至全国比较,官司河流域植物区系具有组成简单的特点;②物种丰富度,物种多样性和均匀度指数都是人工林低于天然次生林,纯林林分低于混交林分;③土壤pH值、土壤有机质、土层厚度和人为干扰是影响防护林群落多样性发育的重要制约因子,其中土壤pH值是主导因子,制约着本区防护林群浇的空间分布格局;④林木高度、郁闭度和灌、草层盖度与物种丰富度、物种多样性指数和均匀度指数相关密切,对群落多样性发育的影响较大;而林木胸径与群落多样性指数相关不明显,对群落多样发育的影响不大。     

Spatial partitioning of biomass and diversity in a lowland Bolivian forest: Linking field and remote sensing measurements

Large-scale inventories of forest biomass and structure are necessary for both understanding carbon dynamics and conserving biodiversity. High-resolution satellite imagery is starting to enable structural analysis of tropical forests over large areas, but we lack an understanding of how tropical forest biomass links to remote sensing. We quantified the spatial distribution of biomass and tree species diversity over 4 ha in a Bolivian lowland moist tropical forest, and then linked our field measurements to high-resolution Quickbird satellite imagery. Our field measurements showed that emergent and canopy dominant trees, being those directly visible from nadir remote sensors, comprised the highest diversity of tree species, represented 86% of all tree species found in our study plots, and contained the majority of forest biomass. Emergent trees obscured 1–15 trees with trunk diameters (at 1.3 m, diameter at breast height (DBH)) ≥20 cm, thus hiding 30–50% of forest biomass from nadir viewing. Allometric equations were developed to link remotely visible crown features to stand parameters, showing that the maximum tree crown length explains 50–70% of the individual tree biomass. We then developed correction equations to derive aboveground forest biomass, basal area, and tree density from tree crowns visible to nadir satellites. We applied an automated tree crown delineation procedure to a high-resolution panchromatic Quickbird image of our study area, which showed promise for identification of forest biomass at community scales, but which also highlighted the difficulties of remotely sensing forest structure at the individual tree level.     

Plant and carabid beetle species diversity in relation to forest type and structural heterogeneity

The aim of this study was to evaluate the influence of forest structure (mainly resulting from human uses) and forest type (the identity of the dominant tree species) on biodiversity. We determined the diversity of two taxonomical groups: the understory vegetation and the edaphic carabid beetle fauna. We selected eight types of forest ecosystems (five replicates or stands per forest type): pine (Pinus sylvestris) plantations of three age classes (10, 40 and 80 years since reforestation), an old-growth relict natural pine forest, and four types of oak (Quercus pyrenaica) stands: mature forests with livestock grazing and firewood extraction, mature forests where uses have been abandoned, “dehesa” ecosystems and shrubby oak ecosystems. The results obtained by a global PCA analysis indicated that both tree size and dominant species influenced the ordination of the 40 forest stands. In general, carabids were more sensitive to changes in forest heterogeneity and responded more clearly to the analysed structural variables than the understory vegetation, although the species richness of both groups was significantly correlated and higher in case of oak forests. Pine forest ecosystems were characterised by the lowest species richness for both taxonomical groups, the lowest plant diversity and by the lowest coefficients of variation and, consequently, low structural heterogeneity. As a result, it was very difficult to discriminate the effects of the spatial heterogeneity and the dominant tree species on biodiversity.     

Centuries-old logging legacy on spatial and temporal patterns in understory herb communities

Understory herb communities in the Southern Appalachians are among the highest biodiversity plant communities in North America. In the mid-1990s, a debate began over whether understory herb communities recover to their pre-disturbance states following logging. Studies showing reduced herb-layer diversity in previously logged forests were criticized for not accounting for intersite environmental heterogeneity. More recent studies have addressed environmental heterogeneity, but have neglected long-term recovery by using “mature forests” as young as 80 years old as the benchmark for diversity comparison, even though old growth stands have disturbance return intervals exceeding 500 years. Here we address concerns clouding previous studies of high-diversity Appalachian herb communities and investigate their long-term recovery by comparing paired sites of old growth forest and forest logged 100–150 years ago. We found that species richness and individual abundance is greater in old growth forests than mature forests and that species composition differed significantly between the two. Turnover in species among old growth and mature forests accounted for 11% of the total species richness and was significantly greater than expected. Species turnover at intermediate (5–50 m) and landscape-scales (>10 km) contributed the most towards total species richness. Herb communities in rich cove forests have successional trajectories that exceed 150 years, with important community changes still occurring long after the forest returns to what has been previously termed a “mature” state. To conserve the diverse herb layer, we conclude that mature forest stands are too young to serve as baselines for recovery, landscape-scale preservation of multiple forest stands is needed to maximize species richness, and maintaining 100–150-year logging rotations will likely lead to loss of biodiversity.     

Plant β-diversity in human-altered forest ecosystems: the importance of the structural,spatial, and topographical characteristics of stands in patterning plant species assemblages

An understanding of spatial patterns of plant species diversity and the factors that drive those patterns is critical for the development of appropriate biodiversity management in forest ecosystems. We studied the spatial organization of plant species in human-modified and managed oak forests (primarily, Quercus faginea) in the Central Pre-Pyrenees, Spain. To test whether plant community assemblages varied non-randomly across the spatial scales, we used multiplicative diversity partitioning based on a nested hierarchical design of three increasingly coarser spatial scales (transect, stand, region). To quantify the importance of the structural, spatial, and topographical characteristics of stands in patterning plant species assemblages and identify the determinants of plant diversity patterns, we used canonical ordination. We observed a high contribution of β-diversity to total γ-diversity and found β-diversity to be higher and α-diversity to be lower than expected by random distributions of individuals at different spatial scales. Results, however, partly depended on the weighting of rare and abundant species. Variables expressing the historical management intensities of the stand such as mean stand age, the abundance of the dominant tree species (Q. faginea), age structure of the stand, and stand size were the main factors that explained the compositional variation in plant communities. The results indicate that (1) the structural, spatial, and topographical characteristics of the forest stands have the greatest effect on diversity patterns, (2) forests in landscapes that have different land use histories are environmentally heterogeneous and, therefore, can experience high levels of compositional differentiation, even at local scales (e.g., within the same stand). Maintaining habitat heterogeneity at multiple spatial scales should be considered in the development of management plans for enhancing plant diversity and related functions in human-altered forests.     

Forest Structure Under Different Management Regimes in the Western Lowlands of Nepal

ABSTRACT

For landscape level conservation and rural sustenance, forests outside of protected areas are becoming increasingly important. Since most outside forests in the lowlands of Nepal are under government control and utilized as open access resources, their feasibility to achieve these objectives is questionable. We compared forests inside (with three decades of strict protection) and outside of protected areas (under government control and proposed as community forests) to assess the ecological sustainability of outside forests and to provide baseline data on structure of outside forests so that effectiveness of community forestry (after implementation) can be evaluated in the future. Our results indicate that outside forests were in a degraded condition with low tree, sapling and seedling densities and lower species diversity. Trees in lower dbh (diameter at breast height) classes were absent in these forests. Although the canopy layer was not significantly different, the ordination of plots and species showed that species composition of understory and the ground layer were different than protected forests. The present condition of outside forests is not only ecologically unsustainable but also cannot fulfill the demands for forest products of local people. The inefficiency of government to monitor against exploitation has been responsible for present conditions. We recommend that community forestry programs, which are successful in improving conditions of degraded forests in the mountains of Nepal, should be widely implemented in lowland areas (Terai) and proposed community forests should be handed over to local communities as soon as it is practicable.     

Lichen diversity in temperate montane forests is influenced by forest structure more than climate

Although the effect of forest management on lichens in temperate forests has been widely examined, little is known about the influence of management-related factors on their biodiversity relative to factors that cannot be altered by management. Here we determined whether forest structure or climate determines lichen diversity in the Bavarian Forest National Park in southeastern Germany, taking spatial variables into account. We investigated 517 single tree stems along 4 transects in 113 pre-stratified plots (8 m in diameter) in this montane forest. We grouped environmental variables into three sets: climate (macroclimate, non-manageable), forest structure (manageable), and space. The explanatory powers of these sets of variables for lichen diversity were compared using variance partitioning for the lichen community, species density, and threatened species density. The relationships of single characteristics of forest structure with lichen species diversity were analyzed using generalized linear models (GLM). Lichen diversity was better explained by stand structures than by climate. Spatial effects influenced the number of species per plot. Among the structural features, the availability of dead wood and sycamore maple as well as forest continuity were most important for the enhancement of lichen diversity. Open canopy structures affected the total diversity positively. Although the availability of large trees was not an influential factor in the GLM at the plot level, high diversity levels were generally associated with large stem diameters at the level of single stems. We provide recommendations for sustainable forest-management practices that aim at specifically enhancing lichen diversity in temperate areas experiencing low levels of air pollution.     

Plant diversity and its maintenance in <Emphasis Type="Italic">Populus euphratica</Emphasis> riparian forests in the Ejina Oasis,China

The dynamic environments in riparian zones support a variety of life-history strategies, which constitute a fundamental mechanism for development and maintenance of biodiversity in riparian forest ecosystems. To demonstrate the effect of life-history strategies on biodiversity, we investigated community-level diversity and its relationship to environmental variability in the riparian Populus euphratica forests of the Ejina Oasis. Communities were divided into 14 associations on the basis of their species importance values. The Simpson’s index, Shannon-Wiener index, Pielou’s evenness index and the regional Whittaker’s index were applied to calculate community diversity. An ordination of the sample plots was carried out by correspondence analysis (CA). Biodiversity was relatively low across the entire study area, but there was high community diversity (βw = 8.09) due to the spatial heterogeneity of habitats in different plots. In addition, we investigated the relationship between biodiversity and several environmental factors, such as water availability, community components and soil conditions. We conclude that the heterogeneity of soil and water availability drives community diversity patterns in riparian zones and that community-level diversity favors the maintenance of species diversity in the P. euphratica forests in the Ejina Oasis.