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1.
林隙微生境及更新研究进展   总被引:19,自引:3,他引:19  
宋新章  肖文发 《林业科学》2006,42(5):114-119
系统评述了国内外关于林隙干扰及更新的最新研究进展.林隙是一种经常发生的小规模干扰,是森林群落时空格局变化的驱动力.不同的林隙干扰状况导致了林隙内光照的差异,进一步造成林隙内温度、湿度、土壤理化性质的变化,形成了微生境的时空异质性.林隙干扰通过改变微生境而对幼苗的出土、存活、定居和生长产生重要影响.林隙内种子库的物种丰富度变化很大,是幼苗分布格局的重要影响因子.更新幼苗的存活率和生长量受林隙大小及幼苗在林隙中位置的影响.指出了这一领域今后的研究方向.  相似文献   

2.
林火干扰对长苞铁杉更新的影响   总被引:1,自引:0,他引:1  
通过模拟实验和样地调查研究了林火干扰对长苞铁杉更新的影响,结果表明:长苞铁杉可以在森林火灾迹地中完成其生活史.长苞铁杉在模拟林火干扰样地的幼苗发生率为13%,在人工刈割杂草后样地为10.5%,而在对照组样地为0,杂草的遮荫作用对长苞铁杉幼苗的发生不利.经过一个完整生长季的生长,模拟林火干扰样地内幼苗存活率为68.1%,...  相似文献   

3.
雪灾干扰下林窗对木荷幼苗更新的影响   总被引:2,自引:1,他引:1       下载免费PDF全文
林窗干扰是维持森林生态系统的重要驱动力之一,对种子萌发、幼苗等自然更新过程、森林物种组成和动态、森林生物多样性的维持具有重要作用。本研究以2008年雪灾干扰后的浙江江郎山木荷林为研究对象,对木荷林窗大小结构、幼苗更新、生长等进行调查研究,结果表明:扩展林窗以50 100 m2的林窗个数最多(占总数的45.45%),各等级林窗中以50 100 m2的林窗占总面积比例最大(占总面积的30.31%)。林窗中木荷幼苗的平均高度和地径较对照林分分别高1.44 cm和0.61 mm,幼树在林窗中的平均高度和地径则比对照林分中分别高45.37 cm和5.00 mm且差异显著;林窗大小对木荷幼苗、幼树的高度和地径生长影响显著,中林窗中幼苗的高度和地径均高于小林窗和大林窗中的幼苗且差异均显著(F=4.893,P=0.007;F=5.203,P=0.004;n=357);幼树的地径在不同大小林窗中差异显著(F=3.569,P=0.037;n=43)。林窗幼苗的更新密度随着林窗面积的增大而增大,在林窗面积达到76 m2时,更新密度达到最大值,而后随着林窗面积的增大下降;中林窗和小林窗中更新苗木以低矮植株(1级、2级)为主,面积100 m2大林窗中,木荷幼苗生长较快。与他人研究的森林天然林窗相比,雪灾干扰后改变了林窗的大小分布结构和面积,50 100 m2的林窗比例较大,一定程度上更利于幼苗更新,具有相对较大的林窗幼苗更新密度;不论林窗大小,林窗内的更新幼苗都比林内多,郁闭度较大的林内或大面积的空地上都不利于更新幼苗的生长。因此,从受灾木荷林窗大小结构、幼苗更新、生长等来看,中林窗是幼苗适宜更新的面积,为木荷灾后恢复与重建提供了科学依据。  相似文献   

4.
采集清流县大丰山长苞铁杉当年生、自然掉落的饱满种子进行育苗试验.结果表明:长苞铁杉育苗在50%日照条件下种子萌发率和幼苗成活率最高,幼株生物量最高,幼株生长最快,整个生命期生长最好;菌根接种对提高长苞铁杉种子萌发率没有明显的作用,但是菌根土育苗能显著提高幼苗的成活率,能极大提高幼苗生长量.  相似文献   

5.
We experimentally investigated interacting effects of canopy gaps, understory vegetation and leaf litter on recruitment and mortality of tree seedlings at the community level in a 20-year-old lowland forest in Costa Rica, and tested several predictions based on results of previous studies. We predicted that experimental canopy gaps would greatly enhance tree seedling recruitment, and that leaf litter removal would further enhance recruitment of small-seeded, shade-intolerant seedlings in gaps. We created a large (320–540 m2) gap in the center of 5 out of 10 40 m × 40 m experimental plots, and applied the following treatments bimonthly over a 14-month-period in a factorial, split–split plot design: clipping of understory vegetation (cut, uncut), and leaf litter manipulations (removal, addition, control). As expected, experimental gaps dramatically increased tree seedling recruitment, but gap effects varied among litter treatments. Litter addition reduced recruitment in gaps, but enhanced recruitment under intact canopy. Species composition of recruits also differed markedly between gap treatments: several small-seeded pioneer and long-lived pioneer species recruited almost exclusively in gaps. In contrast, a few medium-to-large-seeded shade-tolerant species recruited predominantly under intact canopy. Leaf litter represents a major barrier for seedling emergence and establishment of small-seeded, shade-intolerant species, but enhances emergence and establishment of large-seeded, shade-tolerant species, possibly through increased humidity and reduced detection by predators. Periodic clipping of the understory vegetation marginally reduced tree seedling mortality, but only in experimental gaps, where understory vegetation cover was greatly enhanced compared to intact canopy conditions. Successful regeneration of commercially valuable long-lived pioneer trees that dominate the forest canopy may require clear-cutting, as well as weeding and site preparation (litter removal) treatments in felling clearings. Management systems that mimic natural canopy gaps (reduced-impact selective logging) could favor the regeneration of shade-tolerant tree species, potentially accelerating convergence to old-growth forest composition. In contrast, systems that produce large canopy openings (clear-cutting) may re-initiate succession, potentially leading to less diverse but perhaps more easily managed “natural plantations” of long-lived pioneer tree species.  相似文献   

6.
Spatial variation in tree-regeneration density is attributed to the specialization of tree species to light availability for germination and growth. Light availability,in turn, varies across the gap-understorey mosaic. Canopy gaps provide an important habitat for the regeneration of tree species that would otherwise be suppressed in the understory. In subtropical forests, there is still a knowledge-gap relating to how canopy disturbances influence tree-regeneration patterns at local scale, and if they disproportionately favor regeneration of certain species. We aim to analyze whether canopy gaps promote tree regeneration, and tree species are specialized to gaps or understory for germination and growth. We sampled vegetation in 128 plots(0.01 ha), equally distributed in gaps and below canopy, in two subtropical Shorea robusta Gaertn.(Sal) forests in Nepal, recording the number of tree seedlings and saplings in each plot. We compared the regeneration density of seedlings and saplings separately between gaps and the understorey. The mean densities of seedlings and saplings were higher in the gaps at both sites;although there was no difference in the seedling density of the majority of the species between the habitats. No species were confined to either gap or understorey at the seedling stage. We conclude that gaps are not critical for the germination of tree species in Sal forests but these are an important habitat for enabling seedlings to survive into saplings. The classification of trees into regeneration guilds mainly based on germination does not apply to the majority of tree species in subtropical Sal forests. Our results reaffirm that gap creation promotes tree regeneration by favouring seedling survival and growth and can influence forest management for conservation, as well as for plantations.  相似文献   

7.
燕山山地油松人工林林隙大小对更新的影响   总被引:3,自引:0,他引:3  
林隙(gap)又称林窗,指森林群落中老龄树木死亡或偶然因素(如干旱、台风、火灾等)造成成熟阶段的优势种死亡,从而形成林冠空隙的现象(Watt,1947)。美国生态学家Runkle(1982)将林隙分为2种,一种是林冠空隙(canopy gap),指上方无树冠遮荫的部分,即太阳处于正中位置时无树冠遮荫的面积之和;另一种是扩展林隙(expanded gap),指形成该林隙周围树木基部所包围的面积(Runkle,1981)。本研究所指的林隙为应用最为广泛的扩展林隙。近年来的森林动态学研究表明:林隙在森林结构和多样性维持中起着重要的作用,已  相似文献   

8.
Virgin beech Fagus orientalis forests in northern Iran provide a unique opportunity to study the disturbance regimes of forest ecosystems without human influence. The aim of this research was to describe characteristics of natural canopy gaps and gap area fraction as an environmental influence on the success of beech seedling establishment in mature beech stands. All canopy gaps and related forest parameters were measured within three 25 ha areas within the Gorazbon compartment of the University of Tehran’s Kheyrud Experimental Forest. An average of 3 gaps/ha occurred in the forest and gap sizes ranged from 19 to 1250 m2 in size. The most frequent (58%) canopy gaps were <200 m2. In total, canopy gaps covered 9.3% of the forest area. Gaps <400 m2 in size were irregular in shape, but larger gaps did not differ significantly in shape from a circle. Most gaps (41%) were formed by a single tree-fall event and beech made up 63% of gap makers and 93% of gap fillers. Frequency and diversity of tree seedlings were not significantly correlated with gap size. The minimum gap size that contained at least one beech gap-filling sapling (<1.3 m tall) was 23.7 m2. The median gap size containing at least one beech gap-filling sapling was 206 m2 and the maximum size was 1808 m2. The management implications from our study suggest that the creation of small and medium sized gaps in mixed beech forest should mimic natural disturbance regimes and provide suitable conditions for successful beech regeneration.  相似文献   

9.
Tree seedling recruitment was monitored after various types of logging in mixed conifer and deciduous forests of northern British Columbia, Canada. Predicting tree seedling recruitment after disturbance is fundamental to understanding forest dynamics and succession and is vital for forest management purposes. Seedling recruitment success in multi-species northern latitude forests varied as a function of mature tree canopy cover, gap size and position in a gap. Recruitment was abundant within canopy gaps across a wide range of gap sizes (20–5000 m2), but recruit numbers dropped off rapidly under the closed forest canopy and in the open conditions of clearcuts. Inside canopy gaps, recruitment was similar by gap position in small gaps (<300 m2) but, in these northern latitude forests, exhibited a trend of increasing density from the sunny north to shady south end of larger gaps. This was true for all tree species regardless of their shade tolerance ranking. There was no evidence of gap partitioning by any of the tree species during the regeneration phase suggesting that adaptation to the subtleties of gap size during early recruitment are not well developed in these tree species. Favorable locations for emergence and early establishment of germinants were less favorable for growth and survival of established seedlings, i.e. the regeneration niches in these forests were discordant. Tree abundance and species diversity appears to be controlled more by differentiation among growth and survival niches than by the regeneration niches. From the perspective of forest management, abundant natural regeneration of all the dominant tree species of these mixed-species forests can be obtained after partial cutting.  相似文献   

10.
This study examined the variation in the development of naturally regenerated and planted seedlings of Sitka spruce (Picea sitchensis (Bong.) Carr.) within gaps cut in a 32-year-old stand of the same species. The circular gaps were 20 m in diameter and designed to allow sunlight into only half of the gap floor at midsummer given the latitude of 56°45′N. Eight plots (8 m × 3 m) were laid out along a north–south transect through each gap (four within the gap and two each under the closed canopy north and south of the gap). Each plot was sub-divided and seedlings were planted into one part and the other part was left to naturally regenerate. In subsequent seasons, plots were further subdivided into ‘weed free’ and ‘vegetation left untouched’. Results showed that while the two central plots within the gaps had the highest value of canopy openness, the highest accumulated temperature and lowest soil moisture were recorded in plots that received direct sunlight. However, level of germination was significantly higher in the shaded area of the gap than in the part that received direct sunshine suggesting that higher moisture levels in shaded areas are important to successful germination. Minimal germination was recorded in the plots beneath the canopy. Seedling survival was significantly influenced by the influx of competing vegetation, but only in the part of the gaps that received direct sunlight. The success of Sitka spruce regeneration within gaps appears to depend on sufficient moisture and light to support regeneration and early growth, but not too much light to encourage the development of competing vegetation. The permanently shaded areas of the gaps appeared to offer ground conditions with sufficient moisture and light to ensure successful germination and early growth of seedlings, but without excessive competition from other vegetation.  相似文献   

11.
Interactions between forest canopy characteristics and plants in the forest understory are important determinants of forest community structure and dynamics. In the highlands of southwestern, China the dwarf bamboo Bashania fangiana Yi is an understory dominant beneath a mixed canopy of the evergreen Abies faxoniana (Rheder & Wilson) and the deciduous Betula utilis (D. Don). The goal of this study was to better understand the role of bamboo dominance, canopy characteristics, and periodic bamboo dieback on forest development. To achieve this goal, we measured tree seedling, tree saplings, and trees, forest canopy characteristics, and bamboo cover in permanent forest (n = 4) and gap plots (n = 31) in a mixed A. faxoniana and B. utilis forest in Sichuan, China. Dwarf bamboos died off in 1983 in the gap plots, and in three of the four forest plots. Forest development was assessed for the period 1984–1996. The seedling bank in forest and gap plots increased after bamboo die-off. A. faxoniana seedlings increased more than B. utilis in forest plots; the opposite pattern characterized gap plots. The proportion of seedlings on raised micro-sites on the forest floor also changed and new seedling were more abundant on the forest floor. By 1996, bamboo seedling cover and biomass had recovered to ca. 45% or their pre-flowering values. Rates of bamboo seedling recovery were faster beneath canopy gaps and deciduous trees than beneath forest or evergreen trees. Tree mortality exceeded recruitment in plots with dense bamboo; the opposite pattern was found in the plot with little bamboo. The mortality rate for B. utilis trees (2.4% year−1) was higher than that for A. faxoniana (0.8% year−1) and forests with dense bamboos became more open over the census period. Tree mortality was size-dependent and intermediate sized trees had the lowest rates of mortality. Stand basal area increased mainly due to greater basal area gain than loss for A. faxoniana. Interactions between tree species life history, canopy type, and bamboo life-cycles create heterogeneous conditions that influence tree and bamboo regeneration and contribute to the coexistence of A. faxoniana and B. utilis in old-growth forests in southwestern China.  相似文献   

12.
Historical land use and management practices in the southeastern United States have resulted in the dominance of loblolly pine (Pinus taeda L.) on many upland sites that historically were occupied by longleaf pine (Pinus palustris Mill.). There is currently much interest in restoring high quality longleaf pine habitats to such areas, but managers may also desire the retention of some existing canopy trees to meet current conservation objectives. However, fast-growing natural loblolly pine regeneration may threaten the success of artificially regenerated longleaf pine seedlings. We evaluated the establishment and growth of natural loblolly pine regeneration following different levels of timber harvest using single-tree selection (Control (uncut, residual basal area ∼16 m2/ha), MedBA (residual basal area of ∼9 m2/ha), LowBA (residual basal area of ∼6 m2/ha), and Clearcut (complete canopy removal)) and to different positions within canopy gaps (approximately 2800 m2) created by patch cutting at two ecologically distinct sites within the longleaf pine range: Fort Benning, GA in the Middle Coastal Plain and Camp Lejeune, NC in the Lower Coastal Plain. The density of loblolly pine seedlings was much higher at Camp Lejeune than at Fort Benning at the end of the first growing season after harvesting. Following two growing seasons, there were no significant effects of canopy density or gap position on the density of loblolly pine seedlings at either site, but loblolly pine seedlings were taller on treatments with greater canopy removal. Prescribed fires applied following the second growing season killed 70.6% of loblolly pine seedlings at Fort Benning and 64.3% of seedlings at Camp Lejeune. Loblolly pine seedlings were generally less than 2 m tall, and completeness of the prescribed burns appeared more important for determining seedling survival than seedling size. Silvicultural treatments that include canopy removal, such as patch cutting or clearcuts, will increase loblolly pine seedling growth and shorten the window of opportunity for control with prescribed fire. Therefore, application of prescribed fire every 2-3 years will be critical for control of loblolly pine regeneration during restoration of longleaf pine in existing loblolly pine stands.  相似文献   

13.
The dynamics of a seedling bank was studied in four plots located in the Krynica Experimental Forest, southern Poland. Each sample plot was divided into eight sectors, in which germinants and seedlings were mapped, measured, and labelled; their survival was checked twice a year between 1989 and 1996. Maps of relative light intensities on the forest floor in sample plots were produced on the basis of measurements conducted in June and August 1991. During the entire period of study, European beech (Fagus sylvatica L.) made up over 96% of germinants and seedlings. Numbers of beech germinants varied strongly (from 0 to 1178) among plots and years. Each year of abundant germination was followed by one or two years of scarce germination, and there was an increasing trend in the number of germinants in three plots between 1989 and 1996. Seedling emergence was almost independent of relative light intensity and first-season survival was positively, but weakly correlated with it. Long-term survival of seedlings was low and strongly related to relative illumination. Results of analyses of spatial patterns in seedlings suggested that some microsites were more conducive than others to germination and early survival of seedlings. Survival among taller germinants was greater than among shorter ones. A true seedling bank was observed only in two plots, where relative light intensities were 9 and 15% of full sunlight. In a plot with relative light intensity of 4%, the seedling bank was unstable and in the least illuminated plot (<3% of full sunlight) no seedling bank was formed. Results indicate that, even in a strongly shade-tolerant species like beech, formation of a seedling bank requires the occurrence of canopy openings in the close proximity.  相似文献   

14.
  • ? In mixed-species forest stands, large losses in tree species diversity often occur during the regeneration phase. In a former coppice-with-standards, we investigated whether the limiting stage in the recruitment process of advance regeneration is the immediate seedling response to canopy release. Experimental canopy gaps were opened and the survival and growth of advance seedlings (Fagus sylvatica, Acer pseudoplatanus, Acer campestre, Acer platanoides) growing in the gaps or under closed canopy were monitored for three years.
  • ? All species responded positively and rapidly to canopy release. Survival was not affected by gap opening. Diameter increment after gap opening was similar across species, and height increment was greater for Acer platanoides and for Acer pseudoplatanus. Post-release diameter and height growth were mainly determined by pre-release seedling size. Competition from neighbouring seedlings did not affect growth in the three years following canopy opening.
  • ? In the recruitment process of F. sylvatica and Acer sp. advance regeneration, the recovery from canopy release did not appear as a limiting step that would filter against some species. Pre-release size was the main factor accounting for post-release growth and is probably a major determinant of long-term seedling dominance.
  •   相似文献   

    15.
    Eastern white pine (Pinus strobus L.) is a moderately shade-tolerant species that co-occurs with hardwood tree species in many forests of the eastern United States, as well as in pure stands. The species is valued for its timber, as well as for wildlife and recreation. Regeneration of this species is somewhat unpredictable and often occurs in patches of similarly-aged cohorts. We described the regeneration patterns of this species and examined their relation to environmental variables within hardwood forests of southwestern Virginia, USA. An average of 5.3 white pine patches per ha were observed in this study. The majority of patches consisted of saplings (85%), with 9% of patches in pole size classes, and 6% in seedling size classes. The average density of patches was 43.5 stems with an average age of 20 years. The size of patches averaged 80.6 m2. The total density of seedlings and the number of regeneration patches of all sizes of regeneration (seedlings, saplings, and poles) in plots was related to the surrounding density of large white pine trees (potential seed trees). The density of seedlings or patches was not significantly related to current vegetation cover or soil surface cover variables, but more than half of regeneration patches were located in or adjacent to old canopy gaps, most of which were old logging gaps. While seedling regeneration may occur within the understory of these forests near seed trees, advancement to the sapling and pole stage appears to be associated with canopy gap formation.  相似文献   

    16.
    为了阐明华北落叶松人工林不同林隙大小对更新的影响规律,对木兰围场低效华北落叶松人工林林隙内苗木更新特性进行了研究。结果表明:林隙内更新苗的主要种类有华北落叶松、山杨和油松。华北落叶松和山杨的密度在林隙面积为120~140m2时最大,达到2 411株/hm2;而油松的密度在林隙面积为20~40m2时最大,为196株/hm2。随着林隙面积的增大,华北落叶松和山杨大径级和大高度级更新幼苗数量基本呈现增加的趋势,在林隙面积达到160~180m2时达到最大值;而油松在40~60m2时达到最大,径级大于2cm的更新苗密度为69株/hm2。  相似文献   

    17.
    Successful regeneration of northern red oak (Quercus rubra L.) on productive sites is problematic in eastern North American forests. Natural and artificial regeneration often cannot compete with fast-growing, shade intolerant species such as yellow-poplar (Liriodendron tulipifera L.). This study examines 5-year survival, growth, and competitive ability of planted northern red oak seedlings in various group selection harvest sizes in south-central Indiana, USA. Seedling stocktypes consisted of high (BHD; 75 seedlings m?2) and low (BLD; 21 seedlings m?2) nursery-bed-density bareroot seedlings, and small (CS; 11.4 L) and large (CL; 18.9 L) container seedlings. Group selection openings included large (0.400 ha), medium (0.100 ha), and small (0.024 ha) circular gaps in four stands. Larger stocktypes and gap sizes improved seedling height, diameter, and growth; ANOVA indicated only gap size was significant for seedling survival. Logistic regression showed survival was positively correlated to diameter at year 1, and aspect, gap size, and stocktype were significant predictors of survival. Our data indicated no differences in density of natural regeneration among gap sizes, although trends suggest greater numbers of bigger competitors in larger gaps sizes. Yellow-poplar regeneration was the tallest competitor of more than 50% of all northern red oak seedlings. Competitive status of seedlings after 5 years differed only by stocktype, with large container stock in a better competitive position than bareroot stock. However, less than 20% of seedlings in all stocktypes in each gap treatment were considered competitive (i.e., ≥80% of the height of tallest competitor) against their tallest competitor. The use of larger planting stock may offer greater opportunities for successfully regenerating northern red oak seedlings on productive sites but likely would have to be accompanied by treatments to reduce woody competition.  相似文献   

    18.
    海南霸王岭热带山地雨林林隙幼苗库动态规律研究   总被引:18,自引:1,他引:18  
    分析了海南岛霸王岭自然保护区热带山地雨林林隙内树种的种类组成、数量动态以及林隙年龄和林隙面积对树种组成及数量的影响。结果表明 :热带山地雨林林隙树种组成十分复杂 ,数量分布极不均匀 ,种间差异较大 ,2 7个林隙中共有乔木 116种 ,其中数量较多的 10个种的个体数量占到总个体数量的 4 7 0 3% ,而个体数量较少的 5 8个种的个体数仅占总个体数的 6 0 2 %。林隙面积大小对树种幼苗的种类组成和数量有较大影响 ,幼苗种密度和个体密度随林隙面积的增大而减小 ;幼树和成树在不同大小林隙中的种密度和个体密度变化不大。成熟林隙明显比早期林隙和中等年龄林隙有较大的幼苗个体密度和种密度 ,不同年龄林隙内幼树和成树的个体密度变化也不大。幼苗向幼树的转化率在大林隙和成熟林隙内明显高于小林隙和幼龄林隙。依据林隙树木幼苗数量变化的差异 ,可将林隙填充者分为 4种不同类型。  相似文献   

    19.
    The objective of the study was to examine the success of regeneration in gaps of variable size in pine-dominated stands in terms of seedling density. It was based on an experiment in central Lapland containing circular clear-cut gaps of 20, 40 and 80 m in diameter on typical sub-xeric and xeric pine sites with site preparation (patch scarification). The observation period covered the first five years after cutting. The average number of seedlings that had emerged after gap cutting was about 22 000?ha?1 for pine and 7 000?ha?1 for birch. The proportion of regeneration sample plots without any pine seedlings was less than 10%. The seedling density diminished constantly with greater distance from the edge stand but indicating sufficient density up to the largest gap size in the study (diameter 80 m, area ca. 0.5?ha). Site preparation promoted regeneration remarkably, and we conclude that successful regeneration would be achieved in most cases with soil scarification exposing just 10–20% of the soil surface.  相似文献   

    20.
    油茶大规格容器苗质量及其造林效果评价   总被引:1,自引:0,他引:1  
    为了筛选出适宜的容器规格、育苗基质及造林苗木规格,从而为油茶育苗和造林质量的提高提供技术依据,采用芽苗砧嫁接容器育苗方法,就不同规格的容器与不同类型的基质对容器苗质量的影响情况进行了试验研究,并对采用大规格容器苗的造林效果进行了调查与评价。试验结果表明:以大规格容器和轻基质培育的苗木其生长状况与根系质量均良好;运用3年生容器大苗造林的成活率平均为96.32%,其树高1.74 m、地径34.45 mm、冠幅1.07 m^2,当年挂果率达96.67%。研究结果表明:油茶大容器轻基质育苗和造林具有生产可行性,但相关技术还有待进一步优化与推广。  相似文献   

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