Higher breeding densities of the threatened little bustard Tetrax tetrax occur in larger grassland fields: Implications for conservation

Previous studies have found that densities of little bustard Tetrax tetrax breeding males tend to be higher in areas with smaller agricultural fields, presumably due to increased habitat diversity. However, exceptionally high densities have been found in large grassland fields in Portugal, which suggests that the influence of field size varies geographically, and that the role of this factor is not yet fully understood, despite its importance as a key management issue.We studied how field size, together with vegetation structure, influences the presence and density of breeding little bustards in a region of southern Portugal. Fifty-four grassland fields were sampled in 2007 and another 29 in 2008, with sizes ranging from 23 to 172 ha. A total of 183 breeding males were found in 47 of these fields, reaching densities of up to 37 males/100 ha. A higher probability of occurrence of breeding males was found in larger fields with a vegetation height below 40 cm and field size alone explained 46% of the variability in male density. These results suggest that larger continuous areas of suitable habitat attract many males, most likely as a consequence of their lek mating system. We conclude that conservation efforts, in a landscape context of large farm sizes, should: (1) be channelled to farms with large fields; (2) ensure adequate livestock grazing to create suitable habitat and (3) promote management at a landscape level to ensure the most continuous grassland habitat patches possible.     

The importance of traditional meadow management techniques for conservation of vascular plant species richness in Norway

Using information from a regional survey of vascular plants of 130 sites in western Norway, a selection of sites based on a heuristic iterative complementarity-based nature reserve selection procedure was performed. The results indicate that conservation of traditionally managed hay meadows is of major importance as they contributed 60.1% of all native species recorded; afforested grasslands (deciduous woodlands < 70 years old) contributed 26.8%, whereas artificially fertilized hay meadows and intensively cultivated grasslands taken together contributed 13.1% of the species. The species composition of the meadows was significantly nested. Thus, if you conserve the most species-rich meadows, you also conserve most of the species in the less species-rich meadows. Nestedness in meadows was significantly correlated with within-meadow habitat diversity and soil pH. The most species-rich meadows were traditional meadows, characterized by high habitat diversity and high soil pH. These meadows will support nearly all species including habitat specialists and regionally rare species, whilst artificially fertilized hay meadows only support the generalist subset, i.e. common species. Area was not significantly correlated with nestedness suggesting that it is more important to cover many habitats than to preserve large traditional meadows just because they are large.     

Factors relating to soil fertility and species diversity in both semi-natural and created meadows in the West Midlands of England

The post‐war decline in the area and diversity of neutral meadows in Britain, resulting from agricultural intensification, has prompted schemes to restore and create new habitats. Their success relies on understanding the relations between soil fertility and species diversity. We have investigated these relations, using multivariate analysis, in 28 semi‐natural meadows and eight artificially created urban meadows. Mineralizable nitrogen was the most important soil characteristic in the semi‐natural sites; the more N the soil contained the fewer were the species characteristic of traditional meadows. Both potassium and total magnesium favoured diversity, perhaps because their deficiency in many traditionally managed meadows jeopardizes the survival of broad‐leaved species in competition with grasses. Available lead, at sub‐lethal concentrations and measured as a Pb:Ca ratio, appeared to favour diversity in the semi‐natural sites, possibly by inhibiting the uptake of P by competitive grasses and allowing the less competitive species associated with diversity to flourish. The main differences between the soils of the artificial and the semi‐natural meadows were that the former contained more extractable P and less mineralizable N and organic matter. It seems that large soil phosphorus concentrations may be the main reason why relatively few species colonize or survive in grassland on many urban soils.     

Response of microbial community composition and activity in agricultural and grassland soils after a simulated rainfall

Rainfall in Mediterranean climates may affect soil microbial processes and communities differently in agricultural vs. grassland soils. We explored the hypothesis that land use intensification decreases the resistance of microbial community composition and activity to perturbation. Soil carbon (C) and nitrogen (N) dynamics and microbial responses to a simulated Spring rainfall were measured in grassland and agricultural ecosystems. The California ecosystems consisted of two paired sets: annual vegetable crops and annual grassland in Salinas Valley, and perennial grass agriculture and native perennial grassland in Carmel Valley. Soil types of the respective ecosystem pairs were derived from granitic parent material and had sandy loam textures. Intact cores (30 cm deep) were collected in March 1999. After equilibration, dry soil cores (approx. −1 to −2 MPa) were exposed to a simulated Spring rainfall of 2.4 cm, and then were measured at 0, 6, 24, and 120 h after rewetting. Microbial biomass C (MBC) and inorganic N did not respond to rewetting. N₂O and CO₂ efflux and respiration increased after rewetting in all soils, with larger responses in the grassland than in the agricultural soils. Phospholipid fatty acid (PLFA) profiles indicated that changes in microbial community composition after rewetting were most pronounced in intensive vegetable production, followed by the relict perennial grassland. Changes in specific PLFA markers were not consistent across all sites. There were more similarities among microbial groups associated with PLFA markers in agricultural ecosystems than grassland ecosystems. Differences in responses of microbial communities may be related to the different plant species composition of the grasslands. Agricultural intensification appeared to decrease microbial diversity, as estimated from numbers of individual PLFA identified for each ecosystem, and reduce resistance to change in microbial community composition after rewetting. In the agricultural systems, reductions in both the measures of microbial diversity and the resistance of the microbial community composition to change after a perturbation were associated with lower ecosystem function, i.e. lower microbial responses to increased moisture availability.     

The influence of plant diversity on people’s perception and aesthetic appreciation of grassland vegetation

The conservation of biodiversity critically depends on the value that humans attach to it. Apart from an ecological and economic value, an aesthetic value has often been assigned to biodiversity. However, it is not known whether lay people appreciate the diversity of species and not just certain individual species or nature as a whole. We studied in a series of experiments and field studies people’s perception and appreciation of species diversity. We presented meadow-like arrays of different species richness and evenness but random species composition to lay people and asked them to estimate plant species richness and rank the arrays by attractiveness. The experiments were complemented by two larger-scale field studies using natural meadows. Both in the experiments and the field studies the mean perception of species richness by people increased with true species richness, but was slightly overestimated at low and increasingly underestimated at high diversity levels. Lay people’s aesthetic appreciation of both the experimental grassland arrays and the natural meadows increased with true species richness. Communities consisting of the same number of species were perceived to be more species-rich and were appreciated more when their evenness was high. Our results demonstrate that plant diversity in itself is attractive to humans. The current reduction of the diversity of grasslands due to intensification of management may thus reduce the attractiveness of regions where grasslands are a dominant feature of the landscape. This could have negative consequences for tourism and may add an economic argument for the conservation of biodiversity in grasslands.     

Trends in organic carbon and nitrogen contents in agricultural soils in Bavaria (south Germany) between 1986 and 2007

In the last 60 years traditional agriculture in industrialized European countries, which had initially been dependent on available natural resources, has shifted towards a massive intensification of nutrient turnover because of cheap energy and low‐cost synthetic fertilizers. At the same time farm structure has undergone profound changes, resulting in an increase in the number of specialized farms to the detriment of traditional non‐specialized ones. All these trends have had a significant impact on agricultural management. The intensification of agricultural management together with climate change could affect the quantity and quality of soil organic matter (SOM). That could imply decreasing soil fertility, reduced harvest yields, increasing nutrient losses and additional greenhouse gas emission. In order to measure the long‐term development of SOM in agricultural soils a monitoring programme was initiated in Bavaria in 1986. The measurements are based on 92 representative plots located on cropland and 21 plots located on managed permanent grassland. Between 1986 and 2007 the monitoring plots have been sampled four times. The monitoring results suggest a decrease of soil organic carbon content, total nitrogen content and C:N ratio in cropland as well as in grassland in Bavaria between 1986 and 2007. Crops and organic fertilizers are together with the initial SOM content the main causes of the observed changes in SOM quantity and quality. A climatic effect could be neither proved nor excluded. The results in Bavaria are consistent with the reported changes in organic carbon of agricultural soils in Austria, Belgium, France, the Netherlands and England. In Bavaria we should expect declining SOM stocks, particularly soil organic carbon, in agricultural soils if the supply of organic matter remains constant or even decreases.     

The impact of organic and inorganic fertilizers and lime on the species-richness and plant functional characteristics of hay meadow communities

Vegetation responses to fertilizers and lime applied in a seven-year study at paired unimproved and semi-improved mesotrophic hay meadows in Cumbria and Monmouthshire, UK, are described in terms of species-richness and several other plant community variables. Treatments were farmyard manure (FYM) applied at rates of between 6 and 24 tonne ha⁻¹ annually or every third year, inorganic fertilizers giving equivalent amounts of N, P and K, and lime applied either alone or with FYM applied annually or three-yearly. Annual FYM at 24 tonne ha⁻¹ reduced species-richness and the richness of positive indicator species at all sites and increased the proportional cover of nutrient-demanding species, whilst liming in conjunction with 12 tonne ha⁻¹ annual FYM application was equally detrimental at the unimproved site in Wales but not in Cumbria. Inorganic fertilizers were apparently no more detrimental to vegetation quality than equivalent FYM treatments, although medium-term nutrient supply from FYM may have been underestimated. Species-richness was maintained by FYM at ?12 tonne ha⁻¹ year⁻¹ in the semi-natural northern meadow, where such levels had been used in the past, but only by amounts equivalent to ?6 tonne ha⁻¹ year⁻¹ at the Welsh sites which had no recent history of fertilizer use. It is unclear to what extent such differences are attributable to innate differences in plant community type, rather than to site-specific differences in past management. The implications of our findings for defining sustainable fertilizer practices to maintain or enhance the nature conservation value of mesotrophic meadows are discussed.     

Phosphorus availability and farm structural factors: examining scarcity and oversupply in north‐east Germany

Assessing factors influencing phosphorus (P) availability in soils is important in preventing its overexploitation and excessive application in agricultural systems. Despite high historical P applications in the federal state of Brandenburg (Germany), county data on soil test P (STP) reveal considerable disparities in soil available P. In addition, negative soil balances as a result of small mineral P and manure inputs have been observed, raising questions about the factors leading to this situation. Our work focused on identifying possible causes operating at the farm management level by conducting a letter survey in two administrative districts of Brandenburg, the counties Barnim and Uckermark, linking farm management factors (ownership type, farm size, land tenure, animal husbandry with or without grassland and its intensity, presence of a biogas plant and organic production) to farmers’ self‐indicated levels of STP. Small‐ to medium‐sized individual farms tended to have (very) high STP, while large partnership farms and companies/cooperatives were sensitive to factors resulting in low STP. Farms with low shares of land ownership, the presence of grassland, extensive cattle farming and stockless organic farming had lower STP. On the other hand, biogas plants, partly in combination with intensive livestock (cattle) farming, were associated with larger STP. It was concluded that more care should be devoted to the design of agricultural policies and that further (inter‐ and transdisciplinary) research on this topic is needed.     

Impact of grassland farming intensification on the breeding ecology of an indicator insectivorous passerine, the Whinchat Saxicola rubetra: Lessons for overall Alpine meadowland management

The decline of insectivorous farmland passerines has been attributed mostly to global decrease in arthropod availability, as a result of intensification of agricultural practices. The diminution of the Alpine Whinchat, once a widespread insectivorous passerine, has been ascribed to nest losses due to earlier and more frequent mowings. However, potential conjugated effects of deteriorated arthropod food availability had yet to be investigated. We compared food supply and nestling diet in intensively vs. traditionally managed grassland. Abundance and diversity of arthropods were much lower in intensive areas, where small-sized invertebrates, which do not enter nestling diet, were also predominant. Parents breeding in intensive habitats fed less biomass to nestlings than adults from traditional habitats. Nestling diet was less diverse and dominated by less profitable prey items in intensive than in traditional habitats. Feeding rate did not differ between the two habitats, but foraging distances from nest tended to be greater in intensive farmland. There were no significant differences in clutch sizes and hatching success with respect to management intensity, but fledging success was higher in traditional habitats. The recent intensification of farming practices has led to a decrease in the availability of grassland invertebrates, and of important Whinchat nestling food in particular, affecting parents’ foraging efficiency and reproductive success. Conservation actions must not only reduce nest losses by postponing mowing, but should also promote grassland farming that is less detrimental to invertebrates. Dominant at the study site, organic grassland farming does seemingly not provide sufficient conditions for Alpine Whinchats.     

Effects of ecological compensation meadows on arthropod diversity in adjacent intensively managed grassland

An important goal of ecological compensation areas (ECAs) is to increase biodiversity in adjacent intensively managed farmland and the agricultural landscape at large. We tested whether this goal can be achieved in the case of the agri-environmental restoration scheme implemented for Swiss grassland using five large arthropod taxa (bees, true bugs, orthopterans, ground beetles and spiders) representing different ecological and functional groups. The species richness and abundance of all groups and species, respectively, was measured along 100 m transects from ECA-meadows into the adjacent intensively managed grassland at 24 sites. Species richness of all arthropod taxa except ground beetles, and the abundance of 63% of the 234 arthropod species sampled with at least five individuals were higher in ECA-meadows than in their surroundings, while the total abundance of spiders and ground beetles was higher in intensively managed meadows. The abundance of 8% of these species were only increased in the ECA-meadows themselves (“stenotopic” species) but 40% had increased abundance both in the ECA-meadows and the adjacent grassland, declining exponentially with increasing distance from ECA-meadows (“edge species”). The 90%-decay distances for these edge species differed among taxonomic groups (117 ± 18 m for true bugs, 137 ± 24 m for spiders, 152 ± 34 m for bees, 167 ± 5.7 m for orthopterans, 185 ± 34 m for ground beetles; mean ±1 standard error) and independent of taxonomic group were larger for large-sized or predacious species than for small-sized or phytophagous species. Because the average distance between neighbouring ECA-meadows in Swiss grassland is only 73 ± 4 m, the current agri-environment scheme very likely enhances arthropod diversity and possibly associated ecosystem services in the Swiss agricultural landscape at large.     

Small-scale plant species richness and evenness in semi-natural grasslands respond differently to habitat fragmentation

The study explores whether small-scale species diversity, species evenness and species richness in semi-natural grassland communities are similarly associated with present management regime and/or present and historical landscape context (percentage of different land-cover types in the surroundings). Species diversity, evenness and richness were recorded within 441 50 × 50 cm grassland plots in 4.5 × 4.5 km agricultural landscape on Öland, Sweden. Recent and historical land-cover maps (years 2004, 1959, 1938, 1835, and 1800) were used to characterize the present and past landscape context of the sampled vegetation plots. Partial regression and simultaneous autoregressive models were used to explore the relationships between species diversity measures (Shannon diversity, richness and evenness) and different explanatory variables while accounting for spatial autocorrelation in the data. The results indicated that species richness was relatively sensitive to grassland isolation, while the response of species evenness to isolation was characterized by a degree of inertia. Because the richness and evenness components of species diversity may respond differently to habitat fragmentation, we suggest that monitoring projects and empirical studies that focus on changes in biodiversity in semi-natural grasslands should include the assessment of species evenness - as a complement to the assessment of species richness. In addition, our results indicated that the development and persistence of a species-rich and even grassland vegetation was favoured in areas that have historically (in the 19th century) been surrounded by grasslands. Information on landscape history should, whenever possible, be incorporated into the planning of strategies for grassland conservation.     

元阳梯田不同土地利用类型表层土壤的抗冲性

为了研究云南省元阳县梯田核心区土壤的侵蚀特性和保水机制,采用原状土冲刷槽法和实地放水冲刷试验法,结合有林地、灌木林地、荒草地和坡耕地的物理性质等,对元阳梯田核心区上述4种土地利用类型表层土壤抗冲性及其主要影响因素进行研究。结果表明:研究区表层土壤抗冲性由强到弱表现为荒草地>灌木林地>有林地>坡耕地;含沙量随着坡度(15°、30°和45°)和冲刷流量(1.5、2.5和3.0 L/min)的增加而增加,坡耕地具有较弱的抗冲性;土壤中的须根能够增强土壤的抗冲能力,特别是直径<1.0 mm的须根具有较强的固结性;土壤的水稳性团聚体越多土壤的抗冲性也就越好,荒草地和灌木林地表层土壤直径>3 mm的土粒,其结构水稳度为93.33%。     

A comparison of runoff and erosion rates on bare and grassed loamy sand soils

Abstract. Runoff and erosion rates on loamy sand soils were measured at the Hilton experimental site, Shropshire. Little runoff and no erosion occurred on a steep runoff plot inserted into permanent grassland. Most precipitation infiltrated into the soil and was transmitted down slope as interflow within the Ah horizon. In contrast, considerable amounts of runoff and erosion occurred on the bare arable plots. These results suggest that incorporation of strips of permanent grassland into arable systems could assist soil and water conservation. Considering the high erosion rates on bare steep plots, conversion of slopes > 12° to permanent grassland seems appropriate.     

Mycorrhizae activity and diversity in conventional and organic apple orchards from Brazil

The aim of this study was to characterize the impacts of a conventional, an organic apple orchard and a native grassland on the activity and diversity of arbuscular mycorrhizal fungi (AMF) located at the south of Brazil during winter and summer. AMF activity was measured by the mycorrhizal inoculum potential (MIP), mycorrhizal fungal hyphal length (HL), easily extractable and total Bradford-reactive soil protein (BRSP). AMF diversity was represented by richness, Shannon diversity index and number of spores. Orchards reduced the MIP of the soil and increased the HL when compared to the grassland site. The amount of easily extractable BRSP was not different among orchards and between seasons evaluated, with overall mean value of 1.23 mg g⁻¹. However, the amount of total-BRSP was smaller in the conventional orchard (4.55 mg g⁻¹) than in the organic orchard (4.91 mg g⁻¹) and in the native grassland (5.12 mg g⁻¹). T-BRSP and total organic carbon were strongly correlated in the grassland during the winter, suggesting the contribution of this protein for carbon stocks in the native soil of this region. The organic orchard presented the highest AMF richness, but sporulation and Shannon diversity index were larger in the conventional orchard. Our data suggest that the conventional orchard promoted higher impacts on the natural condition of AMF activity, being considered an unadvisable practice to soil conservation.     

Temporal change in land use and its relationship to slope degree and soil type in a small catchment on the Loess Plateau of China

We sought to detect the temporal change (1958–1999) in land use patterns and its relationship to physical landscape parameters in a small catchment in the semi-arid hilly area of the Chinese Loess Plateau. Degree of slope and soil type were selected as stable discriminating parameters that might constrain land use. With the help of GIS and canonical correspondence analysis, the relationship between rural land use distribution or transformation and the selected physical parameters was examined. The land use had undergone a general shift from farmland to woodland or grassland. Canonical correspondence analyses (CCA) indicated that a relationship between land use and the selected physical parameters was evident, farmland coincided with favorable conditions of fertile soil and gentle slope, while grassland and woodland were associated with conditions of poorer soil and steep slope. In the more favorable conditions the main land use change process was the intensification of farming, while in the less favourable conditions it was the abandonment of farmland. A thorough understanding of the relationship between land use temporal or spatial patterns and landscape physical parameters in the Loess Plateau of China, like degree of slope and soil type, will enhance our capability to predict landscape dynamics and lead to more sound and effective land use management strategies.     

高寒草甸区高原鼢鼠新生土丘水土流失特征

高原鼢鼠是青藏高原的优势地下啮齿动物,掘洞造丘是其独特的行为之一。高原鼢鼠新生土丘属于次生裸地容易导致水土流失,研究高原鼢鼠新生土丘水土流失对于合理评价其对生态系统的影响具有重要意义。利用径流小区法和风蚀桥法,测定不同地形(平地和坡地)、不同直径大小(大土丘:直径75 cm,中等土丘:直径50～75 cm,小土丘:直径50 cm)单位面积新生土丘年土壤流失量和有机质流失量,并比较经过1年侵蚀后的土丘与无土丘草地表层(0—10 cm)土壤粒径组成、含水量和营养成分,旨在明确高原鼢鼠不同大小和不同分布的土丘的水土流失状况。结果表明:(1)研究区平地和坡地新生土丘单位面积年均土壤流失量分别为4 039.91,3 731.79 g/m~2,单位面积年均土壤有机质流失量分别为266.83,252.76 mg/m~2,土壤侵蚀级别为中度侵蚀;(2)土丘表层土壤粒径0.25 mm和5 mm的团聚体含量均显著小于无土丘草地(P0.05),0.25 mm的团聚体含量显著大于无土丘草地(P0.05);(3)土丘表层土壤肥力和土壤含水量均显著小于无土丘草地表层土壤(P0.05);(4)雨季坡地新生土丘单位面积泥沙流失量显著高于平地新生土丘(P0.05)。风季平地风蚀厚度大于坡地风蚀厚度,分别为4.33,3.62 cm;(5)风季不同丘型土丘的侵蚀厚度表现为大土丘小土丘中土丘,分别为5.50,3.59,2.82 cm。研究认为高原鼢鼠新生土丘会导致高寒草甸风蚀和水蚀发生,且不同地形和不同大小的土丘水土流失情况各异。     

Effects of abandonment of subalpine hay meadows on plant and invertebrate diversity in Transylvania, Romania

Recent changes in agriculture (intensification or abandonment) have resulted in a critical reduction of semi-natural grasslands in Eastern Europe. Subalpine semi-natural grasslands in Transylvania, Romania, harbour a high diversity of plants and invertebrates, including endemics, and are considered refugia for numerous threatened open-land species. We investigated effects of land abandonment by examining species richness, species abundance, proportion of open-land, endemic and threatened vascular plants, gastropods, and diurnal and nocturnal Lepidoptera in extensive hay meadows (initial stage), and three seral stages of succession (early stage of abandoned hay meadow, naturally growing birch forest, and mature forest) in the mountainous region of Baisoara in Transylvania. A total of 626 species (225 vascular plants, 16 gastropods, 68 diurnal and 317 nocturnal Lepidoptera) were found in the 16 study sites (four replicates per successional stage). The four taxonomic groups differed in their response to the abandonment of hay meadows. Each stage of succession harboured the maximum species richness for one taxonomic group: extensive hay meadows for vascular plants, abandoned hay meadows for diurnal Lepidoptera, birch forests for nocturnal Lepidoptera, and mature forests for gastropods. In all four taxonomic groups the complementarity of species composition increased with successional age, whereas the number of characteristic open-land species decreased with successional age. The four successional stages did not differ in proportion of red-listed plant and diurnal Lepidoptera species. In nocturnal Lepidoptera, however, the proportion of red-listed species increased with successional age. Furthermore, successional stages did not differ in number of plant species endemic to the Carpathians and Eastern Europe. Our results indicate the high conservation value of all stages of subalpine grassland succession for the indigenous biodiversity of Transylvania. To prevent losses of characteristic species, we suggest a rotational grassland management program that maintains different successional stages.     

Earthworm communities in grasslands and horticultural soils

 In a survey of 42 farm sites, comprising grassland and two types of horticultural farms (growing vegetables or flower bulbs), earthworm communities were sampled by hand-sorting and a number of soil physico-chemical characteristics recorded. For heavy metals the availability in the soil solution was estimated based on the measured absolute content. Abundance, biomass and species richness were significantly higher in grassland soils than in horticultural soils, and within the horticultural farms significantly higher in vegetable than in flower-bulb farms. No epigeic species were found in horticultural soils. The differences between the various farm types were probably related to the intensity of management practices, such as soil tillage, harvesting and crop protection measures, that result in less soil organic matter of lower quality. The data from the grassland farms were analyzed in conjunction with the physico-chemical soil characteristics, applying canonical correspondence analysis. Community parameters such as species richness and diversity apparently were not related to soil characteristics, but an appreciable part of the variation in abundance and biomass of several individual species could be explained by factors such as lutum content (particles <16 μm), pH, bio-availability of metal ions and geographical position of the sample site. It is concluded that repeated surveys of this type may become a valuable tool for monitoring soil quality. Received: 16 March 2000     

The effectiveness of conservation measures to enhance nest survival in a meadow bird suffering from anthropogenic nest loss

The main reason for meadow bird declines is supposed to be the intensification of grassland management with earlier first harvest dates and more frequent harvests, resulting in high nest destruction rates. To increase productivity of meadow bird populations in intensified grassland areas a delay of mowing date and individual nest protection measures have been proposed. However, for ground-nesting songbirds such as the whinchat (Saxicola rubetra) the effectiveness of such measures remains widely unknown. In particular, if nest predation rate is high, measures to protect nests from agricultural destruction alone might be questionable. Here, we quantify whinchat nest survival of (1) unprotected nests situated in early mown meadows, (2) protected nests situated in early mown meadows, and (3) nests situated in late mown meadows. Analyses considered the fact that successful and unsuccessful nests are not found with equal probabilities. Periods of reduced nest survival were associated with mowing periods in the different types of meadows. In early mown meadows nest survival rates were low (S < 0.1), and both conservation measures, individual nest protection and delayed mowing, resulted in significantly increased nest survival rates (S > 0.7). Individual nest protection cannot avoid changes in habitat quality of intensively managed meadows, and therefore is only suitable as small-scale and short-term measure to increase nest success until a high portion of late mown meadows is established. Thus, we suggest that a combination of the two measures applied to intensified grassland fields should be provided to maintain viable sizes of endangered meadow bird populations.