Why seedlings survive: influence of plant attributes

Seedling survival and successful forest restoration involves many silvicultural practices. One important aspect of a successful forest restoration program is planting quality seedlings with high survival capability. Thus the nursery needs to create seedlings with plant attributes that allow for the best chance of success once a seedling is field planted. Since the mid-twentieth century, research foresters have critically examined plant attributes that confer improved seedling survival to field site conditions. This review describes the value of commonly measured seedling quality material (i.e. shoot height, stem diameter, root mass, shoot to root ratio, drought resistance, mineral nutrient status) and performance (i.e. freezing tolerance and root growth) plant attributes defined as important in answering the question of why seedlings survive after planting. Desirable levels of these plant attributes can increase the speed with which seedlings overcome planting stress, become ‘coupled’ to the forest restoration site, thereby ensuring successful seedling establishment. Although planting seedlings with these desirable plant attributes does not guarantee high survival rates; planting seedlings with desirable plant attributes increases chances for survival after field planting.     

Enhancing Tree Seedling Supply to Smallholders in Leyte Province, Philippines: An Evaluation of the Production System of Government Nursery Sector and Support to Smallholder Tree Farmers

A decentralized seedling production is regarded as the most effective means of providing smallholders with planting materials. However, the sustainability of smallholder nursery operation is constrained by several factors, including the limited availability of germplasm, lack of technical skills and low seedling demand low viability of seedling market. The government nursery sector could complement the weaknesses of the smallholder nursery group but government nurseries are not operating effectively to enhance the availability of high quality seedlings of a wide species base for smallholder forestry. Seedling production in government nurseries is quantity-orientated with low emphasis on the production of high quality planting materials. Germplasm used in seedling production is mostly collected from unselected sources and seedlings produced are mostly of low physical quality. The species selection in government nurseries is heavily based on the availability of germplasm which resulted in the production of a few species which are commonly raised on private nurseries. The duplication of species of most seedlings raised in government nurseries with those produced in private nurseries and the far distance of government nurseries from the villages has resulted in low uptake of seedlings by smallholder tree farmers. For the government nursery sector to become more effective in providing support in the present paradigm of decentralized seeding production, a change from its production system and a broadening of the role it plays is needed. The social mobilization on seedling production as a scheme of implementing the Green Philippines Program has led the Department of Environment and Natural Resources to acknowledge the seedling production activity of private individual and communal nursery groups. This can be an opportunity for organizing the private and government nursery sectors to promote the operational effectiveness of the overall forest nursery industry.     

Initial seedling morphological characteristics and field performance of two Sudanian savanna species in relation to nursery production period and watering regimes

It has become apparent that some interventions are required to aid the regeneration of woody species in the Sudanian savanna. Direct seeding has been ineffective, thus planting high quality seedlings may be a viable alternative. In this study, we examined the stock quality of two valuable Sudanian savanna species, Acacia macrostachya and Pterocarpus erinaceus. Different nursery production periods were tested as well as the species’ field performance under well-watered and stressed conditions. The results showed that older seedlings (9-month) were morphologically distinct from younger ones (3-month), particularly in the case of P. erinaceus. Eighteen months after planting out, survival and growth of seedlings were not affected by initial seedling size; this was the result of the high root to shoot ratio of seedlings in all age groups at the time of planting. Seedling mortality as high as 30% was observed and attributed to both drought stress and other factors such as herbivory. Regression analyses revealed that initial shoot height was a poor predictor of field performance for both species, but initial root collar diameter accounted for 25% of the variation in diameter of P. erinaceus in the field. We conclude that initial seedling size does not affect survival and growth in the field provided that all sizes of seedling have a high root to shoot ratio at the time of planting. The prediction of field performance could be improved by developing a model that incorporates a wide range of root collar diameter.     

Coating nursery containers with copper carbonate improves root morphology of five native Australian tree species used in agroforestry systems

Seedlings of the native Australian tree species Acacia holosericea, Casuarina cunninghamiana, Eucalyptus argophloia, E. camaldulensis and Melaleuca bracteata, were raised in nursery containers which were coated with CuCO₃ (50 g l^-1, in a water-based paint solution) or left uncoated for three months. Seedling growth of all species in the nursery was generally less in treated containers, although differences were relatively small. More importantly, CuCO₃ treatment reduced the occurrence of lateral roots growing down or coiling around container walls for both three- and six-month-old seedlings. Twenty-four months after planting both three- and six-month-old seedlings on a semiarid site in subtropical southeast Queensland, there was no effect of CuCO₃ treatment on height or basal diameter development of seedlings. However, sample excavations of root systems showed that the poor root morphology exhibited by untreated seedlings when in the nursery had persisted in the field. As such poor root form has the potential to reduce tree growth and wind firmness in the longer term, these results suggest that chemical root-pruning is a helpful nursery practice when raising seedlings for dryland afforestation, and in circumstances where it may be necessary to retain seedlings in the nursery for longer than normal to take advantage of infrequent rainfall events. This revised version was published online in June 2006 with corrections to the Cover Date.     

Effects of boron fertilization in the nursery or after planting on the performance of Norway spruce seedlings on boron-poor sites

Many forest soils in Finland are poor in boron (B), especially otherwise fertile (nitrogen-rich) soils and peatlands. Boron deficiency causes growth disorders affecting young Norway spruce (Picea abies) stands. We studied the effects of B fertilization on the growth and morphology of Norway spruce seedlings. Boron was applied in a nursery and/or after planting in a nursery-field and on a reforestation site, both known to be low in B. The B applied in the nursery increased foliar B concentration in a linear relationship to the amount of B fertilizer but did not affect seedling growth and morphology. Even the highest foliar B concentration (approx. 400 mg kg^?1) did not have harmful effects on the seedlings. Therefore, B fertilization in the nursery can be used for increasing B storage in needles. Boron fertilization at planting increased foliar B concentration in the first season. Afterward, the concentration decreased but remained above the deficiency limit for at least 4–5 years. In the B-poor nursery-field, nonfertilized seedlings began to produce multiple leaders in the second year. On the reforestation site, fewer multiple leaders were found. At the nursery-field site, height growth was stimulated by B fertilization at planting. In conclusion, 200–400 mg m^?2 (2–4 kg ha^?1) of B applied at planting has longer-lasting effects on seedling B status than does B fertilization applied in the nursery, and can improve seedling quality and growth in the first years after planting on B-poor sites.     

Production of planting stock from wildings of four Shorea species

Seedling production from wildlings and early field performance of planted wildlings of four ecologically and commercially important Shorea species were studied in rainforest conditions of South Kalimantan (Indonesian Borneo). We report results from eight trials on Shorea johorensis, S. leprosula, S. faguetiana and S. parvifolia. Five trials were conducted in the nursery and three in the field. Growth medium, container type, wildling size or duration of shading period did not have significant effects on survival or growth of wildlings in the nursery. Keeping the potted wildlings in the greenhouse resulted in significantly higher survival than keeping them under shading net. Duration of greenhouse period had a significant effect on the survival of wildlings, with three weeks being the minimum period resulting in acceptable survival. Early field performance of hardened wildlings was better than that of seedlings raised from seed. Wildlings of 10–40 cm can be grown successfully in the nursery in several growth media and container types. A greenhouse period of 3–5 weeks is necessary for rooting and high survival. Use of wildlings is recommended as a supplementary method for producing dipterocarp planting stock. The techniques tested in the study are simple and appropriate for seedling production in conditions where skilled labor and advanced technology are not available.     

Using morphological characteristics for assessing seedling vitality in small-scale tree nurseries in Kenya

Small-scale tree nurseries are important in fulfilling the goals of reforestation and agroforestry implementation schemes in Kenya and other developing countries. The focus in seedling production has been on quantity, instead of quality, but a change can be seen in recent tree nursery manuals. These manuals are emphasising morphological characteristics as tools for assessing potential field performance of seedlings. However, morphological criteria are debatable and their value is questioned. A survey was carried out among tree nursery operators in the Meru area, in the Eastern province of Kenya, to determine how operators perceived seedling vitality, and how they separated acceptable seedlings from those of poor vitality. Based on the survey, 3 pairs of criteria were chosen, size (tall versus small), colour (green versus yellowish), and sturdiness quotient (sturdy versus lanky). These criteria were tested on survival and growth in a field trial, a controlled bench trial, and in a root growth potential test. The results showed that the nursery operators were aware of quality differences in seedlings, but they did not cull accordingly. The results from the field trial showed that mango (Mangifera indica L.) performed poorly compared to grevillea (Grevillea robusta A. Cunn. ex. R. Br.), probably due to the high altitude. The altitudinal range for mango and grevillea are 0-1,200 m and 0-2,300 m, respectively, and the trial site was located on an altitude of 1,725 m. In grevillea, small seedlings grew better than tall in the field trial, but no differences could be found in the other trials. In mango, sturdy seedlings grew better than lanky ones in the field trial, while in the controlled trials tall seedlings grew better than small ones. The results showed that morphological characteristics as seedling quality assessment criteria could be unreliable as the effect differs with species and planting site.     

Modified exponential nitrogen loading to promote morphological quality and nutrient storage of bareroot-cultured Quercus rubra and Quercus alba seedlings

Abstract

Exponential nutrient loading has been used to improve nursery fertilizer uptake efficiency of conifer seedlings, but the technique has received little attention in the culture of temperate deciduous hardwoods. This study examined responses of northern red oak (Quercus rubra L.) and white oak (Q. alba L.) seedlings to modified exponential nitrogen loading during bareroot nursery culture using a broad range of nutrient supply from 0 to 3.35 g nitrogen (N) per plant per season for 18 weeks in Indiana, USA. Seedling growth and nutritional parameters followed a curvilinear pattern that ranged from deficiency to toxicity with increased fertilization consistent with trends depicted in the proposed model for nutrient loading. Fertilization increased plant dry mass by 113–260% for red oak and 49–144% for white oak. Severe nutrient deficiency occurred under indigenous soil fertility, and limited phosphorus and potassium uptake were found to inhibit seedling growth at higher N supply. The sufficiency and optimum rates were determined to be 0.84 and 1.68 g N per seedling per season, respectively, under the current cultural conditions. Fertilization at 1.68 g N per plant increased N content by 40% in red oak and 35% in white oak. This approach may be used to help refine nursery fertilization practices in hardwood culture to produce high-quality seedlings for field planting.     

Restoration of hard mast species for wildlife in Missouri using precocious flowering oak in the Missouri River floodplain, USA

Increased planting of hard mast oak species in the Lower Missouri River floodplain is critical as natural regeneration of oak along the Upper Mississippi and Lower Missouri Rivers has been limited following major flood events in 1993 and 1995. Traditional planting methods have limited success due to frequent flood events, competition from faster growing vegetation and white-tailed deer herbivory. Results of early growth response of swamp white oak (Quercus bicolor Willd.) seedlings in relation to initial acorn mass and size, and early rapid shoot growth for seedlings produced by containerized root production method (RPM™), are presented. Containerized RPM™ seedlings grown in the greenhouse under optimal conditions demonstrate that seed size had no discernable impact on first-year root or shoot size. Seedling survival for the first two years and acorn production for the first three years after outplanting are presented, comparing use of containerized RPM™ swamp white oak seedlings to nursery stock. Flood tolerant precocious RPM™ oak seedlings in the floodplain provide a source of food for acorn-consuming wildlife ten to fifteen years sooner than oaks originating from natural regeneration, direct seeding or traditional bare root planting. Compared to bare root nursery stock that produced no acorns, some RPM™ swamp white oak seedlings averaged 4.3, 5.2, and 6.3 acorns/seedling in the first three years after fall outplanting. This revised version was published online in August 2006 with corrections to the Cover Date.     

Growth regulation of Scots pine seedlings with different fertilizer compositions and regimes

Scots pine (Pinus sylvestris L.) seedlings were grown in containers filled with peat, using two different fertilizers and three different fertilizer regimes. Seedling shoot and root growth and shoot content of nitrogen, potassium and phosphorus were followed in the nursery and after outplanting in the field. Attempts to regulate growth rate by an exponential nutrient supply were not successful, but the root/shoot ratio was influenced by the fertilization regime. Internal nitrogen concentration was stable only for seedlings with low relative growth rate, while seedlings with high nutrient supply in the nursery showed strong nutrient dilution in the shoot after planting.     

Effects of treeshelters and polyethylene mulch sheets on survival and growth of cork oak (Quercus suber L.) seedlings planted in northwestern Tunisia

Success of plantation establishment depends on the ability of seedlings to withstand animal browsing and competition from vegetation. This study in northwestern Tunisia examined the influence of browsing protection and vegetation control on seedling mortality, diameter, and height growth components of an evergreen sclerophyllous oak species, cork oak (Quercus suber L.) for the first 2 years after planting (2005 and 2006). Browsing protection was controlled by varying both treeshelter type (non-vented, vented, and control) and height (1.2-m and 1.8-m tall). Competing vegetation was controlled through increasing sizes of square-shaped black polyethylene mulch sheets: 0 m, 0.7 m, 1 m and 1.3 m large. Seedling mortality rate was relatively low during the first year (9%) and was not affected by any of the studied factors. During the following year, mortality was lower for seedlings in both types (non-vented and vented) of 1.2-m tall treeshelters than for unsheltered ones. Mortality was only slightly higher for 1.8-m tall shelters than for 1.2-m shelters. Compared to unsheltered seedlings, basal diameter was reduced during both seasons for seedlings inside non-vented treeshelters (1.2-m and 1.8-m tall), whereas for vented shelters it was reduced during the first year and then increased during the second year. Height growth was positively affected by tree shelters during both years. Seedlings inside non-vented 1.8-m tall shelters were the tallest having an average height of up to 140 cm at the end of the second year, while the unsheltered seedlings had only an average height of up to 42 cm. The greater height of the sheltered seedlings was attributed to the increase of: (1) the number of shoot flushes yearly established on the main stem (up to three), and (2) the length of all the growth units produced during each flush. Seedlings inside vented shelters had balanced growth with a low proportion of seedlings unable to support themselves without a stake (7%). However, non-vented shelters had unbalanced height and diameter growths resulting in both a higher height-to-diameter ratio and in a higher proportion of seedlings having stability problems (47%). Polyethylene mulch sheets did not affect early survival, seedling basal diameter, or number of shoot flushes established yearly, but did slightly improve seedling height at the end of the second season, mostly due to the significant improvement in shoot elongation of the first flush. Results suggest that both 1.8 m treeshelters and plastic mulches may enhance the growth of cork oak seedlings planted on harsh sites in northwestern Tunisia.     

Large mulches and manual release enhance growth of ponderosa pine seedlings

Woody and herbaceous vegetation that captures scarce soil moisture often kills or stunts the growth of conifer seedlings in young plantations in the Sierra Nevada of California. Two methods for excluding this vegetation are to apply large (3-m×3-m), long-lasting (at least 5 years) mulches around the seedlings soon after planting, or to repeatedly grub seedlings of competing vegetation in study plots. Both techniques were tested and proved to be effective, significantly increasing ponderosa pine diameter and height relative to the control and to seedlings surrounded by small paper mulches. The effective treatments were large enough to enable conifer seedlings to capture site resources unimpeded by competing plants. Eight species of shrubs, seventeen forbs, four grasses, and one fern grew naturally on the study site.Results suggest that the forester now has two biologically effective means for controlling unwanted vegetation and for attaining rapid early pine growth in environments similar to the study area. Although more expensive, the large heavy polyester mulches ($9.90 per seedling over 5 years) give the forester an alternative to manual release ($2.05 per seedling over 5 years).     

Influence of ectomycorrhizal fungal inoculation on growth and root IAA concentrations of transplanted conifers

We determined whether in vitro plant growth regulator production by mycorrhizal fungi is correlated with conifer seedling growth and root IAA concentrations. Container-grown seedlings of interior Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), lodgepole pine (Pinus contorta Dougl.) and ponderosa pine (Pinus ponderosa Dougl.) were inoculated at seeding with ectomycorrhizal fungi having a high, moderate or low capacity to produce either IAA or ethylene in vitro. Inoculated seedlings were grown for one season in the nursery, harvested in December, cold stored over winter and then transplanted to either a nursery field or a forest site in the spring. Seedling morphology and endogenous IAA in roots were measured immediately after cold storage and again six and 12 months after transplanting. Morphological responses to inoculation varied among different mycorrhizal fungi. Free IAA concentration of roots was increased in some inoculation treatments for all conifer species. In seedlings transplanted to a nursery field, in vitro ethylene-producing capacity of the ectomycorrhizal fungi was highly correlated with more morphological features than in vitro IAA-producing capacity. Both IAA- and ethylene-producing capacity were significantly correlated with more morphological features in seedlings transplanted to a forest site than in seedings transplanted to a nursery field. One year after transplanting, only in vitro IAA-producing capacity was correlated with endogenous IAA concentration of roots of the inoculated seedlings. We conclude that growth responses of conifer seedlings can be partially influenced by IAA and ethylene produced by ectomycorrhizal fungal symbionts.     

A review of ecophysiologically-based seedling specifications for temperate Australian eucalypt plantations

The Australian hardwood plantation estate has significantly expanded in the last decade. Key production issues included: (1) will Eucalyptus nitens Maiden seedlings produced in nurseries on relatively mild sites be susceptible to transplant shock and browsing on cold out-planting sites and can this be managed? And; (2) what specifications should be required of nursery producers for Eucalyptus globulus Labill seedlings for out-planting on mild ex-pasture sites? This review outlines the research conducted in order to address these key questions. Nitrogen deprivation in the nursery induced accumulation of foliar anthocyanin which conferred hardiness to low temperature. Target specifications of 8–10?g/kg foliar nitrogen are recommended. Height of >120?mm, root collar diameter >2.8?mm, seedling container volume of ≥85?cm³, depth of ≥73?mm, density of ≥549?cells/m² and foliar nitrogen concentration of 15–20?g/kg were found to be key specifications for optimal E. globulus growth following planting onto mild ex-pasture sites. Exponential nutrient loading was a useful technique for attaining target foliar N concentrations. Seedling size was found not to affect growth of E. globulus post-planting on mild ex-pasture sites. However, large seedlings were preferentially browsed on high vertebrate-browse-pressure sites in Tasmania (poisons are banned and shooting is not effective on some sites) that did not affect subsequent growth rate but did increase the incidence of double-leaders. The ‘target seedling concept’ is applicable to eucalypts for Australian planting sites: specifications attained in the nursery should be matched to the factors limiting growth on the planting sites.     

Influences of seedling size,container type and mammal browsing on the establishment of <Emphasis Type="Italic">Eucalyptus globulus</Emphasis> in plantation forestry

Rapid early growth of tree seedlings is critical to the success of plantation establishment. We investigated the effects of seedling size (small and large) and container types (small [Lannen 121], medium [Lannen 81] and large [Forestry Tube]) in the nursery and the effects of mammal browsing after planting on growth of Eucalyptus globulus in Tasmania’s Southern Forests. After planting, seedlings were either exposed to browsing or protected from browsing by wire-mesh cages until age 6 months. Low browsing pressure resulted in around 20% and 5–10% of foliage being browsed in the large and small size categories, respectively, between 1 and 3 months after planting the uncaged treatment. 6 months after planting, height growth increment was lower, and 4 years after planting, mortality was higher in uncaged large than caged large seedling treatments. Six and twelve months after planting, seedlings raised in Forestry Tube containers had significantly greater height increment and root collar diameter relative to other treatments. By 4 years after planting, trees of the small seedling treatment had significantly greater diameter than those of the large seedling treatment, but there was no effect of container type treatment. Four years after planting there was no effect on diameter growth by browsing of <30% of foliage up to 3 months after planting, although there was greater incidence of double leaders in trees that had been browsed as seedlings. Small seedlings produced more growth 4 years after planting than large seedlings of E. globulus.     

Slash pine bud dormancy as affected by lifting date and root wrenching in the nursery

Bud dormancy of root wrenched and unwrenched slash pine (Pinus elliottii Engelm.) seedlings growing in a forest nursery was measured on five lifting dates. Determination of bud dormancy was based on days to budbreak (DBB) under optimal growing conditions, mitotic activity in the apical meristem, chilling hours accumulated, and bud morphology. Based on DBB, seedlings were most dormant at Lift 2 on November 24 after exposure to 189 hours below 10 degrees C and 93 hours below 6.7 degrees C. Mitotic activity in the apical meristem was at its lowest 23 days later at Lift 3, possibly indicating the period when seedlings are most resistant to transplanting stresses. Multiple wrenching resulted in a slight shift in the dormancy cycle as wrenched seedlings set bud sooner in the nursery and broke bud sooner at the planting site in the spring than control seedlings. This implies that wrenched seedlings can be successfully lifted from the nursery earlier and will initiate spring shoot growth earlier than control seedlings.     

Comparing large containerized and bareroot conifer stock on sites of contrasting vegetation composition in a non-herbicide scenario

Planting stock selection is an integral part of plantation management, as forest nursery practices influence the physiological status of the seedlings and their capacity to cope with resource availability on different planting sites. We thus compared the 11th-year dimensions and survival of large white spruce (Picea glauca) and black spruce (P. mariana) seedlings produced as 2 + 2 bareroot or 2 + 0 container stock (cell volume of 350 cm³), used to reduce the need for competition control. Using complete split-block designs, we evaluated the seedling competitive potential and response to mechanical release on two sites of contrasting ecological fertility and vegetation dominance in Quebec, Canada. We found that large spruce seedlings can be successfully established on high-competition sites in a context where chemical herbicides cannot be used. These stock types had a limited impact on survival and growth, and both stock responded similarly to mechanical vegetation control. In this context, the choice of stock type should prioritize the highest quality large seedling based on operational considerations such as availability and ease of transportation. Mechanical site preparation was not necessary to promote seedling growth and survival on these sub-boreal sites.     

Silvicultural options to promote seedling establishment on Kalmia–Vaccinium-dominated sites

Seedling growth is often hampered on sites dominated by Kalmia angustifolia. In June 2000, a trial was established on a clear-cut site in Quebec, Canada, with a high cover of Kalmia and Vaccinium species. The objectives were to evaluate how soil scarification and fertilization at the time of planting influence early growth and establishment of black spruce [Picea mariana (Mill.) BSP] and jack pine (Pinus banksiana Lamb.) seedlings. During the first 2 years, scarification reduced Kalmia cover three-fold and doubled the distance from seedlings to the nearest Kalmia stem. Scarification did not increase soil-extractable NH₄-N concentration, and reduced soil potassium, calcium and magnesium. Scarification had no effect on seedling water stress. Seedling growth improved and foliar nutrient concentrations were generally higher in scarified plots than in unscarified control plots. No differences were observed between single- and double-pass scarification for any variables except for ground-level stem diameter of seedlings, which was greater with double-pass scarification (12.1 vs 13.1 mm). Spot fertilization increased seedling growth and foliar nitrogen concentrations. Jack pine growth was greater than black spruce growth, an effect enhanced when seedlings were fertilized.     

Repellent and stocking guards reduce mammal browsing in eucalypt plantations

Mammalian herbivores regularly browse plantation seedlings. In many areas, seedlings require some form of protection if they are to survive and grow into a productive plantation. Two general approaches for reducing browsing damage to seedlings are to apply chemical repellents and tree guards. Both methods have existed for a long time, and new variations are constantly being developed. Seedling stocking guards, a type of tree guard, are being used operationally in Tasmania, Australia, but there is limited data quantifying their effectiveness and concerns with negative effects on tree performance. Conversely, although proven effective, repellents are not being used, but are potentially cheaper and less problematic. We therefore aimed to determine which is more effective under operational conditions, whether this effectiveness can be improved or extended, and if either treatment has any effects on seedling form or survival. We planted Eucalyptus nitens seedlings with combinations of repellent and stocking guards in six operational plantations to examine and compare their effectiveness. Seedlings were monitored for 12 months to examine treatment longevity. We found that both stocking guards and repellent significantly reduced and delayed browsing severity, with their effects being additive. No negative effects on growth were evident after 12 months, but adverse effects on seedling form warrant further investigation. Both of these methods can be easily and relatively cheaply applied in the nursery before planting, making them appealing options to reduce browsing. However, the ideal method for a given site will depend upon local browsing intensity.     

Tubed Seedling Research and Development in Britain

Based initially on Canadian practice, research on the productionand use of tubed seedlings in Britain has been in progress for3 years (1968–70), and was begun because of their potentialadvantages as planting stock. Polythene greenhouses are usedto produce pine and spruce seedlings grown in small plastictubes filled with fertilized peat-sand compost, and plantingis possible 8 weeks after sowing. On ploughed peat-land, step-plantedseedlings are capable of high survival and vigorous early growth,with an effective planting season from mid April to late August.Animal browsing is the main adverse factor encountered, buthas been less serious than expected. Very high planting ratesare possible with a special tool. Seedling use could lead tolarge savings in peatland afforestation costs, and large-scaletrials are planned. In contrast, seedling performance on ploughedmineral soil has been unsatisfactory due to severe frost liftingand poor height growth. Results in reafforestation trials havenot been very promising.