施用氮磷钾对黄土丘陵区山地红枣林土壤酶与土壤肥力的影响

采用野外试验与室内分析,研究了连续4 a施氮磷钾肥对黄土丘陵区山地枣园土壤酶活性及土壤养分含量的影响。结果表明:连续4 aNPK1施肥处理能够明显增强土壤脲酶、磷酸酶、过氧化氢酶和蔗糖酶的活性,提高土壤有机质、全氮、速效氮、速效磷和速效钾含量;在土壤剖面中总的趋势是土壤酶活性和土壤速效氮、速效磷和速效钾含量随土层深度加深而降低。土壤酶活性与土壤养分因子的相关分析表明,在0~20 cm土层,磷酸酶与有机质、速效磷,过氧化氢酶与有机质、全氮呈显著或极显著性正相关。在20~40 cm,脲酶与有机质、全氮、速效氮,磷酸酶与速效钾,过氧化氢酶与速效磷,蔗糖酶与有机质、速效钾呈显著或极显著性正相关。在40~60 cm,脲酶与全氮、速效磷,磷酸酶与有机质、速效氮,过氧化氢酶与速效磷,蔗糖酶与有机质、全氮、速效磷呈显著或极显著正相关。     

Ecological biomass allocation strategies in plant species with different life forms in a cold desert,China

Biomass allocation patterns among plant species are related to their adaptive ecological strategies. Ephemeral, ephemeroid and annual plant life forms represent three typical growth strategies of plants that grow in autumn and early spring in the cold deserts of China. These plants play an important role in reducing wind velocity in the desert areas. However, despite numerous studies, the strategies of biomass allocation among plant species with these three life forms remain contentious. In this study, we conducted a preliminary quadrat study during 2014-2016 in the southern part of the Gurbantunggut Desert, China, to investigate the allocation patterns of above-ground biomass (AGB) and below-ground biomass (BGB) at the individual level in 17 ephemeral, 3 ephemeroid and 4 annual plant species. Since ephemeral plants can germinate in autumn, we also compared biomass allocation patterns between plants that germinated in autumn 2015 and spring 2016 for 4 common ephemeral species. The healthy mature individual plants of each species were sampled and the AGB, BGB, total biomass (TB), leaf mass ratio (LMR) and root/shoot ratio (R/S) were calculated for 201 sample quadrats in the study area. We also studied the relationships between AGB and BGB of plants with the three different life forms (ephemeral, ephemeroid and annual). The mean AGB values of ephemeral, ephemeroid and annual plants were 0.806, 3.759 and 1.546 g/plant, respectively, and the mean BGB values were 0.106, 4.996 and 0.166 g/plant, respectively. The mean R/S value was significantly higher in ephemeroid plants (1.675) than in ephemeral (0.154) and annual (0.147) plants. The mean LMR was the highest in annual plants, followed by ephemeroid plants and ephemeral plants, reflecting the fact that annual plants allocate more biomass to leaves, associated with their longer life span. Biomass of ephemeral plants that germinated in autumn was significantly higher than those of corresponding plants that germinated in spring in terms of AGB, BGB and TB. However, the R/S value was similar in plants that germinated in autumn and spring. The slope of regression relationship between AGB and BGB differed significantly among the three plant life forms. These results support different biomass allocation hypotheses. Specifically, at the individual level, the AGB and BGB partitioning supports the allometric hypothesis for ephemeroid and annual plants and the isometric hypothesis for ephemeral plants.     

Changes in soil organic carbon and nitrogen after 26 years of farmland management on the Loess Plateau of China

Soil carbon(C) and nitrogen(N) play a crucial role in determining the soil and environmental quality. In this study, we investigated the effects of 26 years(from 1984 to 2010) of farmland management on soil organic carbon(SOC) and soil N in abandoned, wheat(Triticum aestivum L.) non-fertilized, wheat fertilized(mineral fertilizer and organic manure) and alfalfa(Medicago Sativa L.) non-fertilized treatments in a semi-arid region of the Loess Plateau, China. Our results showed that SOC and soil total N contents in the 0–20 cm soil layer increased by 4.29(24.4%) and 1.39 Mg/hm2(100%), respectively, after the conversion of farmland to alfalfa land. Compared to the wheat non-fertilized treatment, SOC and soil total N contents in the 0–20 cm soil layer increased by 4.64(26.4%) and 1.18 Mg/hm2(85.5%), respectively, in the wheat fertilized treatment. In addition, we found that the extents of changes in SOC, soil total N and mineral N depended on soil depth were greater in the upper soil layer(0–30 cm) than in the deeper soil layer(30–100 cm) in the alfalfa land or fertilizer-applied wheat land. Fertilizer applied to winter wheat could increase the accumulation rates of SOC and soil total N. SOC concentration had a significant positive correlation with soil total N concentration. Therefore, this study suggested that farmland management, e.g. the conversion of farmland to alfalfa forage land and fertilizer application, could promote the sequestrations of C and N in soils in semi-arid regions.     

黄土高原半干旱区植被重建对不同坡位土壤肥力质量的影响

对黄土高原半干旱山地植被重建后不同坡位土壤肥力质量的变化和空间分布特征进行了研究。结果表明:随着人工植被的建立与恢复,有效提高了土壤持水量;不同坡位土壤pH值及全盐含量与荒山相比明显降低,而土壤肥力显著提高,尤其土壤全氮和速效磷含量增加幅度较大;在坡面位置中,上坡位土壤养分含量与下坡位相比普遍要高,而对于荒山则是下坡位的普遍高于上坡位。从坡顶到坡脚表层土壤的全盐含量依次增加,下坡位全盐含量明显高于其它坡位。土壤脲酶活性随着植被重建增加幅度较大,但各坡位间差异不十分明显;表层土壤过氧化氢酶活性对于植被的重建响应不明显,而20-40cm土层的过氧化氢酶活性却显著提高。坡面土壤肥力质量的变化是土地利用方式、植被类型与景观位置分异综合作用的结果,植被恢复和建设对提高该区土壤肥力质量具有重要的作用。     

行间种植不同草种对幼龄苹果园土壤特性的影响

于苹果园建植第一年春季,果树行间分别种植不同草种,以清耕的土壤为对照,研究果园土壤温度、含水量、pH值、有机质及N、P、K的变化。结果表明,种草后,夏季高温时土壤温度降低、日变幅减小,含水量增加,生长季结束时,果园土壤pH下降,碱解N、全N、速效P、全P、有机质含量增加,而除种植红三叶的土壤速效K显著(P<0.05)升高外,其余处理全K、速效K均无显著变化;种草后对不同深度土壤肥力的影响不同,马蹄金对土壤N、P和狗牙根与高羊茅对有机质的影响在0~20 cm内表现一致,而种植多年生黑麦草、狗牙根、紫羊茅的0~10 cm土壤碱解N、全N、全P和种植红三叶、紫羊茅的0~10 cm土壤有机质升高幅度较10~20 cm大,多年生黑麦草、高羊茅、紫羊茅、红三叶、白三叶的速效P,红三叶和高羊茅的碱解N、全N,马蹄金、多年生黑麦草、白三叶的土壤有机质0~10 cm土壤升高幅度较10~20 cm小,白三叶10~20 cm土壤碱解N增加幅度较0~10 cm小,全N则反之;草种不同对土壤作用不一,马蹄金对土壤肥力改善作用最大,红三叶、白三叶次之,而狗牙根、高羊茅较差。说明种草后可改善土壤性状,而草种是影响果园种草效益的关键因素。     

Plant cover as an estimator of above-ground biomass in semi-arid woody vegetation in Northeast Patagonia,Argentina

The quantification of carbon storage in vegetation biomass is a crucial factor in the estimation and mitigation of CO₂ emissions. Globally, arid and semi-arid regions are considered an important carbon sink. However, they have received limited attention and, therefore, it should be a priority to develop tools to quantify biomass at the local and regional scales. Individual plant variables, such as stem diameter and crown area, were reported to be good predictors of individual plant weight. Stand-level variables, such as plant cover and mean height, are also easy-to-measure estimators of above-ground biomass (AGB) in dry regions. In this study, we estimated the AGB in semi-arid woody vegetation in Northeast Patagonia, Argentina. We evaluated whether the AGB at the stand level can be estimated based on plant cover and to what extent the estimation accuracy can be improved by the inclusion of other field-measured structure variables. We also evaluated whether remote sensing technologies can be used to reliably estimate and map the regional mean biomass. For this purpose, we analyzed the relationships between field-measured woody vegetation structure variables and AGB as well as LANDSAT TM-derived variables. We obtained a model-based ratio estimate of regional mean AGB and its standard error. Total plant cover allowed us to obtain a reliable estimation of local AGB, and no better fit was attained by the inclusion of other structure variables. The stand-level plant cover ranged between 18.7% and 95.2% and AGB between about 2.0 and 70.8 Mg/hm². AGB based on total plant cover was well estimated from LANDSAT TM bands 2 and 3, which facilitated a model-based ratio estimate of the regional mean AGB (approximately 12.0 Mg/hm²) and its sampling error (about 30.0%). The mean AGB of woody vegetation can greatly contribute to carbon storage in semi-arid lands. Thus, plant cover estimation by remote sensing images could be used to obtain regional estimates and map biomass, as well as to assess and monitor the impact of land-use change on the carbon balance, for arid and semi-arid regions.     

Effects of vegetation patterns and environmental factors on woody regeneration in semi-arid oak-dominated forests of western Iran

This study assesses the effects of vegetation patterns and environmental factors on the abundance of natural tree and shrub regeneration in semi-arid forests of the Zagros Mountains, western Iran. We sampled 120 releves at different topographic positions in a protected area of the studied region. Floristic composition, slope, elevation and soil properties were recorded at each releve, and woody seedling density was measured. We have first discerned five floristic groups using two-way indicator species analysis(TWINSPAN), detrended correspondence analysis(DCA), and canonical correspondence analysis(CCA) and then explored the relationships among the floristic group compositions, environmental factors and seedling densities. The indicator species of the five groups were Quercus brantii, Acer monspessulanum, Cerasus microcarpa, Rhamnus arvensis and Astragalus licyoides. Our results indicated that these groups were significantly affected by elevation and soil properties and the soil properties refer to: EC(electrical conductivity), N(nitrogen), K(potassium), OM(organic matter), and bulk density. Woody regeneration was composed of Q. brantii, A. monspessulanum, C. microcarpa, Amygdalus scoparia and Crataegus pontica seedlings. The highest density of seedlings was found for Q. brantii(97.14(±48.00) plants/hm~2) and the lowest for A. scoparia(2.28(±1.50) plants/hm~2). Quercus brantii was the dominant species and the seedling density was positively correlated with soil p H and P(phosphorus) values. Amygdalus scoparia regeneration was negatively correlated with elevation, and the seedling density peaked in C. microcarpa group. There was no significant variation in distribution of C. pontica seedlings among the groups, but the seedling density of this species was positively correlated with slope and K. Cerasus microcarpa seedlings were more abundant in the Q. brantii group than in other groups. This study showed that the regeneration of tree and shrub species was unequally distributed in different floristic groups for some species(A. scoparia and C. microcarpa) but not for other(Q. brantii and C. pontica) and was generally correlated with some environmental factors, particularly elevation, slope and soil nutrients(P and K). These results are a first step to implement future management and restoration strategies for promoting forest regeneration.     

黄土高原农牧交错带退耕坡地苜蓿-铁杆蒿群落养分分布特征

植物群落养分的空间分布反映了生态系统物质循环特征,可以为揭示生态系统资源的有效性及生态系统中植物群落的演替过程提供依据。本研究以黄土高原农牧交错带退耕坡地苜蓿-铁杆蒿群落为对象,分析其生物量和养分含量的空间分布特征。结果表明:退耕地苜蓿-铁杆蒿群落坡上部以苜蓿为主导物种,坡下部以铁杆蒿为主导物种,坡中部由二者共同主导。同一植物的生物量和N含量具有相似的空间结构,并且与植物种类有关;苜蓿和铁杆蒿P含量均以球状模型最佳,K含量的空间分布均以高斯模型最佳。苜蓿-铁杆蒿群落生物量和养分含量具有较强的空间依赖性,其空间变异主要由结构性因素决定。苜蓿和铁杆蒿生物量和养分含量的自相关性随距离变化表现出相似的趋势,但随生物量和养分种类的不同而异,且铁杆蒿生物量和养分含量的自相关性大于苜蓿。随着人工草地向天然草地的演替,群落生物量和养分含量的空间自相关性和空间依赖性逐渐增强。     

Seasonal patterns in water uptake for Medicago sativa grown along an elevation gradient with shallow groundwater table in Yanchi county of Ningxia,Northwest China

Lucerne(Medicago sativa L.) is a deep-rooted perennial leguminous forage with high evapo-transpiration rate exceeding the annual precipitation in semi-arid areas of Northwest China.Groundwater might be the potential water sources of lucerne in the area with shallow groundwater table.In this study,stable isotopic compositions of oxygen and hydrogen(δ18O and δD) of different water sources and xylem sap were analyzed to determine the seasonal(April,June,July and August) and topographic(three slope positions) variations in water sources for lucerne growing in Ningxia eastern semi-arid area adjacent to Mu Us Desert characterized by shallow groundwater table.Iso Source software was used to calculate the probable contribution of different water sources to the total plant water uptake.Stomatal conductance,stem water potential,carbon isotope discrimination(?~(13)C) of whole plant were also determined for evaluating the water status of lucerne growing at different slope positions.The results showed that soil water content increased as the elevation decreased.Oxygen compositions of soil water in the 0–40 cm profile fluctuated considerably.Soil water δ~(18)O values in deep profile(3.5 m) were similar to those in groundwater,implying the recharge of groundwater to this soil layer.Highest water utilization rate from deep soil profile(below 350 cm) was recorded for lucerne grown at the slope position 1(groundwater table depth of 3.5–3.9 m) in April,June and July.The lucernes at slope position 2(groundwater table depth of 5.8–6.4 m) and slope position 3(groundwater table depth 7.1–8.3 m) mostly used water from deep soil layers(below 350 cm) during dry period,and turned to use water from superficial soil layer in wet period.Higher yield,?~(13)C value of whole plant and stomatal conductance were observed for lucerne grown at the slope position 1 than those at other slope positions.These results indicated that groundwater is a significant water source for transpiration of lucerne grown in Ningxia semi-arid area with shallow groundwater table where lucerne grassland is suggested to be established so as to obtain better yield performance.     

Response of C:N:P in the plant-soil system and stoichiometric homeostasis of Nitraria tangutorum leaves in the oasis-desert ecotone,Northwest China

Nitraria tangutorum nebkhas are widely distributed in the arid and semi-arid desert areas of China. The formation and development of N. tangutorum nebkhas are the result of the interaction between vegetation and the surrounding environment in the process of community succession. Different successional stages of N. tangutorum nebkhas result in differences in the community structure and composition, thereby strongly affecting the distribution of soil nutrients and ecosystem stability. However, the ecological stoichiometry of N. tangutorum nebkhas in different successional stages remains poorly understood. Understanding the stoichiometric homeostasis of N. tangutorum could provide insights into its adaptability to the arid and semi-arid desert environments. Therefore, we analyzed the stoichiometric characteristics of N. tangutorum in four successional stages, i.e., rudimental, developing, stabilizing, and degrading stages using a homeostasis model in an oasis-desert ecotone of Northwest China. The results showed that soil organic carbon (SOC), total nitrogen (TN), and total phosphorus (TP) contents and their ratios in the 0-100 cm soil depth were significantly lower than the averages at regional and global scales and were weakly influenced by successional stages in the oasis-desert ecotone. TN and TP contents and C:N:P in the soil showed similar trends. Total carbon (TC) and TN contents in leaves were 450.69-481.07 and 19.72-29.35 g/kg, respectively, indicating that leaves of N. tangutorum shrubs had a high storage capacity for C and N. Leaf TC and TN contents and N:P ratio increased from the rudimental stage to the stabilizing stage and then decreased in the degrading stage, while the reverse trend was found for leaf C:N. Leaf TP content decreased from the rudimental stage to the degrading stage and changed significantly in late successional stages. N:P ratio was above the theoretical limit of 14, indicating that the growth of N. tangutorum shrubs was limited by P during successional stages. Leaf N, P, and N:P homeostasis in four successional stages was identified as 'strictly homeostasis'. Redundancy analysis (RDA) revealed that soil acidity (pH) and the maximum water holding capacity were the main factors affecting C:N:P stoichiometric characteristics in N. tangutorum leaves. Our study demonstrated that N. tangutorum with a high degree of stoichiometric homeostasis could better cope with the arid desert environment.     

Effects of land-use types on the vertical distribution of fractions of oxidizable organic carbon on the Loess Plateau,China

The oxidizability of soil organic carbon(SOC_) influences soil quality and carbon sequestration. Four fractions of oxidizable organic carbon(very labile(C_1), labile(C_2), less labile(C_3) and non-labile(C_4)) reflect the status and composition of SOC_ and have implications for the change and retention of SOC_. Studies of the fractions of oxidizable organic carbon(OC_) have been limited to shallow soil depths and agroecosystems. How these fractions respond at deep soil depths and in other types of land-use is not clear. In this study, we evaluated the vertical distributions of the fractions of oxidizable organic carbon to a soil depth of 5.0 m in 10 land-use types in the Zhifanggou Watershed on the Loess Plateau, C_hina. Along the soil profile, C_1 contents were highly variable in the natural grassland and shrubland I(C_aragana microphylla), C_2 and C_4 contents were highly variable in the natural grassland and two terraced croplands, respectively, and C_3 contents varied little. Among the land-use types, natural grassland had the highest C__1 and C_2 contents in the 0–0.4 m layers, followed by shrubland I in the 0–0.1 m layer. Natural grassland had the highest C_4 contents in the 1.0–4.5 m layers. Natural grassland and shrubland I thus contributed to improve the oxidizability of SOC_ in shallow soil, and the deep soil of natural grassland has a large potential to sequester SOC_ on the Loess Plateau.     

Impact of Fire on Soil Nitrogen Forms in Central Semiarid Argentina

Burning effects on soil N are particularly important in the Caldenal region of Argentina because N is likely to be limited and is easily lost by volatilization during fire. Burning may cause an initial increase in the mineral N content of the surface soil and, perhaps more important, can reduce the proportion of N in hydrolyzable (that is, more mineralizable) forms. The modification in soil N fractions by fire is directly related to the degree of soil heating and to the proportion and amount of N fractions remaining in the ash. This research was aimed at studying the effects of a controlled burn on total N, inorganic (NO3 and NH4+) and organic (amino acids, ammonium, hexosamines) N forms, as well as hydrolyzable and nonhydrolyzable N compounds of top soils of the southern Caldenal. Soil samples were taken from beneath the canopy of shrubs and from the areas with herbaceous cover, before and after burning. In order to show that a greater proportion of N added to regularly burned soils will result in lower levels of mineralizable N, differences in the changes of the various pools of plant tissue N upon burning at different ignition tem peratures are also reported. The initial unburned top soils had no significant differences in total, inorganic, and organic N form concentrations between shrub and herbaceous vegetation-covered soil. Burning did not affect the net content of total N in soil under woody vegetation, but in the grassland soil a decrease of total N concentration was observed. Nitrate concentration increased and NH4+ concentration decreased in both shrub and grassland soils after burning. Controlled burning caused losses of amino acid and ammonium organic N forms from the soil under grass plants and gains of hexosamine N in both shrub-and herbaceous-coveredsoils. There was an observed tendency to increase the relative proportions of non hydrolyzable N forms in both the shrub and grassland soils after burning. Inorganic and organic N form concentrations were similar in initial shrub litter and grass, but litter had higher concentrations of total and nonhydrolyzable N than those of grass. Shrub litter ash obtained after muffling at 300 C contained higher NH4+-N, aminoacid N, ammonium N, and total and nonhydrolyzable N concentrations than those from grass. An increase of 100 C in the ignition temperature produced clear ashes, rich in mineral N with little concentration of total N and C. The organic N forms affected most by the ignition temperature increase were amino acid N and ammonium N, respectively, particularly in woody litter. The most striking feature was the constituent increase in the percentages of nonhydrolyzable N forms of the vegetation covers on passing from initial to 300 C to 400 C, particularly for the litter and its ashes. These results suggest that the soil mineralizable N pool may be reduced by frequent burning or by more severe burning than observed presently in the Caldenal region.     

Effects of rainfall patterns on annual plants in Horqin Sandy Land,Inner Mongolia of China

Growth of annual plants in arid environments depends largely on rainfall pulses. An increased understanding of the effects of different rainfall patterns on plant growth is critical to predicting the potential responses of plants to the changes in rainfall regimes, such as rainfall intensity and duration, and length of dry intervals. In this study, we investigated the effects of different rainfall patterns(e.g. small rainfall event with high frequency and large rainfall event with low frequency) on biomass, growth characteristics and vertical distribution of root biomass of annual plants in Horqin Sandy Land, Inner Mongolia of China during the growing season(from May to August) of 2014. Our results showed that the rainfall patterns, independent of total rainfall amount, exerted strong effects on biomass, characteristics of plant growth and vertical distribution of root biomass. Under a constant amount of total rainfall, the aboveground biomass(AGB), belowground biomass(BGB), plant cover, plant height, and plant individual and species number increased with an increase in rainfall intensity. Changes in rainfall patterns also altered the percentage contribution of species biomass to the total AGB, and the percentage of BGB at different soil layers to the total BGB. Consequently, our results indicated that increased rainfall intensity in future may increase biomass significantly, and also affect the growth characteristics of annual plants.     

土壤侵蚀对土壤理化性质及土壤微生物的影响

通过对黄土丘陵沟壑区侵蚀环境下4种典型坡面上坡顶、坡肩、坡背、坡脚和坡趾5个不同地形部位¹³⁷Cs的含量、土壤理化性质及土壤微生物指标的测定和分析。结果表明:用来表征土壤侵蚀程度的¹³⁷Cs含量与土壤有机碳、全氮、土壤容重、电导率、土壤微生物功能多样性及细菌菌群之间均存在显著的回归关系,土壤有机碳、全氮及土壤容重随着¹³⁷Cs含量的增加呈增加趋势,土壤电导率、土壤微生物功能多样性呈下降趋势,而土壤细菌呈先下降后增加的趋势。土壤侵蚀过程能够直接影响土壤养分在坡面上的空间分布及土壤结构,通过影响土壤性质,改变土壤微生物群落的生长环境和营养物质的含量,从而影响土壤微生物的生长。     

Estimating aboveground biomass in Mu Us Sandy Land using Landsat spectral derived vegetation indices over the past 30 years

Remote sensing is a valuable and effective tool for monitoring and estimating aboveground biomass （AGB） in large areas.The current international research on biomass estimation by remote sensing technique mainly focused on forests,grasslands and crops,with relatively few applications for desert ecosystems.In this paper,Landsat Thematic Mapper （TM）/Enhanced Thematic Mapper Plus （ETM＋） images from 1988 to 2007 and the data of 283 AGB samples in August 2007 were used to estimate the AGB for Mu Us Sandy Land over the past 30 years.Moreover,temporal and spatial distribution characteristics of AGB and influencing factors of climate and underlying surface were also studied.Results show that：（1） Differences of correlations exist in the fitted equations between AGB and different vegetation indices in desert areas.The modified soil adjusted vegetation index （MSAVI） and soil adjusted vegetation index （SAVI） show relatively higher correlations with AGB,while the correlation between normalized difference vegetation index （NDVI） and AGB is relatively lower.Error testing shows that the AGB-MSAVI model established can be used to accurately estimate AGB of Mu Us Sandy Land in August.（2） AGB in Mu Us Sandy Land shows the fluctuant characteristics over the past 30 years,which decreased from the 1980s to the 1990s,and increased from the 1990s to 2007.AGB in 2007 had the highest value,with a total AGB of 3.352×106 t.Moreover,in the 1990s,AGB had the lowest value with a total AGB of 2.328×106 t.（3） AGB with relatively higher values was mainly located in the middle and southern parts of Mu Us Sandy Land in the 1980s.AGB was low in the whole area in the1990s,and relatively higher AGB values were mainly located in the southern parts of Uxin.In 2007,AGB in the whole area was relatively higher than those of the last twenty years,and higher AGB values were mainly located in the northern,western and middle parts of Mu Us Sandy Land.     

玛曲高寒草地沙化对土壤理化特性的影响

以玛曲高寒草地处于不同沙化发展阶段的潜在沙化地、轻度沙化地、中度沙化地和重度沙化地为研究对象,阶地作为对照,对0～60 cm土壤剖面的机械组成和土壤有机碳、全氮、全磷以及速效钾含量进行了测定,分析结果表明:1)随草地沙化程度的加剧,不同类型沙化草地的黏粒和粉粒含量迅速下降,细砂粒含量逐渐下降而粗砂粒含量逐渐上升。2)随草地沙化程度的加剧,不同程度沙化草地的有机碳和全N含量呈逐渐下降趋势。3)从不同程度沙化草地各层土壤的养分含量来看,阶地和潜在沙化地各土层有机碳和全N含量随土层深度加深逐渐下降,中度沙化地、重度沙化地各土层有机碳和全N含量呈现基本稳定的状态。     

黄河口翅碱蓬湿地土壤氮的季节变化

2009年4～11月,选择位于黄河口滨岸中潮滩和低潮滩上的翅碱蓬湿地为研究对象,对比研究了湿地土壤氮的季节变化特征。结果表明:两种湿地不同土层的NO3--N、NH4+-N和TN含量均具有明显的季节变化特征,但不同类型湿地之间差异较大。二者NO3--N、NH4+-N含量的季节差异主要与其所受潮汐影响程度及其引起的无机氮物理运移和对NH4+-N吸附能力的差异有关。二者在7月较低的NO3--N含量主要与植物对土壤中有效氮的大量吸收与利用有关,而8月较高的NO3--N、NH4+-N含量主要与此间土壤有机氮矿化较强有关。两种湿地不同土层的TN含量与C/N的季节变化规律整体相反。二者TN含量的差异主要受制于不同土层的有机质分布以及潮汐的影响状况,而C/N的差异主要与两种湿地在不同时期受陆源的影响程度有关。     

Shrub modulates the stoichiometry of moss and soil in desert ecosystems,China

Desert mosses, which are important stabilizers in desert ecosystems, are distributed patchily under and between shrubs. Mosses differ from vascular plants in the ways they take up nutrients. Clarifying their distribution with ecological stoichiometry may be useful in explaining their mechanisms of living in different microhabitats. In this study, Syntrichia caninervis, the dominant moss species of moss crusts in the Gurbantunggut Desert, China, was selected to examine the study of stoichiometric characteristics in three microhabitats (under living shrubs, under dead shrubs and in exposed ground). The stoichiometry and enzyme activity of rhizosphere soil were analyzed. The plant function in the above-ground and below-ground parts of S. caninervis is significantly different, so the stoichiometry of the above-ground and below-ground parts might also be different. Results showed that carbon (C), nitrogen (N) and phosphorus (P) contents in the below-ground parts of S. caninervis were significantly lower than those in the above-ground parts. The highest N and P contents of the two parts were found under living shrubs and the lowest under dead shrubs. The C contents of the two parts did not differ significantly among the three microhabitats. In contrast, the ratios of C:N and C:P in the below-ground parts were higher than those in the above-ground parts in all microhabitats, with significant differences in the microhabitats of exposed ground and under living shrubs. There was an increasing trend in soil organic carbon (SOC), soil total nitrogen (STN), soil available phosphorous (SAP), and C:P and N:P ratios from exposed ground to under living shrubs and to under dead shrubs. No significant differences were found in soil total phosphorous (STP) and soil available nitrogen (SAN), or in ratios of C:N and SAN:SAP. Higher soil urease (SUE) and soil nitrate reductase (SNR) activities were found in soil under dead shrubs, while higher soil sucrase (STC) and soil β-glucosidase (SBG) activities were respectively found in exposed ground and under living shrubs. Soil alkaline phosphatase (AKP) activity reached its lowest value under dead shrubs, and there was no significant difference between the microhabitats of exposed ground and under living shrubs. Results indicated that the photosynthesis-related C of S. caninervis remained stable under the three microhabitats while N and P were mediated by the microhabitats. The growth strategy of S. caninervis varied in different microhabitats because of the different energy cycles and nutrient balances. The changes of stoichiometry in soil were not mirrored in the moss. We conclude that microhabitat could change the growth strategy of moss and nutrients cycling of moss patches.     

Herbaceous vegetation response to controlled fire and postfire defoliation in a semiarid rangeland of Argentina

Semiarid rangelands of central Argentina are shaped by fire and grazing. The aim of this study was to quantify the effect of controlled fire on aboveground biomass production of (1) herbaceous species as a whole group and (2) herbaceous species subclassified as desirable or undesirable perennial grasses and as herbaceous forbs at the end of two growing seasons after an experimental fire in a semiarid rangeland of central Argentina. The effect of the length of the resting period after fire on aboveground biomass production of desirable perennial grass species was also assessed; treatments consisted in defoliations at the end of the first and second postfire growing seasons, at the beginning and at the end of the second postfire growing season and only at the end of the second postfire growing season. Total biomass production of the burned treatments was higher than that of the control treatment, the difference being attributed to an increase in desirable perennial grass species biomass. This in turn was negatively affected by defoliation at the end of the first postfire growing season, suggesting that the beneficial effect of fire on biomass production might have been overridden by the stress imposed by an early defoliation. Overall, the results of the present study suggest that grazing should be held back for at least a whole year after the fire to achieve the pursued aim of increasing forage production by the use of controlled fire.     

Auswirkungen einer Nachd��ngung im Sommer auf die Frosth?rte von Salix cinerea L. sowie auf ausgew?hlte N?hrstoffe und Biomarker

One year old rooted cuttings were fertilized with Osmocote exact according to 0,8 g N L^{? ? 1} substrate in February 2009. In August half of the plants received the same fertilizer dose while the other half was not refertilized. In autumn 2009 and in spring 2010 early resp. late frost tests were carried out under controlled conditions, furthermore sprouting was evaluated in spring 2010. Frost damage was measured with the electrolyte leakage method. In the early frost test biochemical stress responses like glucose, sucrose and proline were analyzed. In the new shoots N-, P-, and K-concentration were measured. Refertilized plants did not grow more but showed higher N-, P-, and K-concentrations. These plants had more proline in their buds and roots compared to the non-refertilized ones. Sucrose and glucose contents in the roots of refertilized plants were lower than in non-refertilized ones, in the buds there was no difference. Damage due to early frost was independent from refertilization for new shoots and buds but in roots higher for the refertilized plants. In spring 2010 refertilized plants sprouted earlier compared to non-refertilized ones. In buds damage due to late frost slightly increased in refertilized plants while in roots there was no difference between the fertilization treatments.