Seeding cool-season grasses into unimproved warm-season pasture in the southern Great Plains of the United States

Limited availability of herbage during the cool season creates a problem of a supply of nutrients for livestock producers throughout the southern Great Plains of the USA and, particularly, on small farms where resource constraints limit possible mitigating strategies. Six cool‐season grasses were individually sown into clean‐tilled ground, no‐till drilled into stubble of Korean lespedeza [Kummerowia stipulacea (Maxim) Makino] or no‐till over‐sown into dormant unimproved warm‐season pastures. The dry matter (DM) yields of mixtures of cool and warm‐season herbage species were measured to test their potential for increasing cool‐season herbage production in a low‐input pasture environment. Only mixtures containing Italian ryegrass (Lolium multiflorum Lam) produced greater year‐round DM yields than undisturbed warm‐season pasture with all establishment methods. When cool‐season grass was no‐till seeded into existing warm‐season pasture, there was on average a 0·61 kg DM increase in year‐round herbage production for each 1·0 kg DM of cool‐season grass herbage produced. Sowing into stubble of Korean lespedeza, or into clean‐tilled ground, required 700 or 1400 kg DM ha^?1, respectively, of cool‐season production before the year‐round DM yield of each species equalled that of undisturbed warm‐season pasture. Productive pastures of perennial cool‐season grasses were not sustained beyond two growing seasons with tall wheatgrass [Elytrigia elongata (Host) Nevski], intermediate wheatgrass [Elytrigia intermedia (Host) Nevski] and a creeping wheatgrass (Elytrigia repens L.) × bluebunch wheatgrass [Pseudoroegneria spicata (Pursh)] hybrid. Lack of persistence and low productivity limit the usefulness of cool‐season perennial grasses for over‐seeding unimproved warm‐season pasture in the southern Great Plains.     

Effects of reduced time on pasture caused by prolonged walking on behaviour and production of Mpwapwa Zebu cattle

In many African countries, cattle are often allowed to graze only during the daytime and the daily walk to pasture limits time for foraging. Two groups of cattle were allowed access to pasture for 9 h or 5 h associated with a daily walk of 0·5 h or 4·5 h respectively. Each treatment period lasted for 14 days in a cross‐over design. Observations of behaviour were carried out for 8 days in each treatment period. When only 5 h were allowed at pasture, the animals grazed less (0·61 vs. 0·66 of time) compared with the first 5 h of the 9 h treatment. The average daily milk yield was 323 g (0·078) higher when 9 h were allowed on pasture (P < 0·05 during the second week of treatments). There were no differences in milk yield after the end of the treatments. The daily liveweight gain was 250 g higher when the animals were on pasture for 9 h. These differences could be due to the limited time on pasture or to energy requirements for the walk to pasture. It is concluded that increasing the walking time and reducing time on pasture may compromise animal welfare and performance under the experimental conditions, i.e. on poor quality pastures in a warm humid climate where the main period at pasture is in the middle of the day.     

Variation in root traits associated with nutrient foraging among temperate pasture legumes and grasses

Temperate pasture legumes (e.g. Trifolium and Medicago spp.) often have a higher phosphorus (P) requirement for maximum productivity than pasture grasses. This is partly attributed to differences between legumes and grasses in their ability to acquire P from soil. We are the first to report differences in root morphology traits important for soil P acquisition in a range of novel pasture legumes being developed for use in temperate pastures of southern Australia. Up to a 3·6‐fold range in specific root length (SRL) (79–281 m root g^?1 root) and 6·1‐fold range in root hair length (RHL) (0·12–0·75 mm) was found between the pasture species. The commonly used Trifolium subterraneum and Medicago sativa had relatively low SRLs and short root hairs, while Ornithopus compressus, O. sativus and Biserrula pelecinus had RHLs and SRLs more similar to those of two grass species that were also assessed. Specific root length was highly correlated with average root diameter, and root traits were relatively stable at different plant ages. We surmise that large differences among pasture legume species in the effective volume of soil explored could translate into significant differences in their critical P requirements (i.e. soil P concentration to achieve 90% of maximum shoot yield).     

Effect of structure of a tropical pasture on ingestive behaviour, digestibility of diet and daily intake by grazing cattle

The effect of the structure of a tropical pasture, based on Dichanthium spp., on the ingestive behaviour, in vivo digestibility of the diet and herbage intake by eight Creole tethered heifers was studied. Two levels of nitrogen fertilizer (0 and 50 kg ha^?1) were applied to plots after each grazing cycle and there were 28 d between each of the three grazing cycles. Four heifers grazed individual subplots daily on each plot for 14 d in each of the successive grazing cycles. Simultaneous measurements of bite depth, bite mass, biting rate, short‐term intake rate and daily grazing time were made in two 4‐d periods at the end of each 14‐d period. The in vivo organic matter digestibility (OMD) and daily herbage organic matter intake (OMI, expressed on a kg LW^0·75 basis) were also measured at the same times. Relationships among pasture characteristics and ingestive behaviour were similar to those reported in other short‐term studies: pasture height was highly correlated with bite depth, bite mass and biting rate (r = 0·91, r =0·79 and r = ?0·68, respectively, P < 0·001). Pasture variables had lower correlations with grazing time and short‐term intake rate than with bite depth, bite mass and biting rate. Pasture structure was more highly correlated with OMD than OMI: leaf mass and length and also the extended tiller length were highly correlated with OMD (r = 0·77, r = 0·76 and r = 0·72, respectively, P < 0·001) whereas the crude protein concentration of the herbage was correlated with OMI and digestible OMI (r = 0·50 and r = 0·69, respectively, P < 0·001). Ingestive behaviour variables, as well as OMD, were correlated more with pasture characteristics than was OMI.     

Pasture use and management strategies in the Ankole pastoral system in Uganda

The Bahima ethnic group have been crossbreeding autochthonous Ankole with Holstein‐Friesian cattle. Separate herds (pure Ankole and crossbreds) are common. A survey was conducted to characterize pasture utilization and management in the Ankole region to gain a better understanding of current and emerging practices. Herds are largely grazed by continuous stocking although there are efforts to establish paddocks. Factors considered for pasture allocation to cattle genotypes were high‐quality feed, limited shrubs/weeds and close proximity to homestead for 80, 80 and 30% of farmers, respectively. Cross‐bred cattle were kept on medium/high‐quality, and Ankole cattle on medium/low‐quality pastures. Pastures for cross‐bred cattle yielded 0·17 t ha^?1 more dry matter than pastures allocated to Ankole cattle. Crude protein (CP) content of Ankole pasture was significantly (P < 0·05) lower than CP of cross‐bred pasture (6·3% compared with 7·3%). Neutral detergent fibre content was similar (72·8% vs. 69·8%) for both pasture types. Farmers’ perceptions of indicator plant species were important for allocation of pasture. Hyparrhenia rufa, Brachiaria spp., Themeda triandra and Chloris gayana were identified as the preferred pasture species. Sporobolus pyramidalis and Cymbopogon afronardus were the most undesirable pasture species. Brachiaria spp. offers an opportunity for utilization as hay for dry season supplementation.     

Effects on performance and carcass composition of turning beef cattle out to pasture early in the spring

Abstract The effects of turning beef cattle out to pasture earlier in the spring than is conventional in Northern Ireland on performance until slaughter in late summer were examined. Thirty‐eight steers, which were initially 525 kg live weight (LW), were divided into blocks of two according to breed type, previous plane of nutrition and LW and were allocated to two treatments at random within each block. Steers on Treatment 1 were turned out to pasture on 14 March and grazed a permanent perennial ryegrass‐based pasture with a mean sward surface height of 8·4 (s.e. ± 0·36) cm until Treatment 2 steers were turned out on 2 May. Thereafter, all steers grazed as a single group until slaughter at the same estimated carcass fat content, which occurred between June and September. Between the two turnout dates, steers on Treatment 2 were given grass silage ad libitum. For the early and late turnout treatments, respectively, LW gains between the two turnout dates were 1·5 and 0·7 kg d^?1, carcass gains from 14 March until slaughter (mean 143 d) were 0·69 and 0·53 (s.e. ± 0·022) kg d^?1, and carcass weights were 370 and 347 (s.e. ± 2·7) kg. It is concluded that, under the conditions of this experiment, turning beef cattle out to pasture in early spring may offer economic benefits in beef production in situations in which this is practicable.     

Evapotranspiration and plant–atmospheric coupling in a Brachiaria brizantha pasture in the Brazilian savannah region

The savannah of central Brazil occupies 208 million ha, of which about 54 million ha have been converted to cultivated pasture. The aim of this study was to provide the dynamics of evapotranspiration (ET) in a Brachiaria brizantha cultivated pasture in the Brazilian Savannah region at the beginning of the rainy season and determine biological and environmental factors that influence ET by using the decoupling approach. A meteorological station and an eddy covariance system were placed at the studied site. The above‐ground biomass production of the B. brizantha pasture responded rapidly to the onset of the rainy season. Leaf area index (LAI) increased from 0·4 to 1·1 between 15 November and 3 December, with a corresponding increase in live biomass. Average ET during the study period was 2·6 ± 0·9 mm. Daily ET was significantly correlated with maximum vapour pressure deficit and precipitation. The average of daytime decoupling factor obtained over 2 d without rain was 0·50 ± 0·06. The B. brizantha pasture deployed in this region showed an intermediate coupling which takes ET to be influenced by both atmospheric conditions and by the available radiation at the canopy.     

Effects of incorporating cowpea in a subtropical grass pasture on forage production and quality and the performance of cows and calves

The increasing cost of N fertilizer has stimulated an interest in sourcing protein from warm‐season legumes among beef cattle producers in the tropical/subtropical areas of the world. The objective of this study was to evaluate effects of two strategies of incorporating cowpea [Vigna unguiculata (L.) Walp.] into bahiagrass (Paspalum notatum Flügge) pastures on the herbage characteristics and performance of grazing cow–calf pairs. The study was conducted in Ona, Florida, USA, from May to August in 2007 and 2008. Experimental units were 1·0 ha. Treatments were bahiagrass pasture alone (control), 50:50 bahiagrass–cowpea pasture (cowpea), bahiagrass pasture with a cowpea creep grazing area (0·1 ha, creep grazing) and bahiagrass pasture with a creep‐fed concentrate [(creep feeding; 10 g kg^?1 body weight (BW)]. The cowpea pastures had lower herbage mass [HM, 1·8 vs. 3·7 t ha^?1] and herbage allowance [HA, 0·8 vs. 1·4 kg DM kg^?1 live weight (LW)] compared with the other treatments. Cowpea had greater CP (CP, 160 g kg^?1) and in vitro digestible organic matter (IVDOM), (600 g kg^?1) than bahiagrass (110 and 490 g kg^?1 respectively); however, cowpea HM was only 0·9 t ha^?1 in May and 0·7 t ha^?1 in June, but it did not persist in July and August. Calves receiving the creep feeding treatments had greater average daily gain (0·8 vs. 0·7 kg d^?1) than calves in other treatments. Further research is necessary to exploit the superior nutritive value of cowpea in grazing systems in the south‐eastern USA.     

Open pasture,silvopasture and sward herbage maturity effects on nutritive value and fermentation characteristics of cool‐season pasture

In Appalachian USA, silvopasture offers promise of increased farm productivity. A synchronized, temporal understanding of open pasture (OP) and silvopasture (SP) nutritive characteristics is essential for grazing system development. We examined pasture‐type nutritive‐value relationships when herbage was harvested based on morphological maturity rather than calendar date. Neutral detergent fibre and acid detergent fibre (ADF and NDF) content were greater in silvopasture, while organic matter (OM) was lower (P < 0·05). Digestibility of SP herbage dry matter (DM) and OM tended (P = 0·10) to be lower (418 vs. 471 and 437 vs. 491 g kg^?1 respectively). Neutral detergent fibre digestibility was greater (P < 0·05) for OP than SP forage (538 vs. 480 g kg^?1), and ADF tended to be greater (P = 0·10; 551 vs. 501). Open‐pasture forage fermentation effluent exhibited slightly higher microbial richness and Shannon diversity than SP. However, overall community composition of both bacteria and archaea did not differ between pasture types or sampling times. Pasture types show proximate analyses differences generally favourable to OP, although both have similar overall nutritive value. In addition, the SP sward exhibited a 4‐ to 6‐days delay in reaching equivalent maturity. Conversion of farm woodlots to SP would increase overall herbage production and improve pasture management flexibility.     

Nitrogen recovery and loss in a fertilized elephant grass pasture

Limited information is available regarding the recovery and loss of fertilizer nitrogen (N) applied to intensively managed tropical grass pastures. An experiment was carried out in Brazil to determine the fertilizer‐N recovery and ammonia volatilization loss in an elephant grass (Pennisetum purpureum, Schum.) pasture fertilized with 100 kg N ha^?1 as urea or ammonium sulphate, labelled with ¹⁵N, in late summer (LS) or in mid‐autumn (MA). Herbage mass was highest and litter mass was lowest in LS (P < 0·05). The N concentration of herbage was highest in autumn (P < 0·05) and the total N content in soil was lower in LS than in MA (P < 0·05), reflecting the high N uptake capacity of the grass. Proportionately higher ¹⁵N recovery in litter mass (P < 0·05) was observed in autumn (0·094) than in LS (0·0397) and the ¹⁵N recovery in herbage was 0·046 higher for ammonium sulphate‐fertilized pastures (P < 0·05; proportionately 0·243 for ammonium sulphate and 0·197 for urea). Around 0·60 of the fertilizer‐¹⁵N recovered was retained in soil and in non‐harvestable fractions of the plant. The NH₃ volatilization loss was higher in LS and most of the N loss occurred soon after fertilizer application. Urea and ammonium sulphate fertilizers were equally effective in sustaining herbage dry matter yield in the short term. However, the use of ammonium sulphate, rather than urea, would be preferable for LS applications when the objective is to reduce NH₃ volatilization losses.     

Proximal sensing of the seasonal variability of pasture nutritive value using multispectral radiometry

The nutritive value of pasture is an important determinant of the performance of grazing livestock. Proximal sensing of in situ pasture is a potential technique for rapid prediction of nutritive value. In this study, multispectral radiometry was used to obtain pasture spectral reflectance during different seasons (autumn, spring and summer) in 2009–2010 from commercial farms throughout New Zealand. The analytical data set (n = 420) was analysed to develop season‐specific and combined models for predicting pasture nutritive‐value parameters. The predicted parameters included crude protein (CP), acid detergent fibre (ADF), neutral detergent fibre (NDF), ash, lignin, lipid, metabolizable energy (ME) and organic matter digestibility (OMD) using a partial least squares regression analysis. The calibration models were tested by internal and external validation. The results suggested that the global models can predict the pasture nutritive value parameters (CP, ADF, NDF, lignin, ME and OMD) with moderate accuracy (0·64 ≤ r² ≤ 0·70) while ash and lipid are poorly predicted (0·33 ≤ r² ≤ 0·40). However, the season‐specific models improved the prediction accuracy, in autumn (0·73 ≤ r² ≤ 0·83) for CP, ADF, NDF and lignin; in spring (0·61 ≤ r² ≤ 0·78) for CP and ash; in summer (0·77 ≤ r² ≤ 0·80) for CP and ash, indicating a seasonal impact on spectral response.     

Pasture‐based dairy farm systems increasing milk production through stocking rate or milk yield per cow: pasture and animal responses

A 2‐year whole‐farm study compared pasture‐based systems increasing milk production per ha by increasing either stocking rate (from 2·5 to 3·8 cows ha^?1) or milk yield per cow (from 6000 to 9000 kg cow^?1 lactation^?1) or both. Four treatments (systems), comprising 30 cows each, were compared under the same management and grazing decision rules. The diet was based on grazed pasture, whereas pellets and conserved fodder were fed when deemed necessary. Milk production per ha increased by 0·49, 0·1 and 0·66 in the systems that increased either stocking rate, milk yield per cow or both respectively. Cows in the ‘high milk yield per cow’ systems had a significantly higher body condition score throughout the lactation, but reproductive performance was similar among all groups. Total pasture utilized (11 t DM ha^?1 year^?1) and pasture nutritive value were similar across all systems. This was associated with the grazing rules applied and the ability of accurately supplementing to meet deficits in available pasture. At the whole‐system level, there was a higher marginal efficiency of supplement use when increasing stocking rate than when increasing milk yield per cow or increasing both (0·18, 0·07 and 0·12 kg milk MJ^?1 of metabolizable energy of supplements respectively).     

Diet selection of dry and lactating beef cows grazing extensive pastures in late autumn

The diet selection, made by twelve dry and twelve lactating spring‐calving Salers beef cows at the end of the grazing season on an extensively grazed natural mountain pasture in central France, was compared. Sward structure, selection bites, daily grazing time and digestibility of the diet were measured in mid‐October (P₁) and mid‐November (P₂). Live weights were measured twice monthly throughout the experiment. Daily milk production was assessed in the week before the grazing behaviour measurements. Lactating cows grazed more selectively than dry cows: they took more bites on green patches (0·20 vs. 0·13 of total bites, P < 0·01; Jacob's selectivity index: 0·13 vs. ?0·25, P < 0·001), which is consistent with the higher overall nitrogen concentration in faeces (18·1 vs. 17·3 g kg^?1 DM, P < 0·05). Lactating cows tended to slightly increase their daily grazing time compared with dry cows (on average 9·9 vs. 9·5 h, P = 0·07) and grazed faster in mid‐November (74 vs. 69 bites min^?1, P < 0·05). The lactating cows lost more live weight (?24 vs. ?12 kg cow^?1, P < 0·05) between P₁ and P₂. A positive correlation was found between consumption of green patches by lactating cows and their daily milk production in P₂ (r = 0·574, P = 0·05), whereas no correlation occurred between individual milk production and liveweight loss. Dry cows were less selective, which supports management practices that promote the use of cows with low requirements on extensively grazed pastures in late season.     

Effects of the proportions of high or medium digestibility grass silage and concentrates in the diet of beef cattle on liveweight gain, carcass composition and fatty acid composition of muscle

Abstract An experiment was carried out over 2 years to evaluate the effects of increasing the proportion of cereal‐based concentrates in diets containing high‐digestibility and conventional medium‐digestibility grass silages on the dry‐matter (DM) intake, liveweight gain and carcass composition of beef cattle, and to examine the effects of grazed grass and the ratio of grass silage:concentrates in the diet on the fatty acid composition of selected muscle tissues. Late‐maturing steers (n = 231) were offered diets based on high‐digestibility (HD) (0·743 digestible organic matter (DOM) in DM) or medium‐digestibility (MD) (0·643 DOM in DM) grass silages supplemented with barley/soyabean meal‐based concentrates. The concentrates constituted 0·20, 0·40, 0·60 and 0·80 of total DM of the diets, which were offered ad libitum (AL). The two diets, which contained 0·80 concentrates, were also offered at 0·80 of AL intake. A further group of fourteen animals were given the medium‐digestibility silage only for 5 months and then grazed perennial ryegrass pastures for a further 5 months (silage/pasture treatment). For the diets containing HD silage and 0·20, 0·40, 0·60 and 0·80 concentrate, and 0·80 concentrate at 0·8 of AL intake, the DM intakes were 9·4, 10·2, 10·4, 10·2 and 8·1 (s.e. 0·16) kg d^?1, respectively, and daily carcass gains were 0·67, 0·78, 0·77, 0·79 and 0·62 (s.e. 0·029) kg d^?1, respectively; for those containing MD silage and 0·20, 0·40, 0·60 and 0·80 concentrate, and 0·80 concentrate at 0·8 of AL, the DM intakes were 8·2, 9·3, 10·1, 10·1 and 8·0 (s.e. 0·16) kg d^?1, respectively, and daily carcass gains were 0·38, 0·48, 0·64, 0·77 and 0·56 (s.e. 0·029) kg d^?1 respectively. Increasing the proportion of concentrates in silage‐based diets decreased the concentration of omega‐3 (ω‐3) polyunsaturated fatty acids (PUFA) (P < 0·001) and increased the concentration of ω‐6 PUFA (P < 0·001) in muscle. Cattle on the silage/pasture treatment had the highest concentration of ω‐3 PUFA in muscle (51 g kg^?1 lipid), this value being over three times that for animals given diets containing MD silage and 0·80 concentrate in the diet. These results demonstrate the potential of HD silage made from perennial ryegrass relative to high concentrate diets. The consumption of pasture‐finished beef could make a significant contribution towards increasing the intake of ω‐3 PUFA in the human diet.     

Effects of method, rate and timing of slurry application to grassland on the preference by cattle for treated and untreated areas of pasture

Two experiments were conducted in the UK to investigate preference by cattle for areas of pasture treated or untreated with dairy cow slurry. Experiment 1 examined the effects of method and rate of slurry applied to a grass pasture in March (Spring); Experiment 2 examined the effects of method and timing of slurry application after cutting for silage in June (Summer). In both experiments, beef steers grazed grassland plots that had been treated with slurry on one half only of the plots for 4 weeks. Slurry was applied by shallow injection (I), trailing shoe (T) or conventional surface broadcasting (S) techniques. In Experiment 1, slurry was applied at 40 m³ ha^?1 (HI) or 20 m³ ha^?1 (LO) 21 d before grazing commenced; in Experiment 2, slurry was applied at 20 m³ ha^?1 either 32 d (E) or 10 d (L) before grazing commenced. Sward height was measured and animal behaviour was recorded during 8 × 24 h measurement periods in each experiment. Preference for slurry‐treated or untreated swards was indicated by a comparison of post‐grazing sward height on each sward and the proportion of total grazing time spent on the slurry‐treated swards. In Experiment 1, a smaller proportion of grazing time was spent on slurry‐treated areas of swards compared with untreated areas of swards on all treatments, but the proportion of time spent grazing the slurry‐treated sward was greater on method T than on methods I or S (0·28, 0·31 and 0·39 for methods S, I and T, respectively, s.e.d. = 0·035, P < 0·05). Steers showed no preference between slurry‐treated and untreated swards by 49, 45 and 44 d after slurry application for methods S, I and T respectively. Differences between HI and LO were not significant (P > 0·05). In Experiment 2, aversion to slurry‐treated swards compared with untreated swards was less for treatment E than for L, with a greater proportion of both residence and grazing time spent on the slurry‐treated sward of treatment E than L (0·43 and 0·28, respectively, for residence time, s.e.d. = 0·025, P < 0·001, and 0·47 and 0·30, respectively, for grazing time, s.e.d. = 0·021, P < 0·001). Significant differences were shown between the methods of slurry application for treatment E, with a greater proportion of grazing time spent on the slurry‐treated swards of methods I and T than method S (0·40, 0·52 and 0·49 for S, I and T, respectively, s.e.d. = 0·032, P < 0·001), and a greater difference between post‐grazing heights on slurry‐treated and untreated swards for method S than for I or T (30, 12 and 15 mm, respectively, s.e.d. = 4·8, P < 0·001). For treatment L, differences in the proportion of time spent grazing on slurry‐treated swards compared with untreated swards were not significant between treatments (0·26, 0·34 and 0·31 for methods S, I and T, respectively, s.e.d. = 0·040, P > 0·05) and indifference between slurry‐treated and untreated swards was achieved 77, 60 and 58 d after slurry spreading for methods S, I and T respectively. These results show that, when slurry is applied to pasture in a grazing rotation, the adverse effects on grazing behaviour associated with conventional surface broadcasting may be overcome by the use of novel slurry application techniques.     

‘MaxClover’ grazing experiment: I. Annual yields,botanical composition and growth rates of six dryland pastures over nine years

Six dryland pastures were established at Lincoln University, Canterbury, New Zealand, in February 2002. Production and persistence of cocksfoot pastures established with subterranean, balansa, white or Caucasian clovers, and a perennial ryegrass‐white clover control and a lucerne monoculture were monitored for nine years. Total annual dry‐matter (10.0–18·5 t DM ha^?1) and sown legume yields from the lucerne monoculture exceeded those from the grass‐based pastures in all but one year. The lowest lucerne yield (10 t ha^?1 yr^?1) occurred in Year 4, when spring snow caused ungrazed lucerne to lodge and senesce. Cocksfoot with subterranean clover was the most productive grass‐based pasture. Yields were 8·7–13·0 t DM ha^?1 annually. Subterranean clover yields were 2·4–3·7 t ha^?1 in six of the nine years which represented 26–32% of total annual production. In all cocksfoot‐based pastures, the contribution of sown pasture components decreased at a rate equivalent to 3·3 ± 0·05% per year (R² = 0·83) and sown components accounted for 65% of total yield in Year 9. In contrast, sown components represented only 13% of total yield in the ryegrass‐white clover pastures in Year 9, and their contribution declined at 10·1 ± 0·9% per year (R² = 0·94). By Year 9, 79% of the 6.6 t ha^?1 produced from the ryegrass‐white clover pasture was from unsown species and 7% was dead material. For maximum production and persistence, dryland farmers on 450–780 mm yr^?1 rainfall should grow lucerne or cocksfoot‐subterranean clover pastures in preference to ryegrass and white clover. Inclusion of white clover as a secondary legume component to sub clover would offer opportunities to respond to unpredictable summer rainfall after sub clover has set seed.     

Soil dwelling macro‐invertebrates in intensively grazed dairy pastures in Pennsylvania,New York and Vermont

This study estimates the relative contributions of environment and farm management strategies in influencing soil faunal assemblages and attempts to identify the species with potential to affect sustainability of intensive grazing management systems in the north‐eastern USA. It arises because of the change from confinement feeding of dairy cattle, consequent upon concerns about negative environmental effects, the rising costs for machinery and housing, and reduced profit margins, together with the absence of data from which the consequences of such change on the soil fauna may be predicted. Macro‐invertebrates were sampled in soil from seventy‐eight grazed pastures on twenty‐one dairy farms in Pennsylvania, USA, in the spring of 1994. On five of these farms, macro‐invertebrates were sampled (four pastures per farm) in the spring, summer and autumn seasons of 1994, 1995 and 1996. In 1997, macro‐invertebrates were sampled in soil during spring, summer and autumn from (four pastures per farm) on three farms in New York, and during spring and summer on three farms in Vermont. Species richness ranged from two to twelve species (mean 6·4) per pasture site in Pennsylvania and five to eighteen species (mean 10·7) in New York and Vermont. The communities were dominated at most sites by earthworms. Earthworms were correlated with soil basal and substrate‐induced respiration/carbon ratio, and soil moisture, but were negatively correlated with cows per hectare and herbage biomass in Pennsylvania. Sitona larvae were recorded at nineteen of the twenty‐one farms during the spring of 1994 across Pennsylvania and occurred at populations >5 m^?2 in 68% of the sampled pastures. Sitona larvae were less abundant in New York and Vermont. Elaterid larvae comprised a complex of seven species of which Aeolus melillus (Say) and Melanotus communis (Gyllenhal) comprised 35% and 39%, respectively, of the elaterids collected in Pennsylvania. Agriotes mancus (Say) and Ctenicera destructor (Brown) comprised 41% and 26%, respectively, of four species collected in New York and Vermont. Scarabaeid larvae, comprising a complex of eight species, were detected at only 27% of the seventy‐eight pastures sampled in spring 1994 in Pennsylvania. Five species were collected in ten of the twelve New York pastures and four species in nine of the twelve Vermont pastures. Populations of scarabaeid larvae averaged <25 m^?2 in all three states, except in three Pennsylvania pastures in spring 1994. Detrended canonical correspondence analysis (DCCA) showed pasture standing biomass, legume diversity, pre‐winter stubble height, white clover pasture content, and soil phosphorus levels influenced numbers of invertebrate species more than climatic factors, such as temperature, rainfall, altitude, latitude and seasonal water table. DCCA also showed most pastures to be close to the average of environmental factors. The extremely low density of herbivorous macro‐invertebrates in soil and the absence of pest outbreaks may indicate a stable soil ecosystem.     

Consequences for biodiversity of reducing inputs to upland temperate pastures: effects on beetles (Coleoptera) of cessation of nitrogen fertilizer application and reductions in stocking rates of sheep

Current policies for upland pasture management in the UK encourage the integration of environmental objectives with livestock production through extensification of grazing systems. This study tested the hypothesis that a greater sward height in the summer would increase the diversity and abundance of grassland beetles (Coleoptera) as has been demonstrated for insects of indigenous grasslands. The hypothesis was tested with an experiment on an upland sheep pasture in mid‐Wales. Experimental treatments received different nitrogen fertilizer inputs (0 or 50 kg ha^?1), sheep stocking densities (12 or 9 ewes ha^?1) and average sward heights in summer were constrained to 3·5 or 5·5 cm by conserving surplus grass for silage in subplots. Five treatments, replicated in three randomized blocks, combined the two stocking densities and two sward heights without nitrogen fertilizer inputs, with the fifth combining the higher stocking density, shortest sward height and the nitrogen fertilizer input. Beetles were sampled with twelve pitfall traps in each of the fifteen plots from June to September in 1993 and 1995. In years 1 (1993) and 3 (1995) of the experiment, more Coleoptera species occurred in the tall sward (an average of nine species in addition to the forty‐one species present in the sward with the conventional sward height). Continuously grazed as opposed to ensiled subplots supported more beetle species but fewer individuals. Species composition of ground (Carabidae) and rove (Staphylinidae) beetles varied between treatments more than the arithmetic differences in species number. The experimental results supported the hypothesis but the benefits of taller swards to species diversity were small in the sown pastures of the study compared with indigenous upland grasslands (c. 33% fewer species). Inheritance effects of drainage, fertilizer and lime inputs, and the different species and management of cultivated pastures, may constrain the conservation benefits of altered pasture management compared with indigenous grasslands.     

Changes in plant population density, composition and sward structure of a hill pasture during a pastoral fallow

Appropriate pre-sowing methods for the introduction of improved forage legume and grass germplasm are an important issue for hill pasture improvement in New Zealand. A pastoral fallow, which involves not defoliating pasture for a period generally from late spring/early summer to autumn, could create a potentially favourable environment for introducing improved germplasm. A field study was conducted on two aspects (shady and sunny) of moist, low-fertility hill country with or without added fertilizer (phosphorus and sulphur) in the southern North Island of New Zealand, to investigate the changes in plant population density and sward structure during a full or partial pastoral fallow, compared with a rotationally grazed pasture. A 7-month (October to May) pastoral fallow dramatically decreased the densities of grass tillers by 72% (P < 0·01), white clover (Trifolium repens L.) growing points by 87% (P < 0·01) and other species by 87% (P < 0·05). The decline in tiller density by pastoral fallow was enhanced on the shady aspect. Fertilizer application increased white clover growing-point density on the shady aspect (P < 0·05) and grass tiller density on the sunny aspect (P < 0·05). Decreased plant density during pastoral fallowing was attributed to aboveground biomass accumulation, which altered sward structure, leading to interplant competition and mortality by self-thinning and completion of the life cycle of some matured plants. The plant size-density relationship during pastoral fallowing in this mixed-species sward followed the serf-thinning rule, particularly when the calculation was based on all plant species rather than grass alone. There was no significant (P > 0·05) difference in final plant population density between the 7-month pastoral fallow and a shorter term (October to December) pastoral fallow. It is concluded that pastoral fallowing effectively reduced the plant population density and altered sward structure of a hill pasture. Such changes create a more favourable environment for the introduction of improved forage species.