Cherrybark oak stump sprout survival and development five years following plantation thinning in the lower Mississippi alluvial valley,USA

Cherrybark oak (Quercus pagoda Raf.) stump sprouts were studied for 5 years in a 30-year-old plantation thinned to 70–75% stocking (light thinning) and 45–50% stocking (heavy thinning). Sprouting success, survival, number of sprouts per stump, and sprout height differed little between thinning treatments throughout the 5-year study period. Pre-harvest tree d.b.h. also had no influence on sprout survival and development. A 2-year drought reduced survival and may have influenced sprout development. Sprout clump survival dropped from 90% 1 year following thinning to 46% 3 years after thinning. Although sprout height averaged 337 cm 5 years after thinning, annual sprout growth decreased from 166 cm the first year after thinning to 33 cm in each of the last 2 growing seasons. Results indicated that bottomland hardwood regeneration evaluation models may underestimate the potential of oak stump sprouts to contribute to pre-harvest regeneration assessments. Further study in the role of stump sprouts to regenerate bottomland oak species is needed. The U.S. Government's right to retain a non-exclusive, royalty-free license in and to any copyright is acknowledged     

辽东栎伐桩萌苗的发育规律

通过对黄土高原黄龙山林区辽东栎林固定样地连续6年的调查,研究间伐强度分别为13.4％(弱度间伐)和30.0％(强度间伐)的辽东栎伐桩萌苗的存活与生长规律.结果表明:1)间伐第1年,辽东栎伐桩均有萌生枝发生,每桩萌生枝数量和萌生枝高度在不同强度间无显著差异;2)萌生枝数量受伐桩高度和直径的影响,当伐桩高度较低时,萌生枝数量随伐桩高度的增加而增加,超过20cm后随之减少,呈三次抛物线关系;伐桩萌生枝数量随着伐桩直径增加呈现先增加后减少的趋势,直径30～50cm的伐桩萌生枝数量最多;3)伐桩萌生枝数量与最长萌生枝高度呈显著线性负相关,存在“萌生枝数量——高度权衡”现象;4)间伐第6年,伐桩的存活率为97.01％,强度间伐区萌生枝高度显著高于弱度间伐区;5)调查期间,辽东栎伐桩萌生枝数量逐年下降,伐桩产生的萌生枝数量越多,后期萌生枝死亡的数量也越多,表明伐桩能够自我稀疏,存在“萌代主”现象.在黄土高原地区,辽东栎伐桩萌芽潜力很大,应合理利用;间伐时伐桩高度应尽量低于20cm;伐桩直径在30～50cm更有利于其无性更新;间伐后伐桩应及时除蘖定株,促进萌生枝生长;间伐强度30.0％(强度间伐)更有利于辽东栎伐桩萌生枝的生长.     

Potential long-term growth gains from early clump thinning of coppice-regenerated oak stands

A study was established in the Missouri Ozarks to evaluate coppice regeneration of oak. Five years after a 32-year-old stand was clearcut and regenerated naturally through stump sprouts, the dominant sprout on each stump was identified based upon height. Treated plots were thinned to the single dominant sprout on each stump whereas control plots were not thinned. Twenty-five years later the largest 247, 371, 494, and 618 stems per ha were examined and height of the dominant sprout at age 5 was found to be strongly related to dbh at age 30 in both thinned and unthinned plots. However, in the thinned plots, the largest 494 stems per ha were on average 11%, 28%, and 58% greater, respectively, in height, dbh, and volume compared to similar dominant sprouts in unthinned plots. Logistic regression analysis was used to develop curves for the evaluation of potential gains from clump thinning. In this analysis, the average height of a stand at age 5 was used to estimate thinning gains at age 30.     

Development of water oak stump sprouts under a partial overstory

A 28-year-old water oak (Quercus nigra L.) plantation was thinned from below to either 254 or 462 stems per hectare to determine the influence of a partial canopy on oak stump sprout development. Sprout clump survival, number of living sprouts in a clump, and height and DBH of the dominant sprout in a clump were measured in years 1–5 and 7 after harvest. By year 7, sprout clump survival under the heavily thinned canopy was 23% higher than under the lightly thinned canopy. Within-clump sprout mortality was not influenced by overstory thinning level, but by year 2 height and diameter increment were 15% and 22% greater, respectively, under the heavy overstory removal. Positive height and diameter growth of dominant sprouts continued under both canopy conditions through year 7, but early benefits of the heavy overstory removal on sprout growth diminished. Additional overstory removal or sprout clump thinning would be necessary to sustain sprout clump survival and sprout growth over extended periods.     

林龄、采伐方式对大叶相思萌芽更新的影响*

在海南省琼海县进行了不同林龄、不同采伐方式对大叶相思萌芽更新影响的研究，同时比较了保留不同萌条数量的萌芽林生长表现。结果表明：（１）林龄对伐桩萌芽率没有显著影响，但对伐桩存活率及萌条的径、高生长影响显著或极显著。林龄越大，萌芽更新效果越差；（２）皆伐和隔行采伐对大叶相思伐桩的萌芽率、萌条数量和存活率均无显著影响，但对萌条的径、高生长影响极显著。试验证明对大叶相思林分施以隔行采伐通过萌芽更新建立复层     

林龄、采伐方式对大叶相思萌芽更新的影响

在海南省琼海县进行了不同林龄、不同采伐方式对大叶相思萌芽更新影响的研究，同时比较了保留不同萌条数量的萌芽林生长表现，结果表明：（１）林龄对伐桩萌芽率没有显著影响，但对伐桩存活率及萌条的径、高生长影响显著或极显著。林龄越大，萌芽更新效果越差；（２）皆伐和隔行采伐对大叶相思伐桩的萌芽率、萌条数量和存活率均无显著影响，但对萌条的径、高生长影响极显著。试验证明对大叶相思林分施以隔行采伐通过萌芽更新建立复层林分是可行的；（３）保留不同数量萌条对萌芽林早期（１．５年生）的径、高生长有极显著影响，但对后期（４．５年生）的生长影响不显著。每棵植株伐桩保留４根萌条的萌芽林生物产量最高，更新效果最好。     

不同伐桩直径及高度对马占相思萌芽更新影响的研究*

对速生多用途树民占相思进行了不同伐桩高度（25cm,50cm,75cm和100cm)及不直径（4．1－8．0cm和8．1－12．0cm)萌芽更新影响的研究，结果表明，伐桩的萌芽率及平均萌条数随伐桩高度的增加而增加，随直径的增大而减少，萌条直径，长度及直萌条（可望生长成材之萌条）总断面积随伐桩的高低和大小而增减，伐桩最低的萌芽率比以往的结果高4倍以上，以短轮伐期经营的马占相思人工林，采用50－75cm伐桩高度萌芽更新效果好。     

Influence of thinning time and density on sprout development, biomass production and energy stocks of sawtooth oak stumps

The high potential values of sawtooth oak (Quercus acutissima) in fuelwood or bioenergy are recognized. Sprouting ability, sprout growth, biomass production and energy stocks in coppiced plantations of sawtooth oak were evaluated at the Hongya Mountain Forest Farm in Anhui Province, China. Experimental treatments applied in a split-plot design included three sprout thinning times and four sprout numbers reserved on each stump (1 sprout, 2 sprouts, 4 sprouts stump⁻¹ and check). Sprout growth and biomass production per stump were significantly affected by the treatments and a significant positive relationship between stump basal diameter and sprout numbers produced was observed. After the third growing season, the highest total sprout biomass per stump was achieved in the treatment with thinning excess sprouts at the end of first growing season (December, 2007) and reserving 4 dominant sprouts per stump (T12S4, reaching 8.67 kg stump⁻¹), while the lowest was found in the treatment with thinning the sprouts in August of the first growing season and reserving 1 dominant sprout per stump (T8S1, only 3.40 kg stump⁻¹). Different treatments also influenced gross calorific values (GCV) of the components sampled from 3-year-old sprouts and the mean GCV of stem wood on an oven-dry weight basis was within the range of 18.45 ± 0.15 and 18.83 ± 0.12 kJ g⁻¹. Similar to the sprout biomass production, the greatest total and stem energy stocks per stump were observed in T12S4 treatment, achieving 161.6 and 110.5 MJ stump⁻¹, respectively. Based on the results from this study, thinning excess sprouts to reserve 4 sprouts per stump as early as age 1 could be proposed for the management of sawtooth oak coppice with cutting cycle of 3-5 years and stand density of 5000-6000 stump ha⁻¹.     

The relative importance of different regeneration mechanisms in a selectively cut savanna-woodland in Burkina Faso,West Africa

Savanna plant species are capable of regenerating both sexually and asexually. The importance of each regeneration mechanism depends on the species, the type and the intensity of disturbance. This paper describes the relative importance of sexual and asexual regeneration for the restoration of a selectively cut savanna woodland in Burkina Faso. Plantlets regenerated sexually or asexually were determined by excavating the below ground system and assessing basal and aerial sprouts within 144 quadrats along transects in 2, 5, 10 year-cut and control plots. The result shows that seedling sprouts were predominant (83%) compared to root sucker (4%), coppice (5%), water sprout (2%) and layering (less than 1%). True seedlings constituted a minor component (5%) of the plantlet population. A total of 54 woody species was found in the plantlet population with 52 presenting seedling sprouts. Twelve species were found regenerating from both seeds and root buds; of which Dichrostachys cinerea, Pteleopsis suberosa and Detarium microcarpum had the highest number of plantlets. Feretia apodanthera was the only layering species encountered in the sampled area. Coppices, root suckers and water sprouts showed higher height and bigger collar diameter compared to seedling sprouts and true seedlings. It can be concluded that sexual reproduction is the dominant mechanism of seedling recruitment in this disturbed savanna-woodland, and that its success relies on the ability of plantlets of seed origin to resprout abundantly.     

Characteristics of lowland broadleaved woodland being restocked by natural regeneration

A survey of 78 sites in southern England with approved managementplans for restocking by natural regeneration was made duringthe summers of 1993 and 1994. The following features were recorded:species, stocking, canopy cover and seed-bearing potential oftrees present in the overstorey; species and canopy cover ofthe understorey; ground cover; species, browsing damage, numberand heights of tree seedlings. In general, sites were poorlystocked with overstorey trees having an average of 135 stemsha^–1 and a mean canopy cover of 36 per cent. Similarly,the understoreys were poorly developed with an average coverof only 23 per cent. Twenty-nine species of tree were foundin the overstorey, Quercus spp. and Fraxinus excelsior werethe most common. Many of the stems present had poor seed-bearingpotential. Cover of the ground flora often exceeded 75 per cent.Seedlings were present on most sites, with F. excelsior andBetula pendula being most abundant with mean seedling densitiesexceeding 10000 ha^–1. Most seedlings were >20 cm tall,few exceeded 120 cm. About 30 per cent of seedlings were browsed.The results are related to current guidance and the future prospectsfor use of natural regeneration.     

Survival and development of Liaodong oak stump sprouts in the Huanglong Mountains of China six years after three partial harvests

The survival and development of Liaodong oak (Quercus liaotungensis Koidz) stump sprouts were studied for 6 years under leave-tree, as well as 60 and 75% crown closure uniform shelterwood in the Huanglong Mountains of China. Stump sprout survival rate, number of sprouts, and sprout height under the three partial harvests during the first year post-harvest did not differ. Six years post-harvest, 78.32% of the stumps had living sprouts. No variation in sprout survival was found among the treatments throughout the study period. The number of living sprouts per stump was 13.77 ± 0.50 in the first growing season after harvest, and dropped to 6.49 ± 0.22 after 6 years. This result indicated the occurrence of self-thinning within the sprout clumps. Moreover, the height of dominant sprout increased from 52.88 ± 1.62 cm (in the first year) to 132.82 ± 2.35 cm (in the sixth year) post-harvest. Sprout height in the leave-tree harvest type was higher than those in the two shelterwoods in the fifth and sixth years after logging. Annual height growth was approximately 8–96 cm during the first growing season, and then decreased at a rate of 1–44 cm per year in the following years. A negative relationship was found between sprout height and the number of sprouts per stump. This result indicated a trade-off between the number of sprouts and height growth. The number of sprouts per stump decreased by about 0.2 stem for each 1 m²/ha increased in residual basal area. Therefore, sprouting may play an important role in the restoration of the secondary forest after harvesting.     

Precommercial thinning of beech (Fagus sylvatica L.): Early effects of stump height on growth and natural pruning of potential crop trees

Abstract

This paper reports the early effects of stump height on the growth and natural pruning of potential crop trees after precommercial thinning of a young stand of naturally regenerated beech (Fagus sylvatica L.) in Denmark. The experiment comprises five treatments based on combinations of three grades of precommercial thinning and three stump heights. Treatments include the unthinned control, thinning only of whips and wolf trees (using low stumps), and thinning for potential crop trees using low, medium or high stumps. All treatments were replicated three times. Stump heights averaged 10, 90 or 230 cm, respectively. Pretreatment stem number (live trees) varied from 17,500 to 41,000 ha^?1. In the potential crop tree treatment, post-treatment stem number ranged from 4750 to 9500 ha^?1. Following two growth seasons, the quantity of stump regrowth increased with increasing stump height, the rate of stump regrowth increased with increasing stump height, the diameter growth of potential crop trees increased with decreasing stump height, the increase in stand height did not depend on stump height or post-treatment stem number, and the natural pruning of potential crop trees increased with increasing stump height. It remains to be seen whether these trends hold in the long run, and whether additional economic return from the increase in wood quality with increasing stump height compensates for the reduction in diameter growth.     

Understory light predicts stump sprout growth in mixed multiaged stands in north coastal California

Regeneration of commercial species is central to long-term success of multiaged management for wood production. We examined relationships between understory light, varying overstory tree retention, and growth of coast redwood (Sequoia sempervirens; commercial species) and tanoak (Notholithocarpus densiflorus) stump sprouts initiated by group selection and single-tree selection harvesting in 80–100 year old mixed stands at four sites. Treatments included a complete harvest in 1-ha group selection openings, low-density dispersed retention, and either aggregated or dispersed high-density retention. Post-harvest stand density index and basal area were useful predictors of understory light. Mean and maximum understory light did not differ significantly between treatments with the same density where residual trees were retained in aggregated versus dispersed spatial patterns. However, the dispersed retention had lower minimum light levels when compared to the aggregated retention treatment. Aspect appeared to influence understory light more in dispersed treatments. At all light levels, the dominant sprout within clumps of redwood stump sprouts generally grew faster than dominant tanoak sprouts within tanoak sprout clumps. Differences in sprout height growth between aggregated and dispersed treatments were minimal. Stump size had a significant effect on redwood stump sprout height growth, with sprouts on the largest stumps growing approximately twice as quickly as sprouts on the smallest stumps. In the low density dispersed treatment, redwood sprouts outperformed tanoak sprouts by the greatest margin. Regeneration of redwood and tanoak was most rapid within group selection openings.     

章古台沙地赤松天然更新的初步研究

通过2年的野外调查与比较分析,认为在章古台沙地赤松天然更新是可行的。天然更新方式有3种,即林下更新、林隙更新和林缘更新。天然更新幼树在空间上呈带状、群团状分布;更新幼树年龄在时间上是连续的,降水量大的2004、2005、2008年更新幼树相对较多;更新幼树密度为20 300~100 000株.hm-2。赤松林缘天然更新幼树长势要好于林隙,林隙要好于林下。所以,要充分利用各种条件,进行天然更新和人工促进天然更新。     

Sprouting characteristics of <Emphasis Type="Italic">Quercus pubescens</Emphasis> Willd. and <Emphasis Type="Italic">Quercus frainetto</Emphasis> Ten. trees after a wildfire: To cut or not to cut the burned trees?

The aims of this study were to analyze the sprouting characteristics of Quercus pubescens and Q. frainetto trees after a wildfire and to recommend the appropriate silvicultural treatments in order to achieve dense sprout origin regeneration. This study was carried out in the area that was burned in a wildfire in northeastern Greece. Three years later, 140 Q. pubescens trees and 140 Q. frainetto trees were selected. Seventy trees for each species were individuals that were cut after the fire and were in the form of stumps, while 70 trees were intact top-killed trees, as a result of fire. In each stump and intact tree that was selected, various parameters were measured or counted: the diameter at the level of ground, the number of the developed sprouts (if any) and the height as well as the basal diameter of the tallest sprout, etc. Both species have similar sprouting behavior after a wildfire. Mother tree dimensions positively influenced the number of sprouts and the growth of the tallest sprout per individual. The mean height of the tallest sprout per stump, after three growing seasons from the forest fire, is high in both species (2.08 m for Q. pupescens and 2.29 m for Q. frainetto). Cut stumps had more sprouts than intact trees in both species. Forest practice, in order to maximize the sprouting of Q. pubescens and Q. frainetto trees after a forest fire, should cut the burned trees before the next growing season.     

章古台沙地油松天然更新的初步研究

通过2年的野外调查与比较分析,认为在章古台沙地油松天然更新是可能的.天然更新方式有2种:林隙更新;林缘更新.天然更新幼树在空间上呈带状、群团状分布,一般分布在距母树树冠投影以外的东北方向;更新幼树年龄在时间上一般是不连续的,主要集中在年降水量大的2004、2005、2008年;每100 m2更新幼树57～139株.油松...     

桩径对尾叶桉萌芽更新的影响

试验对5.5年生尾叶桉实生林砍伐后，进行不同桩径的萌芽率、萌芽力、萌芽条的高和径生长调查。结果表明，桩径对萌发率的影响不显著，但对萌芽力有极显著的影响，在桩径3-17cm的范围内，萌芽数量随桩径的增大而增加；萌芽条的高和径的生长也随桩径的增大而显著增大。     

Individual seedling mortality as a function of size, growth and competition in naturally regenerated beech seedlings

Young forest stands that are naturally regenerated are characterizedby a large number of seedlings and high mortality rates. Goodprediction of seedling mortality is therefore necessary to modelthe dynamics of young stands. We used data obtained in a naturalbeech (Fagus sylvatica L.) regeneration in north-eastern France,to fit logistic models to predict seedling mortality and toanalyse how seedling size, seedling growth and local competitionrelate to seedling mortality. Good relationships were obtainedwith recent seedling diameter growth, and with a combinationof local competition and initial seedling size. However, therelationships between the probability of mortality and recentgrowth appeared to be unstable and to vary substantially withlocal competition and among measurement years, precluding theuse of these relationships in regeneration models.