Dynamics of coast redwood sprout clump development in variable light environments

Post-thinning stump sprout response was assessed in a coast redwood [Sequoia sempervirens (D. Don) Endl.] stand from 5 to 9 years after five treatments were initiated. Nine years after treatment, leaf area index (LAI) ranged from 5.9 to 14.1 and the percent above canopy light (PACL) ranged from 0.8 to 3.9. Sprout responses included rapid self-thinning in all treatments with complete sprout mortality at low light regimes. Self-thinning of clumps increased with greater overstory LAI and poorer light regimes. Leaf area of individual sprout clumps also declined in low light regimes. The probability of sprout survival declined rapidly below 10 PACL and coincided with declining leaf area in sprout clumps. A conceptual model is presented that shows the sensitivity of redwood sprouts to light regime. Results have implications for management of multiaged stands and indicate these redwood coppice systems require relatively severe overstory treatments to provide sufficient light for sprout vigor and growth.     

Survival and development of Liaodong oak stump sprouts in the Huanglong Mountains of China six years after three partial harvests

The survival and development of Liaodong oak (Quercus liaotungensis Koidz) stump sprouts were studied for 6 years under leave-tree, as well as 60 and 75% crown closure uniform shelterwood in the Huanglong Mountains of China. Stump sprout survival rate, number of sprouts, and sprout height under the three partial harvests during the first year post-harvest did not differ. Six years post-harvest, 78.32% of the stumps had living sprouts. No variation in sprout survival was found among the treatments throughout the study period. The number of living sprouts per stump was 13.77 ± 0.50 in the first growing season after harvest, and dropped to 6.49 ± 0.22 after 6 years. This result indicated the occurrence of self-thinning within the sprout clumps. Moreover, the height of dominant sprout increased from 52.88 ± 1.62 cm (in the first year) to 132.82 ± 2.35 cm (in the sixth year) post-harvest. Sprout height in the leave-tree harvest type was higher than those in the two shelterwoods in the fifth and sixth years after logging. Annual height growth was approximately 8–96 cm during the first growing season, and then decreased at a rate of 1–44 cm per year in the following years. A negative relationship was found between sprout height and the number of sprouts per stump. This result indicated a trade-off between the number of sprouts and height growth. The number of sprouts per stump decreased by about 0.2 stem for each 1 m²/ha increased in residual basal area. Therefore, sprouting may play an important role in the restoration of the secondary forest after harvesting.     

辽东栎伐桩萌苗的发育规律

通过对黄土高原黄龙山林区辽东栎林固定样地连续6年的调查,研究间伐强度分别为13.4％(弱度间伐)和30.0％(强度间伐)的辽东栎伐桩萌苗的存活与生长规律.结果表明:1)间伐第1年,辽东栎伐桩均有萌生枝发生,每桩萌生枝数量和萌生枝高度在不同强度间无显著差异;2)萌生枝数量受伐桩高度和直径的影响,当伐桩高度较低时,萌生枝数量随伐桩高度的增加而增加,超过20cm后随之减少,呈三次抛物线关系;伐桩萌生枝数量随着伐桩直径增加呈现先增加后减少的趋势,直径30～50cm的伐桩萌生枝数量最多;3)伐桩萌生枝数量与最长萌生枝高度呈显著线性负相关,存在“萌生枝数量——高度权衡”现象;4)间伐第6年,伐桩的存活率为97.01％,强度间伐区萌生枝高度显著高于弱度间伐区;5)调查期间,辽东栎伐桩萌生枝数量逐年下降,伐桩产生的萌生枝数量越多,后期萌生枝死亡的数量也越多,表明伐桩能够自我稀疏,存在“萌代主”现象.在黄土高原地区,辽东栎伐桩萌芽潜力很大,应合理利用;间伐时伐桩高度应尽量低于20cm;伐桩直径在30～50cm更有利于其无性更新;间伐后伐桩应及时除蘖定株,促进萌生枝生长;间伐强度30.0％(强度间伐)更有利于辽东栎伐桩萌生枝的生长.     

Sprouting characteristics of <Emphasis Type="Italic">Quercus pubescens</Emphasis> Willd. and <Emphasis Type="Italic">Quercus frainetto</Emphasis> Ten. trees after a wildfire: To cut or not to cut the burned trees?

The aims of this study were to analyze the sprouting characteristics of Quercus pubescens and Q. frainetto trees after a wildfire and to recommend the appropriate silvicultural treatments in order to achieve dense sprout origin regeneration. This study was carried out in the area that was burned in a wildfire in northeastern Greece. Three years later, 140 Q. pubescens trees and 140 Q. frainetto trees were selected. Seventy trees for each species were individuals that were cut after the fire and were in the form of stumps, while 70 trees were intact top-killed trees, as a result of fire. In each stump and intact tree that was selected, various parameters were measured or counted: the diameter at the level of ground, the number of the developed sprouts (if any) and the height as well as the basal diameter of the tallest sprout, etc. Both species have similar sprouting behavior after a wildfire. Mother tree dimensions positively influenced the number of sprouts and the growth of the tallest sprout per individual. The mean height of the tallest sprout per stump, after three growing seasons from the forest fire, is high in both species (2.08 m for Q. pupescens and 2.29 m for Q. frainetto). Cut stumps had more sprouts than intact trees in both species. Forest practice, in order to maximize the sprouting of Q. pubescens and Q. frainetto trees after a forest fire, should cut the burned trees before the next growing season.     

Sprouting ability of two‐year‐old Betula pendula stumps exposed to different light intensities during five years

Altogether 1 080 two‐year‐old silver birch (Betula pendula Roth) were planted in groups of 36 plants at aspacing of 20 x 20 Cm in spring 1983 on a nursery field at lat. 60° 15 N and long. 16°00 E. In spring 1984 all plants were cut with 10 cm high stumps and light screens were arranged on the stump groups. Relative ¡rradiance (RI) 5, 10, 25, 50, 75 and 100% of full sunlight were used. After five growing seasons, all stumps exposed to 5 or 10% light intensity were dead, and 64–94% of stumps exposed to 25–100% RI were alive. 1.3–1.4 sprouts per living stump were living five years after cutting. The mean height of sprouts was highest on stumps exposed to 100% light. Also dry weights of leaves and sprouts, number of leaves per sprout, and leaf area were highest on stumps exposed to full sunlight. Stumps producting only one sprout five years after cutting have higher height and dry weight values than stumps with more sprouts. The biomass production per hectare and year for silver birch sprouts at a spacing of 0.2 m and with 90% survival was calculated as 5.4 tonnes. The biomass of leaves was 8.1 tonnes/ha/year five years after cutting.     

不同直径伐桩对赤桉萌芽更新影响的初步研究

对速生树种赤桉进行了不同直径伐桩（２．１～６ｃｍ）萌芽更新影响的研究，结果表明：伐桩的萌芽率及平均萌条数随伐桩直径的增大而增大，萌条直径、高度随伐桩直径变化不大．     

印度阿鲁纳恰尔邦温带阔叶混交林树桩周长和高度对受干扰的树形杜鹃再发芽的影响

Sprout contribution to natural regeneration of Rhododendron arboreum following the tree felling for fuel wood and other purposes was studied in two disturbed sites (Paipraw and Falockchar) of Rhododendron forest in Arunachal Pradesh, northeast India. Cut stumps were categorized into seven girth and height classes to evaluate the effect of stump girth and height on production of sprouts. Effect of stump girth on survival of sprouts was also investigated. Results show that highest number of cut stumps is recorded in medium girth classes (50–80, 80–110, and 110–140 cm) in both Paipraw and Falockchar stands. However, density of cut stumps is higher in Paipraw (1 210 stumps·ha⁻¹) than Falockchar (1 154 stumps·ha⁻¹). Nevertheless, a little higher percentage of sprouting is recorded in Falockchar (18%) than that of Paipraw (15%). Similarly, maximum number of cut stumps is observed in the medium height classes (50–80, 80–110 and 110–140 cm) in both the sites. Cut stumps with lower girths show greater ability of sprouting than that of larger girth classes. Significant correlations are observed between stump girth and sprout number and between stump height and number of sprouts. Survival of sprouts also depends on stump girth. Survival of sprout shows significant variations between different sampling times in both the sites. Though overall sprout regeneration of R. arboreum is very poor, sprouts arising from lower girth classes survive well than those of higher girth classes. The findings of this investigation signify that stump sprouting is not playing much role in the natural regeneration of R. arboreum though it has the ability to sprout. Sprout survival is not adequate to restore a stand where indiscriminate tree felling is continued and the species may be considered as a poor coppicer. Therefore, regeneration through seeds and seedlings should be preferred over regeneration through sprouting.     

Influence of thinning time and density on sprout development, biomass production and energy stocks of sawtooth oak stumps

The high potential values of sawtooth oak (Quercus acutissima) in fuelwood or bioenergy are recognized. Sprouting ability, sprout growth, biomass production and energy stocks in coppiced plantations of sawtooth oak were evaluated at the Hongya Mountain Forest Farm in Anhui Province, China. Experimental treatments applied in a split-plot design included three sprout thinning times and four sprout numbers reserved on each stump (1 sprout, 2 sprouts, 4 sprouts stump⁻¹ and check). Sprout growth and biomass production per stump were significantly affected by the treatments and a significant positive relationship between stump basal diameter and sprout numbers produced was observed. After the third growing season, the highest total sprout biomass per stump was achieved in the treatment with thinning excess sprouts at the end of first growing season (December, 2007) and reserving 4 dominant sprouts per stump (T12S4, reaching 8.67 kg stump⁻¹), while the lowest was found in the treatment with thinning the sprouts in August of the first growing season and reserving 1 dominant sprout per stump (T8S1, only 3.40 kg stump⁻¹). Different treatments also influenced gross calorific values (GCV) of the components sampled from 3-year-old sprouts and the mean GCV of stem wood on an oven-dry weight basis was within the range of 18.45 ± 0.15 and 18.83 ± 0.12 kJ g⁻¹. Similar to the sprout biomass production, the greatest total and stem energy stocks per stump were observed in T12S4 treatment, achieving 161.6 and 110.5 MJ stump⁻¹, respectively. Based on the results from this study, thinning excess sprouts to reserve 4 sprouts per stump as early as age 1 could be proposed for the management of sawtooth oak coppice with cutting cycle of 3-5 years and stand density of 5000-6000 stump ha⁻¹.     

不同伐桩直径及高度对马占相思萌芽更新影响的研究*

对速生多用途树民占相思进行了不同伐桩高度（25cm,50cm,75cm和100cm)及不直径（4．1－8．0cm和8．1－12．0cm)萌芽更新影响的研究，结果表明，伐桩的萌芽率及平均萌条数随伐桩高度的增加而增加，随直径的增大而减少，萌条直径，长度及直萌条（可望生长成材之萌条）总断面积随伐桩的高低和大小而增减，伐桩最低的萌芽率比以往的结果高4倍以上，以短轮伐期经营的马占相思人工林，采用50－75cm伐桩高度萌芽更新效果好。     

Cherrybark oak stump sprout survival and development five years following plantation thinning in the lower Mississippi alluvial valley,USA

Cherrybark oak (Quercus pagoda Raf.) stump sprouts were studied for 5 years in a 30-year-old plantation thinned to 70–75% stocking (light thinning) and 45–50% stocking (heavy thinning). Sprouting success, survival, number of sprouts per stump, and sprout height differed little between thinning treatments throughout the 5-year study period. Pre-harvest tree d.b.h. also had no influence on sprout survival and development. A 2-year drought reduced survival and may have influenced sprout development. Sprout clump survival dropped from 90% 1 year following thinning to 46% 3 years after thinning. Although sprout height averaged 337 cm 5 years after thinning, annual sprout growth decreased from 166 cm the first year after thinning to 33 cm in each of the last 2 growing seasons. Results indicated that bottomland hardwood regeneration evaluation models may underestimate the potential of oak stump sprouts to contribute to pre-harvest regeneration assessments. Further study in the role of stump sprouts to regenerate bottomland oak species is needed. The U.S. Government's right to retain a non-exclusive, royalty-free license in and to any copyright is acknowledged     

Effects of thermal wounding, shading and exogenous auxin on some sprouting responses of coast redwood seedlings

The effects of wounding, shading, and indole-3-acetic acid (IAA) on sprouting responses were examined in coast redwood (Sequoia sempervirens D. Don (Endl)) seedlings. In August, two-year-old seedlings were given one of three thermal wounding treatments made with a blowtorch at the stem base. After 8 months, the heat-wounded seedlings were decapitated above the burl and IAA (0.00% to 1.00% by weight in lanolin) was applied to the cut surface every week for 8 weeks. Throughout the 8-week hormone treatment, the decapitated seedlings were subjected to one of three shading treatments (no shade, 70% shading and complete darkness). At the beginning of June, sprouts were harvested from each stump and measurements made of the length of the tallest sprout, number of sprouts, dry weight of all sprouts, and mean dry weight per sprout. All sprouting parameters showed negative responses to IAA. Shading decreased sprout number and sprout dry weight. Wounding increased maximum sprout length and dry weight per sprout but decreased the number of sprouts. Only the 2-way interaction between IAA and wounding was not significant for any of the responses.     

东北东部林区次生林主要阔叶树种的萌芽更新规律

对固定样地调查,采用伐桩的径、高度、萌芽的数量、萌芽的生长情况等指标,分析东北东部林区主要阔叶树种的伐桩萌芽更新规律。结果表明:在一定的伐桩径范围内,伐桩萌生枝数随着伐桩径的增大而增多;增长到一定程度后随着伐桩径的增大而减少;得出不同树种萌芽能力较强的年龄范围的伐桩径级。对于不同阔叶树种,伐桩高与当年生萌生枝数量的关系存在着不同的变化规律,研究地点具有产生萌芽能力的阔叶树种可划为3种类型:对数型、直线型、指数型,阔叶树的萌生枝高生长与伐桩径和伐桩高无明显关系。第2年伐桩萌生枝的数量与伐桩径的关系在总体趋势上与第1年基本相同,同时次年仍有新的萌生枝产生,但数量很少。与第1年不同之处在于萌生枝开始死亡,一方面,伐桩产生的萌生枝越多,死亡的数量也增大;另一方面,第2年的伐桩存活率在不同种类树种间差别较大:水曲柳高达96·2%,槭树为23·3%,榆树类为21·8%。所探讨的树种在研究区域内都有“萌代主”的能力,并且不同树种以及同一树种不同年龄,“萌代主”的能力和单个伐桩保存的萌生大树的数量不同。     

林龄、采伐方式对大叶相思萌芽更新的影响*

在海南省琼海县进行了不同林龄、不同采伐方式对大叶相思萌芽更新影响的研究，同时比较了保留不同萌条数量的萌芽林生长表现。结果表明：（１）林龄对伐桩萌芽率没有显著影响，但对伐桩存活率及萌条的径、高生长影响显著或极显著。林龄越大，萌芽更新效果越差；（２）皆伐和隔行采伐对大叶相思伐桩的萌芽率、萌条数量和存活率均无显著影响，但对萌条的径、高生长影响极显著。试验证明对大叶相思林分施以隔行采伐通过萌芽更新建立复层     

桩径对尾叶桉萌芽更新的影响

试验对5.5年生尾叶桉实生林砍伐后，进行不同桩径的萌芽率、萌芽力、萌芽条的高和径生长调查。结果表明，桩径对萌发率的影响不显著，但对萌芽力有极显著的影响，在桩径3-17cm的范围内，萌芽数量随桩径的增大而增加；萌芽条的高和径的生长也随桩径的增大而显著增大。     

Natural regeneration in oak wilt centers

Openings, 0.2 ha and less, and 3–12 years old, caused by the oak wilt fungus Ceratocystis fagacearum had abundant oak reproduction, averaging 4700 stems per ha. Species of other woody reproduction made up over 27 000 stems per ha. Seedling sprouts comprised 54%, seedlings 30%, and stumps sprouts the remainder of the reproduction. No infections were found except a few on stump sprouts of diseased oak stumps. Oakhickory reproduction was abundant on all site qualities, but especially so on the poorer sites with over 8000 stems per ha. Sugar maple reproduction was much more abundant on high quality sites, also with over 8000 stems per ha. The reproduction in the oak wilt openings was judged to be similar to that of selection cuttings in disease-free but otherwise similar stands, and should respond in like manner.     

林龄、采伐方式对大叶相思萌芽更新的影响

在海南省琼海县进行了不同林龄、不同采伐方式对大叶相思萌芽更新影响的研究，同时比较了保留不同萌条数量的萌芽林生长表现，结果表明：（１）林龄对伐桩萌芽率没有显著影响，但对伐桩存活率及萌条的径、高生长影响显著或极显著。林龄越大，萌芽更新效果越差；（２）皆伐和隔行采伐对大叶相思伐桩的萌芽率、萌条数量和存活率均无显著影响，但对萌条的径、高生长影响极显著。试验证明对大叶相思林分施以隔行采伐通过萌芽更新建立复层林分是可行的；（３）保留不同数量萌条对萌芽林早期（１．５年生）的径、高生长有极显著影响，但对后期（４．５年生）的生长影响不显著。每棵植株伐桩保留４根萌条的萌芽林生物产量最高，更新效果最好。     

Predicting average growth and size distributions of Douglas-fir saplings competing with sprout clumps of tanoak or Pacific madrone

The average growth and size distributions of Douglas-fir (Pseudotsuga menziesii) saplings in three plantations were studied for 7 years (1983–1989) after thinning of associated sprout clumps of tanoak (Lithocarpus densiflorus) or Pacific madrone (Arbutus menziesii); in some cases understory vegetation (shrubs and herbs) was experimentally suppressed. Biologically based nonlinear equations explained 66, 90, and 53% of the variation in the average annual increment of Douglas-fir height, diameter-squared, and cover, respectively. Equations for annual increment of cover of hardwood and understory vegetation explained only 10 to 12% of the variation, because these parameters exhibited a high degree of variability. Model simulations demonstrated that, for the same initial levels of cover, tanoak had faster rates of cover growth than madrone and also caused greater limitations in growth of Douglas-fir. Suppression of understory vegetation increased growth of Douglas-fir only when hardwood cover was absent. Weibull functions adequately described size distributions for Douglas-fir in 92% of the individual-tree data sets. Regression functions of hardwood cover and average Douglas-fir size explained 51, 93, and 24% of the variation in the Weibull A, B, and C parameters, respectively. Model simulations with predicted Weibull parameters demonstrated that hardwood competition caused a positive skewing in size distributions for height and stem diameter of Douglas-fir.