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1.
A shortage of available livestock for utilizing grassland biomass in Central Europe is challenging for the management of both semi‐natural grasslands and previously intensified (limed, fertilized and reseeded) upland grasslands. An alternative method of grassland management is mulching, in which aboveground biomass is cut, crushed and subsequently spread on the surface. This paper reports on an experiment to compare three different mulching frequencies (one, two and three times per year) with an unmanaged treatment and traditional management of two cuts per year (control) on a previously improved upland meadow. Plant species composition was monitored over 13 years. Traditional management of two cuts with biomass removal was the most suitable method for maintaining plant species richness and diversity, and both were reduced significantly in the once‐mulched and especially in the unmanaged treatment. Tall dicotyledonous weeds such as Urtica dioica, Cirsium arvense and Aegopodium podagraria were promoted by the unmanaged treatment and by mulching once a year. Higher frequency of defoliation had positive effects on the spread of short forbs such as Taraxacum spp., Plantago lanceolata and Trifolium repens. After eight years, there were changes in sward structure in the unmanaged and mulched‐once‐a‐year treatments, with increase in the tall/short species ratio. In conclusion, repeated mulching cannot substitute fully for traditional two‐cut management in improved upland meadows without decreasing plant species richness and diversity, and changing the sward structure. Although mulching once a year may prevent invasion by shrubs and trees, it also supports the spread of weedy species similar to no management.  相似文献   

2.
A 3‐year experiment was conducted in Central Europe to examine the effects of three managements, viz. continuous cattle grazing from April to September, mowing once in July and abandonment of grazing, and two levels of fertilizer application, no fertilizer and 400 kg ha?1 of a NPK fertilizer, on changes in plant species composition and summer biomass of dry matter (DM) in a calcareous mountain grassland containing many plant functional types. Different managements led to changes in species composition due to species‐specific responses. Low creeping and rosette species were associated with grazing, while grasses and tall forbs correlated with mowing and abandonment of grazing, probably because of their ability to outcompete rosette species. There was a negative relationship between the number of species and above‐ground biomass and a positive relationship between number of species and below‐ground biomass, suggesting that these species‐rich communities allocate more to below‐ground organs when not grazed. The application of fertilizer had no effect on species composition but it indirectly increased competitive asymmetry for light and increased the number of plant extinctions. It is concluded that continuation of cattle grazing is an acceptable form of grassland management at the study site and that species and functional group compositions can rapidly change with changing environmental conditions such as abandonment of grazing or application of fertilizer.  相似文献   

3.
Changes in plant density were evaluated monthly in the first three vegetation seasons after imposing different grazing management treatments on abandoned semi‐natural grassland in the Czech Republic. There was no agricultural management in the 5 years before the start of the experiment in 1998. A completely randomized block experiment was established with the following five treatments: unmanaged control, intensive continuous grazing, extensive continuous grazing and a harvest in June followed by either intensive or extensive continuous grazing for the rest of the growing season. The sward was maintained at a target height of 5 and 10 cm under the intensive and extensive grazing managements respectively. An almost immediate increase in the densities of all sward components, especially grass tillers, occurred after the introduction of grazing on the previously abandoned grassland in comparison with the unmanaged control treatment. Trifolium repens was able to colonize and increase the number of its stolon growing‐points in all managed treatments, particularly in intensively grazed patches during the second and third experimental seasons. Delay to defoliation in both treatments containing a harvest in June resulted in an increase in the number of forb plants, particularly in the number of Taraxacum spp. plants, most probably due to an enabling of its seed production. It is evident that increases in plant density as a function of intensive defoliation are not restricted to the frequently documented effect on grass tillers but also can occur in many legume and forb species in species‐rich grasslands.  相似文献   

4.
Dandelion (Taraxacum officinale agg.) is a common forb species in grasslands in Europe. Although sometimes regarded as a valuable forage herb, it may become a weed, especially in arable land. There is limited information on the response of Taraxacum to long‐term grassland management practices. Therefore, we analysed cover and dry‐matter standing biomass of Taraxacum in a long‐term (1998–2012) grazing experiment on an Agrostis capillaris grassland. The following treatments were laid out on formerly abandoned grassland: (i) intensive grazing (IG); (ii) extensive grazing (EG); (iii) first cut followed by intensive grazing (ICG); (iv) first cut followed by extensive grazing (ECG); and (v) unmanaged grassland (U). During the first 10 years, all defoliation treatments (i–iv) supported the presence of Taraxacum, and the lowest proportion was recorded in the unmanaged treatment (U). During the final 7 years of the study, combined cutting and grazing promoted Taraxacum cover more than that of grazing only (ICG > IG > ECG > EG). Cover of Taraxacum was negatively affected by increasing sward height where Taraxacum plants had lower fitness. Due to the relatively strong relationship between percentage cover of Taraxacum and its dry‐matter biomass, percentage cover could be used as a simple method for the assessment of biomass of Taraxacum in a sward. Results are discussed in the context of adapting the management of A. capillaris grassland as a simple method for control of Taraxacum abundance, particularly in situations of extensification or abandonment.  相似文献   

5.
The plant species number and composition, iind yield of herbage biomass of an agriculturally improved hay meadow were assessed after 4 years under various combinations of grazing, fertilizer applicution. cutting date and seed addition treatments in a replicated split-plot design. Grazing treatments consisted of either autumn grazing with cattle and sheep, spring grazing with sheep or both regimes. Fertilizer application treatments consisted of either 25 kg ha?1 N plus 12–5 kg ha?1 P and K or no fertilizer. Cutting date treatments consisted of cuts on either 14 June, 2i July or 1 September. Seed addition treatments consisted of either no addition or sowing with a range of meadow species in the autumn. Data analysis was by correspondence analysis and analysis of variance. Species number decreased with fenilizer use and when the cutting date was 1 September. A range of species was affected by the main treatments and there were some first-order interactions, mainly between cutting date and fertilizer application. Rhinanthus minor was particularly favoured by the seed addition treatment. Species attnbutes in the regenerative and established phase were related to treatments and their effect on species composition. The National Vegetation Classification communities were associated with particular treatment regimes. The 21 July cutting date favoured ‘improved’ over ‘unimproved-traditional’ swards, with spring grazing favouring ‘unimproved-traditional’ swards. Lowest yields of herbage biomass were associated with autumn and spring grazing, the 14 June cutting date and no fenilizer treatments. The fenil-izer, 1 September cutting date and auiumn grazing treatments gave the highest yields. The implications of these results are discussed in terms of the conservation management required to return agriculturally improved mesotrophic grassland to a species composition similarto that of traditionally managed grassland.  相似文献   

6.
This study quantifies the impact of four different grazing regimes (heavy, moderate, light and ungrazed) on the vegetation dynamics of rangeland ecosystems along the southern boundary area of the Mu Us Desert, China. As the grazing intensities decreased, the soil quality, canopy cover, height, density, above‐ and below‐ground biomass, litter, root/shoot ratio and native plant (Aneurolepidium dasystachys) and grass abundances significantly increased; the above‐ground biomass of grasses increased, but the above‐ground biomass of forbs decreased. Ungrazed grassland has significantly improved from grasslands experiencing three other levels of grazing pressure, especially in the grassland biomass. Species richness increased as the grazing intensity decreased in the grazing grasslands, but peak species richness appeared under moderate and light grazing against lower productivity. Grazing exclusion causes desirable transitions in plant communities of desert steppe rangelands. Therefore, appropriate and efficient grazing exclusion is an available way to counteract local grassland degradation and promote rangeland sustainability.  相似文献   

7.
We compared the role of grasses and forbs in managed grassland under nutrient‐poor conditions via the experimental removal of one of these two species groups and an evaluation of the removal effects over a 5‐year chronosequence (beginning 4 years after removal start), focusing on above‐ground biomass and community composition. There was evidence for the complementarity of grasses and forbs, as shown by better relative performance of both groups in the mixture, measured as the proportional deviation of their above‐ground biomass from that predicted using single‐group plots. Relative performance of forbs was better than that of grasses and did not change significantly throughout the experiment. Relative performance of grasses decreased, reaching a minimum value of 0·03, but then increased to 0·71, in parallel with increasing legume proportion in the above‐ground biomass (from 0·9 to 5·1%). The botanical composition of the grass group did not respond to forb removal. Among the forbs, subordinate species sensitive to light competition became more abundant after removal of grasses; the forb species richness increased on average by 1·6 species per plot. Our results show that the composition and diversity of the forb group is affected by the presence of grasses, and the presence of forbs in a grassland community assures higher temporal stability in above‐ground production. Under increasing depletion of soil nutrients, the grasses derive substantial benefits from the presence of legume species and possibly also from the other forb species. The study supports the need for realistic, long‐term experiments to better understand plant community dynamics.  相似文献   

8.
Little is known about the immediate effect of high nitrogen (N), phosphorus (P) and potassium (K) application rates on sown grasslands cut twice per year. We asked how quickly plant species composition, biomass yield, biomass chemical properties and nutrient balance respond to N, P and K application. An experiment using unfertilized control, P, N, NP and NPK treatments was established on seven‐year‐old cut grassland in the Czech Republic in 2007 and monitored over four years. Annual application rates were 300 kg N ha?1, 80 kg P ha?1 and 200 kg K ha?1. The immediate response of plant species composition to N application was recorded and was found to be different to the response over the four years of the study period. Highly productive grasses (Dactylis glomerata, Festuca arundinacea and Phleum pratense) were promoted by N application in 2008 and then retreated together with legumes (Medicago sativa, Trifolium pratense and Trifolium repens) in all N treatments where the expansion of perennial forbs (Urtica dioica and Rumex obtusifolius) and annual weeds (Galinsoga quadriradiata, Impatiens parviflora, Lamium purpureum and Stellaria media) was recorded. At the end of the experiment, Festuca rubra was the dominant grass in the control and P treatment, and species richness was lowest in all treatments with N application. Mean annual dry‐matter yield over all years was 3.5, 3.9, 5.8, 5.6 and 6.8 t ha?1 in the control, P, N, NP and NPK treatments, respectively. Concentrations of N in the biomass ranged from 20.0 to 28.7 g kg?1 in the P and N treatments; concentrations of P ranged from 3.2 to 3.7 g kg?1 in the N and P treatments; and concentrations of K ranged from 24.1 to 34.0 g kg?1 in the NP and NPK treatments. The N:P, N:K and K:P ratios did not correctly indicate the nutrient limitation of biomass production, which was primarily N‐limited, and K‐limitation was only recorded for high production levels in treatments with N applications. On the basis of the nutrient‐balance approach, the balanced annual application rates were estimated as 140 kg N ha?1, 30 kg P ha?1 and 100 kg K ha?1. We concluded that high N, P and K application rates can very quickly and dramatically change species composition, biomass production and its chemical properties in sown cut grasslands. High N application rates can be detrimental for tall forage grasses and can support the spread of weedy species.  相似文献   

9.
Livestock grazing can be a means to maintain biodiversity in grasslands, but the outcome for vegetation structure and species composition depends on livestock type and grazing regime. This study aims at disentangling the effects of plant functional‐group abundance and livestock type on the above‐ and below‐ground biomass and N allocation in temperate pastures. We investigated the effects of cattle, sheep and mixed stocking on above‐ground biomass (AGB) and belowground biomass (BGB) and plant N pools in a replicated grazing experiment in two pasture community types with different plant functional‐group abundance (diverse vs. grass‐dominated swards). In the six treatments, AGB was reduced up to 80% compared with an ungrazed control. Cattle reduced AGB to a larger extent than sheep in diverse pastures (80 vs 44% reduction) while sheep grazing tended to do so in grass‐dominated pastures (57 vs 46% reduction); mixed stocking led to intermediate values. Grazing reduced AGB more than the N pool in AGB, thus lowering the biomass C/N ratio relative to the ungrazed control. Neither BGB nor the N pool in BGB differed between the grazing treatments and the control plots. We conclude that livestock type and functional‐group abundance are interacting factors that influence plant biomass and N pools in swards of managed temperate pastures. The contrasting biomass removal rates of cattle and sheep could be used to increase the structural heterogeneity and total plant species pool of pastures by keeping different livestock species in neighbouring patches.  相似文献   

10.
To determine the impacts of climate change and defoliation on the community structure and plant diversity of a semi‐natural temperate grassland, monoliths of a permanent grassland were exposed to ambient or elevated atmospheric CO2 concentrations (ambient + 235 ppmv) and temperature (ambient + 3°C) from October 1998 to December 2000. The monoliths were subjected to two different cutting frequencies, either two or six cuts per year. The grassland community structure changed during the course of the experiment and was more responsive to changes in management than to changes in climate. Increased cutting frequency stimulated plant diversity by enhancing the number of forb species, but plant diversity was not significantly affected by climate change. The contribution of individual plant species to the vegetation cover revealed species‐specific responses to climate change and cutting frequency, but for most species significant interactions between climate change and cutting frequency were present. There were no clear‐cut effects of treatments on the total annual yield and the proportion of forbs present, as significant interactions between climate change and cutting frequency occurred. It is concluded that differential grassland management will modify plant species‐specific responses to climate change and resulting changes in the botanical composition of mixed‐species, temperate grasslands.  相似文献   

11.
Studying the seed production of herbaceous species can help to conserve grassland habitats and re‐create new high‐value grassland surfaces. Studies on grassland seed production have focused mainly on individual species and traits, without characterizing their relative importance at the plant community level. The aim of this study was to investigate the entire seed production process of the main species in a temperate grassland. Fertile shoots (FS) of twenty‐nine grasses and forbs were collected over 4 years and analysed for sixteen traits that determine inflorescence size, seed production and seed quality. The per cent viability played a predominant role in determining the total production of viable seeds. Forbs showed a range of reproductive strategies, including variable distribution of flowers among growth periods, number of inflorescences per FS and relationship between seed size and FS density in the grassland. The flower production for grasses was concentrated in the first growth period, but this limitation was mitigated by a higher seed dormancy. The number of viable seeds per FS and seed size were important components of the reproductive strategy of forbs, with heavy‐seeded species being characterised by high individual densities in the community, but producing few seeds per FS. Light‐seeded species showed an opposite pattern. The results suggest that when using seeds from semi‐natural grasslands for ecological restoration, special attention should be paid to the seed amount, germinability and viability of forbs, as they seem to depend more on seed reproduction and have a lower ovule to seed transformation efficiency.  相似文献   

12.
Tall fescue (Festuca arundinacea) is a Eurasian forage grass extensively planted in the United States. However, an endophytic fungus in tall fescue, Epichloë coenophiala, causes health problems in cattle. We predicted that cattle prefer to graze alternative forages when available. We also predicted that cattle use tall fescue more intensively in recently burned areas, as fire can increase forage quality. We tested these predictions in four diverse‐forage pastures in Iowa, comparing use by cattle of tall fescue and four alternative forages (non‐fescue cool‐season grasses, native warm‐season grasses, non‐leguminous forbs and legumes) to their availabilities at the pasture scale. We also examined how tall fescue influences the distribution of grazing at a fine scale (0.1‐m2 quadrats). Tall fescue was the most abundant forage (46% of plants), but composed only 26% of grazed vegetation. In contrast, legumes composed 12% of available forage but 25% of grazed vegetation. Other forages were used in proportion to availability. At a fine scale, total grazing frequency (proportion of plants grazed) was lower in quadrats containing abundant tall fescue, and higher in quadrats with abundant warm‐season grasses. Grazing frequency of tall fescue and other cool‐season grasses was greatest in recently burned quadrats, but total grazing frequency did not increase after burning. Our results show that although cattle graze tall fescue, particularly following burns, they limit their use of this grass. Given that tall fescue is underused, creates health risks for cattle, and degrades wildlife habitat quality, it may be advisable to reduce tall fescue in pastures.  相似文献   

13.
Social Network Analysis (SNA) is used for the first time to investigate the relations between plant functional groups and social structure of grazing sheep. The research was conducted on a grassland in Crete's island Lefka Ori, Greece during 2016. A flock of 20 sheep of Sfakion race was analysed as a network and the sheep as nodes. The focal sampling technique was applied during four experimental periods of four consecutive days × eight hours/day. The plant species selected by sheep during grazing were categorized into four functional groups (grasses, broad‐leaved forbs, shrubs and trees), and the proximity and conflict relations that were developed among the flock members were recorded. The tools of SNA were defined and interpreted in the sheep flock, and the Visone software is used to calculate the network variables of proximity and conflicts relations of nodes. Correlations between the plant functional groups and network variables were examined by Spearman's bivariate correlation test. Sheep collectivity was enhanced while grazing grasses and simultaneously, a tendency for hierarchization within the flock was identified. Similarly, the proximity of sheep increased during forbs’ grazing while their individualism was strengthened. When grazing shrubs, the proximity of sheep was only in part maintained while the grazing of trees seems as the most deconstructive forage option of sheep proximity relations. In general, the grazing of all plant functional groups decreases conflicts among sheep except that of trees, which was insignificant for developing such relations.  相似文献   

14.
Reduction in grazing intensity and the use of traditional instead of commercial breeds have frequently been recommended to meet biodiversity and production goals in sustainable grazing systems. The impact of such practices within a range of contrasting grassland vegetation types was studied. This paper describes the effects on foraging behaviour in a study conducted over three years on mesotrophic or semi-natural grasslands in UK (steers), Germany (steers), France (heifers) and Italy (sheep). Three treatments were performed: (i) a moderate grazing intensity using a commercial breed, (ii) a more lenient grazing intensity with a commercial breed and (iii) a more lenient intensity with a traditional breed. Livestock at all sites preferentially selected bites containing legumes and forbs, and also short rather than tall vegetative patches. Grazing intensity affected not only diet consumed, largely reflecting the different availabilities of dietary components, but also some differences in diet selection. Livestock grazing the more productive mesotrophic grasslands more frequently exploited short patches of higher nutritive value, which is expected to reinforce the spatial heterogeneity of the pastures. Studies in the UK and Germany also revealed that steers showed a more pronounced selection for short patches at the lenient grazing intensity. More homogeneous grazing by livestock on the semi-natural grasslands with fine-scale heterogeneity is likely to decrease their spatial heterogeneity. There were few differences in the choices made by commercial and traditional breed livestock. North Devon steers in the UK expressed a greater selection for tall grass-forb bites than Charolais × Holstein crossbreds, whereas traditional breeds appeared slightly less selective than commercial breeds at the other three sites.  相似文献   

15.
In Maqu County, Gansu Province, China, there are two types of grazing management. Under multi‐household (MH) management, grassland is jointly managed by two or more households without fences between pastures. Under single‐household (SH) management, fenced‐off parcels of grassland are used. SH management was imposed in the belief that it would alleviate grassland degradation. Comparable land parcels with similar stocking rates subject to MH and SH management were identified and surveyed to determine the species present and species density (m?2). MH land had greater number values than SH but the differences were not significant. A further analysis indicated that the number of households using MH land influenced species richness. There was also evidence to suggest that the quality of the forage available deteriorated under SH management. It was concluded that MH grazing resulted in greater species richness than SH and is perhaps an important contributor to biodiversity conservation and grassland management for the region.  相似文献   

16.
Biochar and hydrochar application to soil holds promise for climate change mitigation. This study provides first insights into the nutrient concentration and removal of grassland vegetation after addition of various carbon compounds together with pig slurry. Four treatments: control (no carbon application), feedstock, hydrochar and biochar from Miscanthus x giganteus were applied at a permanent grassland site near Giessen, Germany. Changes in plant functional groups, biomass production and nutrition status were monitored over 2 years. Total biomass production was not affected by the carbon amendments. However, biochar favoured growth of forbs over grasses, while legume growth was increased by all carbon amendments. The initial nutrient concentrations of the carbon compounds were enriched according to their degree of carbonization, potentially providing nutrients to plants. The plant biomass from hydro‐ and biochar amended plots, added up over 2 years, exhibited higher potassium concentrations compared to biomass from feedstock and control plots. All carbon amendments led to lower sodium concentrations in total biomass, compared to the control. Uncarbonized feedstock led to increased manganese concentrations in total biomass, while the concentrations of all other heavy metals were not influenced by any carbon amendment, compared to the control. From a plant and animal nutritional point of view, none of the carbon amendments reduced grassland yield or fodder quality. The study suggests that hydrochar and, even more so, biochar may provide a source of potassium to plants.  相似文献   

17.
Oat and ryegrass intercropping in pastures is widely used in regions with subtropical climates. The aim of this study was to evaluate the tiller size/density compensation mechanisms in monoculture and intercropping swards of black oats (Avena strigosa Schreb cv. IAPAR 61) and annual ryegrass (Lolium multiflorum Lam. cv common) under intermittent grazing. Treatments (black oat, annual ryegrass and their mixture) were assigned according to a complete randomized block design with four replicates. Ryegrass, oat and intercropped pastures were grazed when the swards reached a height of 17, 25 and 23 cm, respectively, and with a level of defoliation of 40%. The aerial biomass was determined with a rising plate meter, and the tiller population density (TPD) was estimated by counting tillers in three 10 cm diameter PVC rings per paddock. The mass per tiller was estimated based on the aerial biomass and the TPD of each paddock. Total herbage production did not differ among treatments, with values around 7400 kg DM ha?1. TPD decreased and mass per tiller increased linearly in the monoculture treatments. Tiller size/density compensation was observed in the three plant communities (treatments) according to the self‐thinning rule. In addition, no relationships were found when each species was analysed individually in the intercrop treatment. The results suggest that species in grass mixed swards adjust their population to keep a relatively constant leaf area index (LAI) over the grazing seasons, and that would help pastures to stabilize herbage production.  相似文献   

18.
Reduction of grazing intensity and the use of traditional instead of commercial breeds has frequently been recommended to meet biodiversity and production goals in sustainable grazing systems in Europe. To test the impact of such practices across a range of contrasting grassland types, integrated measurements of foraging behaviour, agronomic production and botanical, structural and invertebrate biodiversity were made over three years on four sites in the UK, Germany, France and Italy. The sites in the UK and Germany were mesotrophic grassland with high productivity and low to moderate initial levels of plant diversity, and were grazed by cattle. The French site was a semi‐natural, species‐rich grassland grazed by cattle. The Italian site contained a wider range in plant diversity, from species‐rich to mesotrophic grassland, and was grazed by sheep. The treatments were: MC, moderate grazing intensity with a commercial breed – this was designed to utilize herbage growth for optimum livestock production; LC, lenient grazing intensity with a commercial breed – this was designed to increase biodiversity by not fully utilizing herbage growth; and LT, lenient grazing intensity with a traditional breed – this was also designed to increase biodiversity. Neither fertilizers nor pesticides were applied. The nutritive value of the herbage and the performance of the livestock were measured. Mean stocking rates were proportionately 0·30–0·40 lower and mean sward heights and herbage mass on offer were 0·30–0·50 higher on the LC and LT treatments compared with the MC treatment. The proportion of live and dead material, and leaves and stems in the herbage, its chemical composition and nutritive value were little affected by the treatments. Individual livestock performance, measured as liveweight gain, showed no consistent response to treatment. In Germany, performance on the MC treatment was slightly lower than on the LC and LT treatments but no such difference was found on the sites in the other countries. Livestock breed did not have a strong effect on livestock performance. In the UK and France the traditional breeds had a lower performance but this was not the case in Germany or Italy. Livestock performance per ha of the LC and LT treatments was up to 0·40 lower than of the MC treatment. It is concluded that biodiversity‐targeted extensive grazing systems have potential to be integrated into intensive livestock production systems because the individual livestock performance reaches a similar level compared to a moderate grazing intensity. Traditional breeds did not have a production advantage over commercial breeds on extensively managed pastures.  相似文献   

19.
20.
An experiment was undertaken to examine the effect of supplement type on herbage intake, total dry matter (DM) intake, animal performance and nitrogen utilization with grazing dairy cows. Twenty‐four spring‐calving dairy cows were allocated to one of six treatments in a partially balanced changeover design with five periods of four weeks. The six treatments were no supplement (NONE), or supplementation with either grass silage (GS), whole‐crop wheat silage (WS), maize silage (MS), rapidly degradable concentrate (RC) or slowly degradable concentrate (SC). Cows were rotationally grazed with a mean herbage allowance of 20·5 kg DM per cow per day, measured above 4 cm. Forage supplements were offered for approximately 2 h immediately after each morning milking, with cows on NONE, RC and SC treatments returning to the grazing paddock immediately after milking. Cows on treatment MS had a significantly higher supplement DM intake than the other treatments but a significantly lower grass DM intake than the other treatments, resulting in no significant difference in total DM intake when compared with cows on treatments WS, RC and SC. Concentrate type had no significant effect on herbage intake, milk yield, milk composition or yield of milk components. The yield of milk fat and milk protein was significantly higher on treatments MS, RC and SC compared with treatments NONE, GS and WS. The results indicate that despite a relatively high substitution rate, maize silage can be a useful supplement for the grazing dairy cow.  相似文献   

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