Effects of long‐term grazing management on dandelion (Taraxacum officinale) in Agrostis capillaris grassland

Dandelion (Taraxacum officinale agg.) is a common forb species in grasslands in Europe. Although sometimes regarded as a valuable forage herb, it may become a weed, especially in arable land. There is limited information on the response of Taraxacum to long‐term grassland management practices. Therefore, we analysed cover and dry‐matter standing biomass of Taraxacum in a long‐term (1998–2012) grazing experiment on an Agrostis capillaris grassland. The following treatments were laid out on formerly abandoned grassland: (i) intensive grazing (IG); (ii) extensive grazing (EG); (iii) first cut followed by intensive grazing (ICG); (iv) first cut followed by extensive grazing (ECG); and (v) unmanaged grassland (U). During the first 10 years, all defoliation treatments (i–iv) supported the presence of Taraxacum, and the lowest proportion was recorded in the unmanaged treatment (U). During the final 7 years of the study, combined cutting and grazing promoted Taraxacum cover more than that of grazing only (ICG > IG > ECG > EG). Cover of Taraxacum was negatively affected by increasing sward height where Taraxacum plants had lower fitness. Due to the relatively strong relationship between percentage cover of Taraxacum and its dry‐matter biomass, percentage cover could be used as a simple method for the assessment of biomass of Taraxacum in a sward. Results are discussed in the context of adapting the management of A. capillaris grassland as a simple method for control of Taraxacum abundance, particularly in situations of extensification or abandonment.     

Consequences for biodiversity of reducing inputs to upland temperate pastures: effects on beetles (Coleoptera) of cessation of nitrogen fertilizer application and reductions in stocking rates of sheep

Current policies for upland pasture management in the UK encourage the integration of environmental objectives with livestock production through extensification of grazing systems. This study tested the hypothesis that a greater sward height in the summer would increase the diversity and abundance of grassland beetles (Coleoptera) as has been demonstrated for insects of indigenous grasslands. The hypothesis was tested with an experiment on an upland sheep pasture in mid‐Wales. Experimental treatments received different nitrogen fertilizer inputs (0 or 50 kg ha^?1), sheep stocking densities (12 or 9 ewes ha^?1) and average sward heights in summer were constrained to 3·5 or 5·5 cm by conserving surplus grass for silage in subplots. Five treatments, replicated in three randomized blocks, combined the two stocking densities and two sward heights without nitrogen fertilizer inputs, with the fifth combining the higher stocking density, shortest sward height and the nitrogen fertilizer input. Beetles were sampled with twelve pitfall traps in each of the fifteen plots from June to September in 1993 and 1995. In years 1 (1993) and 3 (1995) of the experiment, more Coleoptera species occurred in the tall sward (an average of nine species in addition to the forty‐one species present in the sward with the conventional sward height). Continuously grazed as opposed to ensiled subplots supported more beetle species but fewer individuals. Species composition of ground (Carabidae) and rove (Staphylinidae) beetles varied between treatments more than the arithmetic differences in species number. The experimental results supported the hypothesis but the benefits of taller swards to species diversity were small in the sown pastures of the study compared with indigenous upland grasslands (c. 33% fewer species). Inheritance effects of drainage, fertilizer and lime inputs, and the different species and management of cultivated pastures, may constrain the conservation benefits of altered pasture management compared with indigenous grasslands.     

Changes in plant densities in a mesic species-rich grassland after imposing different grazing management treatments

Changes in plant density were evaluated monthly in the first three vegetation seasons after imposing different grazing management treatments on abandoned semi‐natural grassland in the Czech Republic. There was no agricultural management in the 5 years before the start of the experiment in 1998. A completely randomized block experiment was established with the following five treatments: unmanaged control, intensive continuous grazing, extensive continuous grazing and a harvest in June followed by either intensive or extensive continuous grazing for the rest of the growing season. The sward was maintained at a target height of 5 and 10 cm under the intensive and extensive grazing managements respectively. An almost immediate increase in the densities of all sward components, especially grass tillers, occurred after the introduction of grazing on the previously abandoned grassland in comparison with the unmanaged control treatment. Trifolium repens was able to colonize and increase the number of its stolon growing‐points in all managed treatments, particularly in intensively grazed patches during the second and third experimental seasons. Delay to defoliation in both treatments containing a harvest in June resulted in an increase in the number of forb plants, particularly in the number of Taraxacum spp. plants, most probably due to an enabling of its seed production. It is evident that increases in plant density as a function of intensive defoliation are not restricted to the frequently documented effect on grass tillers but also can occur in many legume and forb species in species‐rich grasslands.     

Effects of sheep stocking on the plant community and agricultural characteristics of upland Anthoxanthum odoratum–Geranium sylvaticum meadow in northern England

In the UK, upland hay meadows (Anthoxanthum odoratum–Geranium sylvaticum grassland) with high plant biodiversity are rare and confined to submontane areas of northern England. We report results from a 5‐year experiment to test suggestions that recent biodiversity declines were attributable to increased sheep‐stocking density and a longer spring stocking period, thus delaying the shut up date for the growth of the hay crop. Longer stocking periods and higher stocking densities decreased the forage mass at 8 July, but they increased herbage N content and digestibility, reduced plant species diversity and reduced populations and seed production of Rhinanthus minor. Compared with unstocked swards, the similarity of the vegetation to Anthoxanthum odoratum–Geranium sylvaticum grassland was reduced by 16·9% when stocking with sheep continued until 27 May each year, and by 8·3% when sward heights were maintained at 3 cm compared with 5 cm. Increased mean sward height and height of R. minor were positively correlated with accumulated temperatures. Results support suggestions that recent reductions in the nature value of these grasslands might be a consequence of high stocking densities persisting until later in the spring, carried out during a 1‐year period with warmer temperatures.     

Cutting frequency vs. N application: effect of a 20‐year management in Lolio‐Cynosuretum grassland

The aim of this study was to determine the effects of different long‐term management options on re‐establishment of traditional species‐rich grassland in a Lolio‐Cynosuretum grassland. The experiment was set up in 1987 in a low‐fertilized mown pasture in the Eifel Mountains (Germany). Two and four cuts per year each with three levels of nitrogen fertilizer application were compared to no application of P and K fertilizer. Biomass production increased as a result of N application, whereas crude protein content and digestibility of organic matter improved in the four‐cut treatments. During a 20‐year period, we found a significant shift in dominant grass species without any significant effect on species richness. Lolium perenne and Elytrigia repens were replaced by Holcus lanatus and Alopecurus pratensis, especially in the two‐cut treatments with N application. In the unfertilized plots, several individuals of Platanthera bifolia appeared when swards were cut only twice per year, probably as a result of nutrient depletion together with low cutting frequency. An increase in number of cuts mainly supported plant species well adapted to intensive management, even under low N application and relatively low plant‐available P content in the soil. Therefore, we conclude that cutting frequency was a key driver on plant species composition in this experiment. The re‐establishment of traditional species‐rich grassland from intensively managed Lolio‐Cynosuretum was not achieved within 20 years of cutting management combined with absence of P and K application.     

Spatial and structural uniformity of lowland agricultural grassland in England: a context for low biodiversity

The contention that agricultural grassland within the UK is spatially and structurally uniform across wide differences in livestock farming intensity is examined. Total nitrogen (N) input was used as a surrogate for management intensity. Farms with average annual N inputs of >200 kg ha^?1 were categorized as highly intensive, those with N inputs of 50–200 kg N ha^?1 as moderately intensive and those with inputs of <50 kg N ha^?1 were categorized as extensive. Four farms within each management category were selected in two discrete regions: one in south‐west England, typifying a landscape dominated by agricultural grassland, the other in south‐east England with a more mixed‐farming landscape. Specific N input and management data, and sward botanical composition and structure, were obtained on four fields per farm. The south‐west region had a higher proportion of farms with dairy cattle than in the south‐east England region. The average N input and stocking rates were higher within the intensive‐management category farms in south‐west England than in south‐east England. Old permanent grasslands contained variable amounts of sown species, such as Lolium perenne, and were dominated by a few (generally <5) non‐sown grass species. Grassland that received >75 kg N ha ^?1 year^?1 contained <7 plant species and <3 forb species m^?2. Grassland communities with >12 plant species and >5 forb species m^?2 were only found in study fields that received <50 kg N ha^?1 year^?1. Grasslands with >10 forb species m^?2 were found only in sites receiving <15 kg N ha^?1. Of importance at a landscape scale was ubiquity of species‐poor and similar grassland plant communities across the management intensity range within and between livestock farms. Grassland in the extensive management category was similar in structure in terms of sward height and/or sward density to more intensively managed grasslands. This study revealed, albeit in two lowland sample regions in the UK, the ubiquity of grass‐dominated, species‐poor and structurally uniform grasslands, irrespective of apparently broad differences in farming intensity. The consequences of this spatial uniformity on grassland biodiversity are likely to be profound.     

Grassland responses to grazing disturbance: plant diversity changes with grazing intensity in a desert steppe

This study quantifies the impact of four different grazing regimes (heavy, moderate, light and ungrazed) on the vegetation dynamics of rangeland ecosystems along the southern boundary area of the Mu Us Desert, China. As the grazing intensities decreased, the soil quality, canopy cover, height, density, above‐ and below‐ground biomass, litter, root/shoot ratio and native plant (Aneurolepidium dasystachys) and grass abundances significantly increased; the above‐ground biomass of grasses increased, but the above‐ground biomass of forbs decreased. Ungrazed grassland has significantly improved from grasslands experiencing three other levels of grazing pressure, especially in the grassland biomass. Species richness increased as the grazing intensity decreased in the grazing grasslands, but peak species richness appeared under moderate and light grazing against lower productivity. Grazing exclusion causes desirable transitions in plant communities of desert steppe rangelands. Therefore, appropriate and efficient grazing exclusion is an available way to counteract local grassland degradation and promote rangeland sustainability.     

Plant species richness, functional type and soil properties of grasslands and allied vegetation in English Environmentally Sensitive Areas

To facilitate the maintenance and restoration of semi‐natural grasslands, it is important to understand their relationships with soil properties. Semi‐natural grasslands typically have a high incidence of stress‐tolerant species (measured here by high stress radius values), but not all have high species richness. Species richness and stress radius values were related to soil pH, Olsen extractable phosphorus (P), extractable potassium (K) and magnesium (Mg), total nitrogen (N) and organic matter (OM) at 571 sites representing a wide range oftemperate grasslands. Highest species richness (>30 m^?2) occurred at pH > 6 and 4–15 mg l^?1 P, but species richness was also highly variable at 4–15 mg l^?1 P. At pH < 5, species richness was low (<20 m^?2). Stress radius values were highest (mainly calcareous and heath grasslands and mires) at pH c. 8·0 and < 5·0, and at the lowest soil P levels (<5 mg l^?1). A wide range of stress radius values occurred at low soil P levels because appropriate management is also needed to maintain semi‐natural grasslands. Reducing soil P is difficult in practice, so grassland restoration in the presence of elevated soil‐extractable P levels merits re‐assessment.     

Productivity,grazing utilization,forage quality and primary production controls of species‐rich alpine grasslands with Nardus stricta in northern Spain

Species‐rich alpine grasslands with Nardus stricta are important communities for both animal production and environmental conservation in Europe. We selected two contrasting types of Nardus grasslands (mesic and wet) within a rangeland of northern Spain and measured annual above‐ground net primary productivity (ANPP), botanical components, forage utilization and their respective seasonal patterns, during a 5‐year period. We analysed their chemical properties and recorded soil moisture and temperature in order to construct models able to explain grassland productivity. Mean annual ANPP of mesic Nardus grassland was about half (216 g DM m^?2 year^?1; ±29·8 s.e.) that of the wet grassland (406 g DM m^?2 year^?1; ±54·3 s.e.), with significant intra‐ and interannual variability. Mesic grassland, with a more important contribution of forbs and legumes over graminoids in its botanical composition, was the preferred forage source of grazing livestock and showed better chemical properties in spring and early summer. In summer and autumn, wet grassland had a higher utilization owing to its ability to maintain high biomass production. This was partially explained by soil moisture, a limiting factor of mesic grassland productivity. Our results provide new and relevant information on key aspects of species‐rich alpine Nardus grasslands, potentially useful for the definition of management options for these habitats of priority conservation.     

Reintroduction of grazing management after deforestation of formerly abandoned grassland and its effect on early vegetation changes in the Western Carpathians (Slovakia)

Although the process of reforestation of grassland has been widely studied in Europe, little is known about the effect of deforestation on grassland development. Thus, the specific objective of this study was to evaluate early changes in plant species composition, functional group, yield and biomass quality after deforestation of long‐term abandoned pastures. The experiment was established immediately after deforestation on sparse herbaceous vegetation (mean initial cover 27%) with the following treatments: grazing management only (G0), cutting and grazing aftermath (CG), grazing after seeding of grassland mixture (GS), grazing after a burning treatment in which branches were burned after deforestation (GB) and unmanaged control (U). Very rapid recovery of bare ground by germination and/or sprouting of grassland species was similar under all types of grazing management. Total plant species richness increased in all managed treatments except GB. Similarities according to redundancy analyses in plant species composition were found among G0, CG and GB treatments, especially for forbs with correlated rosette or creeping growth. The woody species, tall grasses and tall forbs had higher abundance in the U treatment. The restoration of grassland following deforestation of formerly reforested grassland area by grazing management was a relatively fast process, and swards were created after 3 years. The highest biomass yield was observed under treatments GS and GB. Forage quality of all managed treatments was sufficient for the demands of beef cattle grazing. However, for subsequent grassland preservation, some type of grazing management is necessary to prevent reforestation, which can occur immediately after deforestation in unmanaged places.     

The conservation management of upland hay meadows in Britain: a review

Upland hay meadows conforming to MG3 in the National Vegetation Classification of the UK are a rare habitat in Britain and are largely confined to upland valleys in northern England. Agricultural intensification, particularly ploughing and reseeding and a shift from hay‐making to silage production over the last 50 years, has resulted in large losses of species‐rich upland hay meadows. Remaining species‐rich meadows have been the focus of much nature conservation effort resulting in many of the species‐rich sites being protected by statutory designations or through voluntary agri‐environment scheme agreements. Research and monitoring has tended to confirm that species richness is maximized by management involving spring and autumn grazing, a mid‐July hay cut, no inorganic fertilizer and possibly low levels of farmyard manure. Deviations from this regime result in a loss of species richness. Restoration of semi‐improved grassland to swards resembling species‐rich MG3 also requires a similar regime but is also dependent on the introduction of seed of appropriate species. The role of Rhinanthus minor as a tool for manipulating meadow biodiversity during restoration management is discussed. Suggestions for future research are outlined.     

Response of plant species composition,biomass production and biomass chemical properties to high N,P and K application rates in Dactylis glomerata‐ and Festuca arundinacea‐dominated grassland

Little is known about the immediate effect of high nitrogen (N), phosphorus (P) and potassium (K) application rates on sown grasslands cut twice per year. We asked how quickly plant species composition, biomass yield, biomass chemical properties and nutrient balance respond to N, P and K application. An experiment using unfertilized control, P, N, NP and NPK treatments was established on seven‐year‐old cut grassland in the Czech Republic in 2007 and monitored over four years. Annual application rates were 300 kg N ha^?1, 80 kg P ha^?1 and 200 kg K ha^?1. The immediate response of plant species composition to N application was recorded and was found to be different to the response over the four years of the study period. Highly productive grasses (Dactylis glomerata, Festuca arundinacea and Phleum pratense) were promoted by N application in 2008 and then retreated together with legumes (Medicago sativa, Trifolium pratense and Trifolium repens) in all N treatments where the expansion of perennial forbs (Urtica dioica and Rumex obtusifolius) and annual weeds (Galinsoga quadriradiata, Impatiens parviflora, Lamium purpureum and Stellaria media) was recorded. At the end of the experiment, Festuca rubra was the dominant grass in the control and P treatment, and species richness was lowest in all treatments with N application. Mean annual dry‐matter yield over all years was 3.5, 3.9, 5.8, 5.6 and 6.8 t ha^?1 in the control, P, N, NP and NPK treatments, respectively. Concentrations of N in the biomass ranged from 20.0 to 28.7 g kg^?1 in the P and N treatments; concentrations of P ranged from 3.2 to 3.7 g kg^?1 in the N and P treatments; and concentrations of K ranged from 24.1 to 34.0 g kg^?1 in the NP and NPK treatments. The N:P, N:K and K:P ratios did not correctly indicate the nutrient limitation of biomass production, which was primarily N‐limited, and K‐limitation was only recorded for high production levels in treatments with N applications. On the basis of the nutrient‐balance approach, the balanced annual application rates were estimated as 140 kg N ha^?1, 30 kg P ha^?1 and 100 kg K ha^?1. We concluded that high N, P and K application rates can very quickly and dramatically change species composition, biomass production and its chemical properties in sown cut grasslands. High N application rates can be detrimental for tall forage grasses and can support the spread of weedy species.     

Impacts of elevated atmospheric CO2 and temperature on plant community structure of a temperate grassland are modulated by cutting frequency

To determine the impacts of climate change and defoliation on the community structure and plant diversity of a semi‐natural temperate grassland, monoliths of a permanent grassland were exposed to ambient or elevated atmospheric CO₂ concentrations (ambient + 235 ppmv) and temperature (ambient + 3°C) from October 1998 to December 2000. The monoliths were subjected to two different cutting frequencies, either two or six cuts per year. The grassland community structure changed during the course of the experiment and was more responsive to changes in management than to changes in climate. Increased cutting frequency stimulated plant diversity by enhancing the number of forb species, but plant diversity was not significantly affected by climate change. The contribution of individual plant species to the vegetation cover revealed species‐specific responses to climate change and cutting frequency, but for most species significant interactions between climate change and cutting frequency were present. There were no clear‐cut effects of treatments on the total annual yield and the proportion of forbs present, as significant interactions between climate change and cutting frequency occurred. It is concluded that differential grassland management will modify plant species‐specific responses to climate change and resulting changes in the botanical composition of mixed‐species, temperate grasslands.     

Model predicting dynamics of biomass, structure and digestibility of herbage in managed permanent pastures. 2. Model evaluation

A mechanistic model, simulating the dynamics of production, structure and digestibility of managed permanent pastures, was developed. Its evaluation consisted of (i) studying model response to a range of grassland communities, cutting frequencies and site characteristics, and (ii) testing the model against experimental data, focusing on biomass accumulation and digestibility during three different cutting cycles, herbage production under a frequent cutting regime, and sward dynamics during the winter. The model realistically predicted the dynamics of biomass, structure and digestibility of herbage for various communities of permanent pastures, in different sites and under different management conditions for upland areas of the Auvergne region in France. The predicted responses to environmental conditions and cutting regimes were close to field observations and experimental results. Although the model successfully predicted the dynamics of average herbage production, it lacked precision in predicting the low biomass production observed in relation to the weather conditions found in a few specific years. The model was able to predict the dynamics of the sward during winter and is, therefore, fit for producing multiple‐year simulations. To improve the prediction of variability of biomass production and to predict the medium‐ to long‐term dynamics of permanent pastures, the model could be refined by adding seasonal and multiple‐year variation in nitrogen availability and in the proportion of grass functional groups in the grassland community.     

The conservation management of mesotrophic (meadow) grassland in northern England. 1. Effects of grazing, cutting date and fertilizer on the vegetation of a traditionally managed sward

The results are reported from an experiment on the effects of cutting date (14 June, 21 July and I September), fertilizer application (none or 80 kg ha^?1 N plus 40 kg ha^?1 P and K) and grazing treatments (none, autumn or autumn plus spring) on the vegetation of an upland mesotrophic grassland in Upper Teesdale. northern England, UK. Effects on plant species number and cover are reported for 4 years (1989–93) of treatment. Effects on ‘species -attributes’ are given for the fourth year. The cessation of grazing combined with the use of fertilizer progressively reduced species number by about 25%. Under traditional management (no fertilizer, cutting date on 21 July, autumn and spring grazing) the species number and cover remained relatively static over the 4 years. Comparison between treatments in the fourth year showed a reduction in species number under the fertilizer application, cutting date on 1 September and no-grazing treatments. Fertilizer use together with cutting date on 1 September particularly lowered species number and cover. Analysis of variance was used to assess the effect of treatment on species that occurred frequently in the sward. A cutting date of 1 September favoured Agrostis capillaris. Alopecurus pratensis, Poa trivialis, Phleum pratense and Trisetum flavescens, The absence of grazing favoured Dactylis glomerata and Holcus lanatus. The use of fertilizer particularly favoured A. pratensis and H. lanatus. Ordination methods were used to assess the effect of treatment on the less frequent species. These were primarily associated with the treatment combination that matched ‘traditional’ management. Deviations from this ‘traditional’ regime acted separately, rather than in combination, and favoured different grass species. Traditional management was associated with ruderal, stress-tolerant ruderal and competitive ruderal strategists and with longer seed germination times, heavier seeds, some of which needed scarifying or chilling to break dormancy, and transient seed banks that germinated in the autumn. The original sward was an Anthoxanthum odora-turn-Geranium sylvatirum grassland, Briza media subcommunity (MG3b). After 4 years, Festuca ovina-Agrostis capillaris-Galium saxatile grassland, Holcus lanatus-Trifolium repens subcom-munity (U4b) and Lolium perenne-Alopecurus pratensis-Festuca pratensis grassland (MG7c) were found in many of the fertilized and late-cutting treatments.     

The impact of cutting,liming and fertilizing on characteristics of abandoned upland meadows in the Czech Republic

The responses of three upland grassland phytocoenoses to three different management treatments were studied in Southern Bohemia. The sites were at an advanced successional stage, with all three being meadows abandoned since World War II: (i) Molinion, (ii) degraded Calthion with dominant Carex brizoides and (iii) plant community with the dominant species Calamagrostis villosa. Three different treatments (cutting, liming and fertilizing) were applied to the three grassland types, and their effects on the diversity, plant cover and biomass production were investigated over a 3‐year period. Analyses showed significant differences between meadow types in time as well as response to treatments. Independent of the treatments, the biomass production and Shannon index increased significantly in all meadows by varying degrees during the investigation period, whereas the cover decreased slightly in the case of the Carex community and the Molinion and remained unchanged in the case of the Calamagrostis community. Changes in the ground cover (as percentage) in response to management treatments varied between the meadow types. Limed and fertilized plots of the Molinion and Calamagrostis communities had higher percentage cover values compared to the control, but the cover of the C. brizoides community was not affected. In contrast, cut plots of the Molinion had slightly reduced cover. The changes in biomass and species diversity showed no significant correlation with the types of treatment used.     

Effect of low‐dose N application and early mowing on plant species composition of mesophilous meadow grassland (Arrhenatherion) in Central Europe

Semi‐natural temperate grasslands in Europe make an important contribution to biodiversity, but their existence has been endangered by both agricultural intensification and abandonment. In the Czech Republic, Arrhenatherion meadows are a typical example, with their conservation made difficult by the scarcity of research on effective management practices. We examined effects of two potentially useful approaches, application of 56 kg N ha^?1 and earlier first mowing (i.e. in the middle of May, about 2 weeks earlier than usual), on species composition of an Arrhenatherion grassland in central Bohemia over a 7‐year period (1998–2004). These treatments did not influence species richness, but resulted in small changes in canopy structure in terms of cover of individual species and particular plant functional groups. N application resulted in increases in nitrophilous species and tall graminoids, especially Alopecurus pratensis, Festuca pratensis, Heracleum sphondylium and Ranunculus acris, and in decreases in short graminoids. Earlier mowing decreased tall herbs, especially H. sphondylium, and might thus be useful in compensating for increases in some nitrophilous species owing to N application. However, earlier mowing had negative impacts on spring‐flowering species in terms of reduced cover.     

Biomass production of upland vegetation types in England and Wales

Abstract The objective of the study was to describe the annual and seasonal production of live biomass of six plant species or communities of the uplands of the UK –Calluna vulgaris, Vaccinium myrtillus, Nardus stricta, Molinia caerulea, Eriophorum vaginatum and Agrostis–Festuca grassland. The species or communities are important for the management of the uplands to meet both biodiversity and agriculture objectives. The annual and seasonal production of live biomass was determined for sites within six regions of England and Wales in 1995, 1996 and 1997. The six regions were North‐east England, North‐west England, North Pennines, South Pennines, South‐west England and Wales. In each region, the Institute of Terrestrial Ecology's Land Classification Scheme was used to select three or four 2 km × 2 km squares that contained a mosaic of C. vulgaris and upland grassland plant species. Uniform plots of 1 ha were selected from each 2 km × 2 km square, within which three measurements were made on each sampling occasion. Biomass of each species or community was measured within 1·0 × 0·5‐m quadrats using the same standard procedures. Measurements of annual production of current season's shoots of the pioneer, building, mature and degenerate phases of C. vulgaris and of V. myrtillus were made in October, and of the annual production of live biomass of N. stricta, M. caerulea and E. vaginatum in August. A simulated grazing regime was imposed for the Agrostis–Festuca grassland with cuts being made to a height of 4 cm at 8‐week intervals. The annual production of live biomass produced was estimated as the sum of four harvests taken during the growing season. Seasonal measurements of the amount of live biomass of all the species were also made in April, June, August and October. For C. vulgaris, there were significant (P = 0·05) differences between the pioneer, building, mature and degenerate phases in the annual production of current season's shoots, with greater amounts being produced by the phases of greater age and with higher biomass. There were significant year (P < 0·05) and region (P < 0·05) effects, with the lowest production in 1995 and in the South‐west of England. The values were higher than those observed in previous studies in Scotland, and these were interpreted as being associated with a longer growing season rather than higher summer temperatures. There was no difference between years and regions in the annual production of V. myrtillus. For the grass species, there were small differences in annual production of live biomass between years and regions. However, the annual production of live biomass of N. stricta was significantly (P < 0·01) lower in 1995 than in 1996 and 1997 and significantly (P < 0·001) lower in the South Pennines region than elsewhere. The effect of a range of weather and other environmental variables on the annual production of live biomass of each species of grass was explored, but they provided only a limited insight into the reasons for the observed differences and were found to be of limited value as predictors.     

Changes in the area of permanent grassland and its implications for the provision of bioenergy: Slovakia as a case study

Throughout Europe, grasslands are managed primarily for agricultural production but also provide a range of ecosystem services, the magnitude of which is influenced by their area, management and the abiotic properties of the ecosystem. The grassland area in Europe has been affected by significant changes in recent decades, including abandonment and conversion to arable land. This study presents an assessment of changes in the permanent grassland area with reference to Slovakia and provides an assessment of the bioenergy potential of permanent grassland surplus to agricultural needs. In 2015, of the total permanent grassland area (868,000 ha) 42% had ceased to be used as feed for milk or meat production and had been abandoned. Results from field experiments show that low‐level improvement of permanent grasslands, e.g., by oversowing commonly sown agricultural grassland species and low‐dose fertilization (up to 30 kg N ha^?1 year^?1) could provide herbage biomass of 1.7 million tonnes of herbage dry matter per year. This would be sufficient to generate an energy output of approximately 31.3–31.9 PJ/year (4.5% of the current gross inland energy consumption of Slovakia). Low‐intensity use of the currently surplus permanent grassland is also considered to have potential beneficial outcomes in terms of delivery of ecosystem services. Results are also discussed in the context of countries beyond the case‐study area.     

Spontaneous colonization of restored dry grasslands by target species: restoration proceeds beyond sowing regional seed mixtures

Dry grasslands are endangered habitats across Europe, and their restoration requires increasing attention, including their re‐creation on arable land. We studied spontaneous colonization by eleven target species on thirty‐five dry grassland sites restored with regional seed mixtures 1–13 years ago in the eastern part of the Czech Republic, a region where species‐rich dry grasslands often occur in the vicinity of the restored grasslands. Occurrence of the target species (not sown in the restored grasslands) was quantified inside and outside each restored grassland up to distances of 100 and 500 m, and the distance to the nearest population outside was assessed. Data were processed using univariate (Spearman rank‐correlation coefficient, generalized linear models) and multivariate (redundancy analysis) statistics. The establishment of all target species was closely related to their occurrence in the surroundings. Occurrence of species at a distance of up to 500 m, minimum distance to the nearest population and time since start of restoration significantly influenced the occurrence of all the species in the restored grasslands, while distance up to 100 m influenced all except two species. The most successful were wind‐dispersed Cirsium canum and Inula salicina and, the least successful was the myrmecochorous Euphorbia virgata. Seeding regional seed mixtures may be considered an appropriate first step in the restoration of species‐rich grasslands on arable land. Following diversification of the vegetation in restored grassland, spontaneous establishment of additional species may occur if the species occur in the vicinity. Results are discussed in the context of grassland restoration and agri‐environmental objectives.