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1.

Stem form and taper changes after thinning and thinning combined with N fertilization were studied in 23 Norway spruce [Picea abies (L.) Karst.] and 46 Scots pine (Pinus sylvestris L.) stands in Sweden observed for 5-28 yrs. Average taper was calculated for the stem section 1.3-6.0 m above ground level and as the ratio between the diameter at breast height and total tree height for periods of 5-10 yrs. At the start of the experiment, before the first thinning, the dominant height was 12-15 m. Once 65% of the basal area had been removed in a single thinning from below, the remaining trees showed a strong increase in taper compared with trees in unthinned stands. Stems in stands treated with heavily recurrent thinnings from below also increased in taper, although the increases were not always statistically significant. Trees in thinned, N-fertilized Scots pine stands in middle and northern Sweden developed a more pronounced taper compared with stems in equally thinned, unfertilized stands.  相似文献   

2.
Forest management strongly influences the interactions between ungulates and their food resources. Different ungulate-adapted measures have been proposed in forestry to improve forage availability or to reduce browsing damage. However, the potential and feasibility of such measures are inadequately known. We studied the effects of harvest timing and slash treatment in final felling and commercial thinning on the availability of Scots pine Pinus sylvestris forage and its use by ungulates during winter in the Swedish boreal forests. Pellet group counts showed that moose (Alces alces) was the dominating species using the post-harvest stands. Under conventional slash treatment, final felling stands held on average 226 kg pine forage ha?1 after harvesting and commercial thinning stands 137 kg ha?1. Ungulate-adapted slash treatment increased the available forage biomass by 20 %, but had no significant effect on consumption of forage by ungulates. Time since harvest had the strongest effect on forage consumption; for example, under conventional slash treatment, there was a tenfold increase in consumption (3 vs. 33 kg ha?1) following final felling as exposure time increased from 2–3 to 4–5 months. Consumption was higher in thinned stands than in final felling stands for the first 3 months but not later. To increase ungulate use of the forage made available at harvest, pine-dominated stands should be harvested in the late autumn or early in the winter.  相似文献   

3.
Precommercial thinning is regarded as one of the most important measures for influencing timber quality in stands of Scots pine (Pinus sylvestris L.). The present study considers the influence of both thinning intensity and tree height at the time of thinning on diameter of the thickest branch and crown ratio. A total of 115 plots (either naturally regenerated, planted or sown, and either thinned down to 528–8000 stems?ha?1 or untreated) included in 20 sites in southern Sweden were analysed. The average tree height after thinning varied from 1.2 to 8.3?m. An increasing number of remaining stems resulted in a reduction in branch diameter, although the reduction appeared to be only minor if the number of stems after thinning was more than 3000 stems?ha?1. It was found that late thinning reduced the diameter of the thickest branch. The crown ratio decreased with stand height, number of stems after thinning and average height at thinning. The results were consistent for all trees and for the 500 thickest trees per hectare.  相似文献   

4.
Incoming shortwave global radiation (Q g) and photosynthetically active radiation (PAR; Q pa as a fraction of full daylight, relative irradiance (%Q), were measured at the same time in young stands of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). Measurements were made on three levels above ground: 20 cm above ground and 50 and 15 % of stand height. Stands of three heights (75, 150 and 300 cm) were studied during two months. The stands were created by arranging young trees cut from natural stands, in nine quadratic spacings: 0.1, 0.2, 0.3, 0.4, 0.5, 0.7, 1.0, 1.4 and 2.0 m. The leaf area index (L) was estimated. Differences in %Q‐values for Q pa and Qg in the same species and at the same stand height and level of light measurements above ground were significant only in 150 and 300 cm stands. In 75 cm high stands of Scots pine, the %Q was <60% at spacings <0.2 m and in 75 cm high stands of Norway spruce, the %Q was <60% at spacings <0.3 m. Only at 0.1x0.1–0.3x0.3 m, %Q was <20% in Norway spruce stands. In 150 cm stands %Q<20% was measured at spacings <0.7 m in Norway spruce and <0.5 in Scots pine. In 300 cm stands of Norway spruce it was measured up to 1.2 and in Scots pine <1.1 m. Light extinction coefficients, K and Kg for spruce and pine stands were 0.17–0.40 and 0.16–0.31 respectively. Some practical implications of the study are presented. Lack of light (%Q<10%) as a single factor of seriously suppressed growth and development of broad‐leaved plants and suckers by competition in young stands of spruce and pine only occur in dense stands 0.3x0.3 m‐1.1xl.l m (8000–100000 stems/hectare). Competition by light on a regenerated area generally occurs in the level of 50% of tree height (150–300 cm) and higher due to the rapid growth of broad‐leaved trees (sprouts) compared with planted conifers but the light intensity at these levels %Q>10%.  相似文献   

5.
Natural mortality in a 30-year period was examined in thinning and fertilisation experiments with 48 blocks in Scots pine (Pinus sylvstris L.) and 23 blocks in Norway spruce (Picea abies (L.) Karst.) with up to 12 different treatments. Of about 90,000 living trees at start of the experiments 18.7% were registered as dead by natural mortality in the 30-year observation period. In non-thinned stands about 20% of the volume growth disappeared by natural mortality, in thinned stands about 10%. In normally thinned pine stands (repeated thinning from below with moderate intensity) the annual mortality of the basal area at start of an average 7-year period was 0.34%. In spruce stands, on more fertile sites, the corresponding figure was about 0.6%. In an effort to model the mortality, severe damage not leading to final felling was identified in 1.7% of the observation periods. It was assumed that this part of the mortality, representing 24% of the total volume mortality, could be recovered by active thinning. The probability for severe damage increased sharply with stand top height, as shown in a logistic regression. The more sparse mortality was expressed as a function of site fertility, stand density, disturbance by thinning and form of treatment (thinned from above or below or non-thinned). The naturally dead trees were approximately of mean size in normally thinned stands while the self-thinning in non-thinned stands tended to occur amongst smaller than average trees. Diagrams were presented for basal area development and stem number reduction in the non-thinned stands.  相似文献   

6.
The effects of timing and intensity of precommercial thinning were studied in three Scots pine artificially regenerated stands on Vaccinium forest sites in southern Finland. A two-level factorial design (3×3) was used in each stand: thinning at dominant height of 3, 6 and 9 m to 1000, 1600 and 2200 stems ha?1. The effects of the treatments were analysed after a period of 23–25 yrs when the dominant height was 14–15 m. Early thinning resulted in the highest standing volume and amount of merchantable wood, and also in slightly accelerated height development. Thinning to 1000 stems ha?1 caused a considerable production loss, but there were no differences between the densities of 1600 and 2200 stems ha?1. Branches became thicker after early thinning, but the differences between the treatments were negligible for crop trees. Crown ratio was lowest as the result of early or moderate thinning (2200 stems ha?1).  相似文献   

7.

New silvicultural regimes with high within-stand competition require new functions for estimation of standing stock and growth of biomass components, since the allometry of trees is changed by light competition. This paper presents functions for estimation of the aboveground biomass dry weights for stem wood, stem bark, branches and leaves of young (diameter at breast height <10 cm) Scots pine (Pinus sylvestris L.), Norway spruce [Picea abies (L.) Karst.] and birch (Betula pendula Roth. and Betula pubescens Ehrh.) trees growing in dense mixed stands. The functions were derived from a sample consisting of 84 Scots pine, 43 Norway spruce and 66 birch trees from six stands in northern Sweden with high stand densities (>10000 st ha-1). The logarithmically transformed power function displayed a good ability to stabilize the variance of dry weights and showed a good fit to the material (0.37< R 2 <0.99). A comparison with the most commonly used biomass functions in Sweden today showed that they overestimated the weight of stem wood and branches, while the weight of foliage was underestimated. The nature of these discrepancies suggested that the precision of biomass estimations might also be improved for young trees at wider spacing.  相似文献   

8.
This study compared visitor preferences of forestry professionals across six European countries (Sweden, Denmark, Great Britain, Austria, Romania and Portugal) using a questionnaire survey. The 598 interviewees were asked to rank photographs depicting recently thinned experimental plots in a 13-year old stand of pedunculate oak (Quercus robur L.) according to the criterion: “Which forest environment do you prefer as a visitor?” The plots represented five different residual stem densities: 7000 (no thinning, very high stem density), 5300 (heavy thinning, high stem density), 1000 (very heavy thinning, medium stem density), 300 (extremely heavy thinning, low stem density/open stand) and 100 (solitary trees, very low stem density/very open stand) stems ha?1. The results indicated geographical variation in the preferences for different thinning practices in young stands of oak. Portuguese, Austrian and Romanian respondents generally favoured thinned, but dense stands, whereas Danish and British respondents preferred very heavily thinned stands. Swedish respondents preferred open stands resulting from extremely heavy thinning. Photographs taken along rows were favoured to photographs across rows, indicating a preference for scenes offering perspective and accessibility. The results indicate a variation of visitor preferences among forestry professionals for different silvicultural regimes. We interpret this in the context of national traditions and forestry paradigms that influence the shaping of preferences.  相似文献   

9.
The three‐parameter Weibull function met specified statistical standards for goodness of fit as a model for the diameter distribution of moderately thinned Norway spruce stands in Denmark. Weibull distributions estimated by percentile estimators fit the majority of 522 observed diameter distributions (material made available by the Danish Forest Experiment Station) at the 10% level of significance in the Chi‐square and Kolmogoroff‐Smirnow tests. The Weibull distribution was less suited to describe the diameter distribution in unthinned stands and heavily thinned stands. Weibull parameter predictions were developed and used in connection with a Danish yield table for Norway spruce on clay‐rich coastal soils. The use of the Weibull distribution to predict stem frequencies in different diameter classes before and after a specified thinning of stands is demonstrated.  相似文献   

10.
Early management of the regenerated seedlings shapes the future stand properties. To address these issues, pre-commercial thinning (PCT) and control treatments were applied to planted Norway spruce (Picea abies L. Karst) and naturally regenerated birch (Betula pendula Roth., Betula pubescens Ehrh.) stands in forest experiments in southern Sweden (lat. 56–57?N) containing 1.1–5.5?m tall saplings. The treatments were retention of 1000 or 2000?stems?ha?1 of Norway spruce, with no birch or birch at 1000?stems?ha?1. Treatments were replicated with and without annual removal of birch sprouts from stumps. The periodic annual increment (PAI) over five years was calculated for total stand volume and individual trees. The mean PAI of dominant trees was significantly higher both following all PCT treatments than controls, and following low rather than high-density PCT. Birch retention did not affect growth of the dominant trees but PAI was lower in plots with uncontrolled sprouting. The PAI of birch was significantly higher in low-density Norway spruce plots than in control plots and the high-density plots. The treatment response was significant even in stands with initial heights of only 1–2?m.  相似文献   

11.
Eleven seral, postfire forest stands in southern Yukon (Canada) were sampled to determine where western white spruce (Picea albertiana ssp. albertiana) seedlings occurred with respect to distance to the nearest lodgepole pine (Pinus contorta var. latifolia) tree. Seedling-to-nearest tree distances were assessed at 10-cm increments up to 220?cm. On average, seedlings occurred 54?cm from the nearest pine (n?=?490), but peak frequencies were 20–50?cm away, compared to a potential separation distance of 103?cm. Greatest average seedling density occurred 10–20?cm from pine (0.81?m?2), with values between 10 and 120?cm decreasing logarithmatically with increasing distance from pine (r?=?0.994, p?n?=?11). Spruce seedling densities were <0.02?m?2 beyond 120?cm. The differences in frequency, which represented moderately strong aggregation (Clark-Evans Index 0.34–0.52), and density suggest greater spruce recruitment near lodgepole pine was facilitated by more favorable ecological conditions than further away, for example, greater nutrient availability. The bias in seedling-to-nearest tree distances occurred regardless of stand age (57–165 years), pine density (1599–5935 stems?ha?1), or understory vegetation type, although the bias may be weakened by the abundant presence of feathermosses (Hylocomium splendens) on the forest floor.  相似文献   

12.
The effects of silvicultural treatments on carbon sequestration are poorly understood, particularly in areas like the Mediterranean where soil fertility is low and climatic conditions can be harsh. In order to improve our understanding of these effects, a long-term thinning experiment in a stand of Mediterranean maritime pine (Pinus pinaster Ait.) was studied to identify the effects of thinning on soil carbon (forest floor and mineral soil), above and belowground biomass and fine and coarse woody debris. The study site was a 59-year-old pinewood, where three thinnings of differing intensities were applied: unthinned (control), moderate thinning and heavy thinning. The three thinning interventions (for the managed plots) involved whole-tree harvesting. The results revealed no differences between the different thinning treatments as regards the total soil carbon pool (forest floor + mineral soil). However, differences were detected in the case of living aboveground biomass and total dead wood debris between unthinned and thinned plots; the former containing larger amounts of carbon. The total carbon present in the unthinned plots was 317 Mg ha?1; in the moderately thinned plots, it was 256 Mg ha?1 and in the case of heavily thinned plots, 234 Mg ha?1. Quantification of these carbon compartments can be used as an indicator of total carbon stocks under different forest management regimes and thus identify the most appropriate to mitigate the effects of global change. Our results indicated that thinning do not alter the total soil carbon content at medium term, suggesting the sustainability of these silvicultural treatments.  相似文献   

13.
Abstract

The success of natural regeneration on patch clear-cuts in Norway spruce stands in Southern Finland was examined in terms of stocking and tree height. The experiment was established in mature spruce stands in the submesic Myrtillus (MT) site type. In each of the eight study sites, three plots were treated with clear-cutting and planting, clear-cutting with partial tree retention and planting, and patch clear-cutting without site preparation, respectively. In three of the study sites, one plot with patch clear-cutting and site preparation was established. Each plot was 1 ha, out of which three patches of 40×40 m were clear-cut in the patch treatments. 10–11 years after cutting, the patches had on average 1316 crop trees ha–1, of which 91% were spruces. Some 27% of the stands were up to the target stocking level (≥1600 ha?1), and 36% were at least satisfactory (≥1300 ha?1). Site preparation did not yield greater stocking levels on patches, but that result is ambiguous due to a difference in initial stocking. The average spruce tree height in the patches (0.76 m) was much smaller than in the case of clear-cutting and planting (2.42 m). In conclusion, the patches had been restocked tolerably well with spruce and birch for practical purposes during the 10- to 11-year period, but the regeneration process had been very slow compared to clear-cutting and planting.  相似文献   

14.
Long-term management impacts on carbon storage in Lake States forests   总被引:2,自引:0,他引:2  
We examined carbon storage following 50+ years of forest management in two long-term silvicultural studies in red pine and northern hardwood ecosystems of North America’s Great Lakes region. The studies contrasted various thinning intensities (red pine) or selection cuttings, shelterwoods, and diameter-limit cuttings (northern hardwoods) to unmanaged controls of similar ages, providing a unique opportunity to evaluate long-term management impacts on carbon pools in two major North American forest types. Management resulted in total ecosystem carbon pools of 130-137 Mg ha−1 in thinned red pine and 96-177 Mg ha−1 in managed northern hardwoods compared to 195 Mg ha−1 in unmanaged red pine and 224 Mg ha−1 in unmanaged northern hardwoods. Managed stands had smaller tree and deadwood pools than unmanaged stands in both ecosystems, but management had limited impacts on understory, forest floor, and soil carbon pools. Total carbon storage and storage in individual pools varied little across thinning intensities in red pine. In northern hardwoods, selection cuttings stored more carbon than the diameter-limit treatment, and selection cuttings generally had larger tree carbon pools than the shelterwood or diameter-limit treatments. The proportion of total ecosystem carbon stored in mineral soil tended to increase with increasing treatment intensity in both ecosystems, while the proportion of total ecosystem carbon stored in the tree layer typically decreased with increasing treatment intensity. When carbon storage in harvested wood products was added to total ecosystem carbon, selection cuttings and unmanaged stands stored similar levels of carbon in northern hardwoods, but carbon storage in unmanaged stands was higher than that of thinned stands for red pine even after adding harvested wood product carbon to total ecosystem carbon. Our results indicate long-term management decreased on-site carbon storage in red pine and northern hardwood ecosystems, but thinning intensity had little impact on carbon storage in red pine while increasing management intensity greatly reduced carbon storage in northern hardwoods. These findings suggest thinning to produce different stand structures would have limited impacts on carbon storage in red pine, but selection cuttings likely offer the best carbon management options in northern hardwoods.  相似文献   

15.
A series of 15 field experiments was established to quantify the growth response of first‐thinning stands of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst) to whole‐tree harvesting and to estimate the need for nutrient compensation. The experiments were undertaken in Finland, Norway and Sweden and represent a wide range of site conditions. The site index (H 100) of Scots pine stands varied from 19 to 29 m, and that of Norway spruce stands from 28 to 36 m. Total amounts of biomass and nutrients removed were calculated based on data obtained from felled sample trees. During the first 5‐yr period the growth response to the removal of logging residues varied considerably in both pine and spruce stands. Regression analyses did not reveal any functions that explained the variation in results satisfactorily. In cases where whole‐tree harvesting influenced tree growth negatively, this effect was counteracted by compensatory fertilization. It was concluded that to determine the response of remaining trees to harvesting intensity reliably, the post‐harvest period analysed must be longer than 5 yrs.  相似文献   

16.
The Warner Mountains of northeastern California on the Modoc National Forest experienced a high incidence of tree mortality (2001–2007) that was associated with drought and bark beetle (Coleoptera: Curculionidae, Scolytinae) attack. Various silvicultural thinning treatments were implemented prior to this period of tree mortality to reduce stand density and increase residual tree growth and vigor. Our study: (1) compared bark beetle-caused conifer mortality in forested areas thinned from 1985 to 1998 to similar, non-thinned areas and (2) identified site, stand and individual tree characteristics associated with conifer mortality. We sampled ponderosa pine (Pinus ponderosa var ponderosa Dougl. ex Laws.) and Jeffrey pine (Pinus jeffreyi Grev. and Balf.) trees in pre-commercially thinned and non-thinned plantations and ponderosa pine and white fir (Abies concolor var lowiana Gordon) in mixed conifer forests that were commercially thinned, salvage-thinned, and non-thinned. Clusters of five plots (1/50th ha) and four transects (20.1 × 100.6 m) were sampled to estimate stand, site and tree mortality characteristics. A total of 20 pre-commercially thinned and 13 non-thinned plantation plot clusters as well as 20 commercially thinned, 20 salvage-thinned and 20 non-thinned mixed conifer plot clusters were established. Plantation and mixed conifer data were analyzed separately. In ponderosa pine plantations, mountain pine beetle (Dendroctonus ponderosae Hopkins) (MPB) caused greater density of mortality (trees ha−1 killed) in non-thinned (median 16.1 trees ha−1) compared to the pre-commercially thinned (1.2 trees ha−1) stands. Percent mortality (trees ha−1 killed/trees ha−1 host available) was less in the pre-commercially thinned (median 0.5%) compared to the non-thinned (5.0%) plantation stands. In mixed conifer areas, fir engraver beetles (Scolytus ventralis LeConte) (FEN) caused greater density of white fir mortality in non-thinned (least square mean 44.5 trees ha−1) compared to the commercially thinned (23.8 trees ha−1) and salvage-thinned stands (16.4 trees ha−1). Percent mortality did not differ between commercially thinned (least square mean 12.6%), salvage-thinned (11.0%), and non-thinned (13.1%) mixed conifer stands. Thus, FEN-caused mortality occurred in direct proportion to the density of available white fir. In plantations, density of MPB-caused mortality was associated with treatment and tree density of all species. In mixed conifer areas, density of FEN-caused mortality had a positive association with white fir density and a curvilinear association with elevation.  相似文献   

17.
Optimal management of Korean pine plantations in multifunctional forestry   总被引:1,自引:0,他引:1  
Korean pine is one of the most important plantation species in northeast China.Besides timber,it produces edible nuts and plantations sequester carbon dioxide from the atmosphere.This study optimized the management of Korean pine plantations for timber production,seed production,carbon sequestration and for the joint production of multiple benefits.As the first step,models were developed for stand dynamics and seed production.These models were used in a simulation–optimization system to find optimal timing and type of thinning treatments and optimal rotation lengths.It was found that three thinnings during the rotation period were optimal.When the amount or profitability of timber production is maximized,suitable rotation lengths are 65–70 years and wood production is 5.5–6.0 m~3 ha~(-1) a~(-1).The optimal thinning regime is thinning from above.In seed production,optimal rotation lengths are over 100 years.When carbon sequestration in living biomass is maximized,stands should not be clear-cut until trees start to die due to senescence.In the joint production of multiple benefits,the optimal rotation length is 86 years if all benefits(wood,economic profits,seed,carbon sequestration) are equally important.In this management schedule,mean annual wood production is 5.5 m~2 ha~(-1) and mean annual seed yield 141 kg ha~(-1).It was concluded that it is better to produce timber and seeds in the same stands rather than assign stands to either timber production or seed production.  相似文献   

18.

Mean age, mean and top heights and yield were studied in 20 mixed stands of birch ( Betula pubescens Ehrh. and B. pendula Roth) and Picea abies (L.) Karst. and nine mixed stands of birch and Pinus sylvestris L. in south-eastern Norway. Each mixed stand and the adjacent pure coniferous stand (control) were growing under the same site conditions and had not been commercially thinned. There were no significant differences in mean age at breast height or in top heights between birch and conifers in the mixed stands, while mean height was significantly higher for birch than for spruce. A growth index was calculated based on total volume and age at breast height. For the spruce sites the growth in young mixed stands (birch < 17 m) was superior to that of pure spruce, while the difference was insignificant in older stands. The growth index correlated positively with the ratio between generatively and vegetatively regenerated birch trees, and negatively with the age of the oldest species in the mixture and with site quality. For the pine sites there was no significant difference in the growth index between mixed birch-pine and pure pine stands. A mixture effect of birch on the volume yield of spruce or pine could not be demonstrated.  相似文献   

19.
Growing concern about increasing concentrations of greenhouse gases in the atmosphere, and resulting global climate change, has spurred a growing demand for renewable energy. In this study, we hypothesized that a nurse tree crop may provide additional early yields of biomass for fuel, while in the longterm leading to deciduous stands that are believed to better meet the demands for other ecosystem services. Ten different species combinations were planted, with two different stocking densities, at three different sites in Denmark. Significant differences, with regard to biomass production, were observed among the different sites (P?P??1?yr?1 more biomass. The additional biomass production was similar to what was obtained in stands with conifers only (Sitka spruce, Douglas-fir and Japanese larch), which produced 4.9–6.1?t ha?1yr?1 more biomass than the pure beech stands. No effects of initial planting density (P?=?.19), or of initial weeding (P?=?.81), on biomass production were observed. Biomass production of the broadleaved crop was in most cases reduced due to competition. However, provided timely thinning of nurse trees, the qualitative development of the trees will allow for long-term timber production.  相似文献   

20.
Diameter, height and volume increment of Scots pine (Pinus sylvestrisL.) and Norway spruce (Picea abies (L.) Karst.) stands wereinvestigated in long-term thinning experiments in southern Finland.The measurement period was on average 19 years, and thinningsconsisted of five different treatments: (1) unthinned; (2) selectivethinning from below where thinning intensity was determinedaccording to the number of stems; (3) selective thinning frombelow where thinning intensity was determined according to standbasal area; (4) systematic thinning where corridors were openedat regular intervals in the stand; and (5) half-systematic thinningwhere corridors were opened at regular intervals and the remainingstrips thinned from below, i.e. a combination of selective andsystematic thinning. The selective (basal area) thinning ofScots pine resulted in a volume growth reduction of about 12per cent, but in Norway spruce it resulted in no marked reductioncompared with the unthinned plots. In both tree species, volumeincrement in the half-systematic and selective (basal area)thinning differed only slightly from each other. The systematicthinning resulted in the lowest volume increment. In the systematicthinning, the total yield (m3 ha–1) somewhat decreasedwith increasing corridor width. In half-systematic thinning,on the other hand, the total yield was not clearly related tocorridor width. A corridor accelerated the diameter incrementof the edge trees. The edge effect progressively diminishedwhen moving deeper into the strip and ended at a distance ofabout 3–4 m from the corridor edge. After the thinning,the height increment first decreased, but later on it recovered.The unthinned plots had more dead trees than the other treatments,but there were no significant differences between the othertreatments. It can be concluded that the prevailing thinningmethod in the Nordic countries, i.e. half-systematic thinning,most probably results in no significant growth and yield losses.  相似文献   

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