Affiliation: | 1.Graduate School of Environmental Science, Hokkaido University, Kita 10, Nishi 5, Kita-ku, Sapporo, Hokkaido, 060-0810, Japan ;2.Faculty of Fisheries Sciences, Hokkaido University, 3-1-1 Minato, Hakodate, Hokkaido, 041-8611, Japan ;3.Salmon and Freshwater Fisheries Research Institute, Hokkaido Research Organization, 3-373 Kitakashiwagi, Eniwa, Hokkaido, 061-1433, Japan ; |