Fine-scale vegetation patches decline in size and cover with increasing rainfall in Australian savannas |
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Authors: | Ludwig John A. Tongway David J. Eager Robert W. Williams Richard J. Cook Garry D. |
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Affiliation: | (1) Tropical Savannas Co-operative Research Centre and CSIRO Wildlife and Ecology, PMB 44, Winnellie, Darwin, NT, 0822, Australia;(2) CSIRO Wildlife and Ecology, PO Box 284, Canberra, ACT, 2601, Australia |
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Abstract: | Fine-scale vegetation patches (<5 m in width) are critically important in many landscapes because they function to obstruct surface flows of water and wind. These obstructions increase the infiltration of runoff and the capture of nutrients in runoff sediments and in wind-blown soil and litter. The importance of redistribution of runoff into runon patches from spaces between patches (fetches) is likely to be greater in drier than in wetter environments. In this paper we examine the hypothesis that the ratio of fetch to patch decreases as rainfall increases, and that this trend will be most evident on intermediate-textured soils because these soils are more prone to runoff. We measured fine-scale patches on 38 sites with sand, loam or clay soils. Sites were located along a 1000-mm rainfall gradient in the savannas of northern Australia. The width and intercept length of patches and the fetch between patches was measuring along line transects of 100–120 m oriented down slope. We found that the ratio of fetch to patch area did not decrease with decreasing rainfall, but increased on both sand and loam soils. This result was because with increasing rainfall mean spacing between patches disproportionally increased while mean patch size and cover declined. The cover of patches was negatively correlated with tree canopy cover, which significantly increased with rainfall. This negative correlation suggests that in higher rainfall savannas the size and spacing of ground-layer patches is controlled by the tree layer, and that as rainfall decreases this control decreases and runoff-runon processes increasingly structure the landscape. For savannas on clay soils these trends were not significant except that on the highest rainfall sites the cover of ground-layer patches was nearly 100% while trees were absent. |
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Keywords: | ground-layer landscape function patch spacing runoff soil texture tree-layer |
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