The components of fruit size in strawberry

High Correlation coefficients have been found in the application of the formula:

$\text{Fresh weight of ripe berry}\text{(g)}\text{=}\text{(Total number of achenes?C)F}\text{(Number of achenes per cm}{}^2\text{of surface}$

to data for a large number of strawberries from nine cultivars. Of the terms in the equation (Total no. of achenes — C) was found to be different in different cultivars and to be related to the position of the berry on the inflorescence, but there was no constant relationship between the achene numbers in the different ranks of berry. Whether the numbers of achenes per berry can be altered by environmental or cultural treatments was not resolved. F, a term having the dimensions of weight per area, represents weight per unit area of berry surface and seems to be constant amongst plants of a cultivar subject to uniform treatment, but differs between cultivars and treatments.The number of achenes per cm² of surface on ripe berries is a measure of berry development in the fruiting season and may be affected by environmental and managerial factors and by internal competition between fruits and leaves on the same plant.The interactions of these factors on berry size enable two stages to be recognised at which size is determined, namely during flower formation, when achene numbers and possibly F are fixed, and during fruit development after pollination when the term for spatial distribution of achenes is settled.It is suggested that an examination of the effects of different treatments on each of the size components might enable a distinction to be drawn between the influence on yield of genetic characteristics on the one hand and intrinsically variable characteristics subject to external influence on the other hand, thus clarifying the type of research and/or development needed to enable growers to achieve maximum yields.