小麦药壁表皮细胞与药室内壁细胞超微结构

电镜观察研究表明，在大液泡小孢子时期，小麦药壁表皮细胞含有发达的线体和高尔基体，开始产生嗜锇颗粒。药室内壁细胞具活力旺盛的叶绿体和丰富的细胞器。在花粉粒时期，药壁表皮细胞内充满嗜锇颗粒。药室内壁和花粉囊内也有少量与表皮细胞内相同的嗜锇颗粒。此外，在Ｕ氏体、绒毡层膜、花粉粒周围有大量的与药壁表皮细胞同质的小嗜锇颗粒。花粉成熟时，这些颗粒又减少。据此推测，药壁表皮、药室内壁在花粉发育过程中，尤其是发育后期有提供营养的作用。

Ultrastructure of Epidermis and Endothecium Cells of Auther Wall in Wheat

The result of ultrastructure observations for the epidermis and endothci-um cells of the auther wall in wheat with transmission electron microscopy(TEM)showed that,at vacuolate microspore stage,the epidermis cells of auther wall containedenergic chloroplasts and rich organells. At pollen stage,the epidermis cells was ftill ofthe osmiophilic globules,also there were small numbers of osmiophilic globules,similarto those in epideris cells,in endothecium and pollen sacs.In addition,there were num-berous small osmiophilic globules around the Ubisch bodies,orbicular wall and pollenwall,which resembled,in composition,to those in the epidermis cells.When the pollenbecame mature,these globules decreased.The results suggested that epidermis and en-dothecium of auther wall in wheat probably provide nutrient for the pollen development,specially for the later period of the development.