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The stand dynamics of a mixed coppice forest of shade-tolerant and intermediate species
Institution:1. Graduate School of Science and Technology, Niigata University Ikarashi 2-8050, Niigata 950-21, Japan;2. Faculty of Agriculture, Niigata University Ikarashi 2-8050, Niigata 950-21, Japan;1. Center for Orthopaedics and Trauma Surgery, University Hospital Giessen and Marburg, Location Marburg, Baldingerstrasse, 35043 Marburg, Germany;2. Center for Orthopaedics and Trauma Surgery, University Medical Center Cologne, Cologne, Germany;3. Department of Orthopaedic Sports Surgery, St. Josef-Hospital, Ruhr-University Bochum, Bochum, Germany;1. Erasmus Medical Center Stem Cell Institute, Department of Cell Biology, Erasmus Medical Center, Rotterdam, the Netherlands;2. Centre for Inflammation Research, Queens Medical Research Institute, University of Edinburgh, Edinburgh, UK
Abstract:We investigated stand dynamics of a mixed coppice forest of shade-tolerant and intermediate species, Fagus crenata Bl. and Quercus crispula Bl., in five stands with canopy-stem densities from 420 to 1320 ha−1 (estimated stand age: 37–62 years). The mean basal area (BA), exposed-crown area (ECA) and crown depth (CD) per stem were significantly larger in F. crenata in the three lower density plots. Also Q. crispula had considerably less overtopped stems in the two lower density plots. The mean annual BA increments of the two species were similar in the earlier 20 years; however, in the most recent 5 years, F. crenata had a higher relative growth rate for basal area (RGRBA) than Q. crispula. The most critical variable of the RGRBA was the ECA for F. crenata and the relative height (RH) for Q. crispula. We predict that dominance of F. crenata will gradually increase over that of Q. crispula as stands mature. This seems to be caused by lower shade-tolerance despite the similar initial growth rates of Q. crispula compared with F. crenata.
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