A1,A2型高粱雄性不育性核质互作遗传方式研究

用A1,A2不育系与相应的恢复系杂交,再用A1,A2不育系、保持系与A1,A2杂交种交叉杂交,对获得的F1,F2,BC1F1,BC1F2群体进行了育性遗传分析。结果表明,A1,A2雄性不育性均由两对基因控制,但A1,A2基因互不等位。A1雄性不育系的完整遗传模式为S1N2(ms1ms1ms1'ms1'ms2ms2ms2'ms2'),相应的保持系基因型为N1N2(ms1ms1ms1'ms1'ms2ms2ms2'ms2'),A2雄性不育系的遗传模式为N1S2(MS1MS1ms2ms2ms2'ms2'),相应的保持系基因型为N1N2(MS1MS1ms2ms2ms2'ms2')。A1,A2相应的恢复系核基因型各有6种,其表达方式有明确的对应性,在相应的不育位点,只要有其中一对就可实现对不育系的恢复。分析认为,高粱细胞核、细胞质内均有多种育性基因存在,但是细胞质内的不育基因和细胞核内的不育基因有些是有关系的,有些则是没有关系的,保养有细胞质和细胞核同时具备同样性质的不育基因时,不育特性才能表达,如有一方有可育基因存在,则表现可育,呈对应互作关系。

Genotype and Genetic Modle of A1 ,A2 Male - sterility in Sorghum

A_1 and A_2 male-sterile lines were crossed with restorer lines,A_1 and A_2 male-sterile and maitainer lines were crossed with the hybrids of A_1 and A_2 cytoplasm. The genetics of sexuality in F_1,F_2,BC_1F_1,BC_1F_2 produced were analysed during 1978～1992. The following conclusions were obtained.The male-sterility of A_1 and A_2 cytoplasm was both controlled by two nuclear genes.The genotype of A_1 cytoplasm male sterility is S_1N_2(ms_1ms_1ms_1~'ms_1~'ms_2ms_2ms_2~'ms_2') ,and the maintainer line is N_1N_2 (ms_1ms_1ms_1~'ms_1~'ms_2ms_2ms_2~'ms_2~'). The genotype of A_2 cytoplasm male sterility is N_1S_2 (MS_1MS_1ms_2ms_2 ms_2~'ms_2~') and the maitainer is N_1N_2(MS_1MS_1ms_2ms_2ms_2~'ms_2~'). The nuclear genotype of restorer lines of A_1 and A_2 are of 6 types separatly. There are correspondence clealy in the sex expression genes and cytoplasm.