贮藏蒜苔中细胞内含物由衰退茎组织向顶端珠蒜的再分配

离体蒜苔贮藏过程中,顶端珠蒜的形成依赖干衰退茎苔的自我牺牲。由于并无叶片光合产物,也无根系吸收的矿质水分的参与,顶端珠蒜生长所需的物质全部来自衰退茎苔内含物的撤离(包括顶端珠蒜外面的筒状苞片)。最后茎苔中央髓部几乎成为空腔,周缘皮层则由残存的骨架所支撑。而在蒜苔顶端形成一丛鲜脆多汁的珠蒜。茎苔中主要内含物输出率如下:全 N 为80—92%,全 P 为80—90%,K 为70—75%,干物质(除去呼吸消耗)为70—80%,水分(不考虑最后胞壁的失水)为60—70%,撤离是相当彻底的。茎苔内全 N、全 P、K 和水分的输出与干物质的输出有着密切的线性关系(r=0.95-0.98),表明细胞内含物的再分配是单向的和协同的,这应是衰退营养组织中内含物完全撤退的特征。蒜苔上较组织内糖、非蛋白 N、P 和 K 的含量高于下段,周缘皮层中 K、氨基酸的含量高于中央髓部,表明茎苔细胞内含物的输出是由内向外,由下而上进行的,这种逆浓淡梯度的转移表明有主动机制的参与。

REDISTRIBUTION OF CELL CONTENTS FROM DETERIORATING TISSUES TO DEVELOPING APICAL CLOVES IN GARLIC SCAPE DURING STORAGE

In an excised garlic scape on long storage,formation of new apical cloves can take place at the expense of the withering stalk proper. The material supply needed for new growth is solely derived from the cell contents of the various deteriorating tissues of the stalk(including the terminal bract)without involvement of photosynthesis and mineral absorption.The medullary portion of the stalk eventually becomes evacuated,transforming itself into a hollow cavity,whereas the pe- ripheral portion shrinks into a f...