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Leaf area development, biomass production and yield of four spring barley varieties grown in a Mediterranean environment (southern Spain) in response to an early application of foliar sulphur or etephon have been studied. Both sulphur and etephon produced similar results compared to the control. Thus, whereas the maximum leaf area index on untreated plots was reached at the beginning of shooting, the growth of foliar area in the treated ones was extended until anthesis and its decline during maturity was similarly retarded. This led to a significant improvement in the biomass at anthesis (which was closely correlated with grain yield) as well as in the leaf area duration during grain filling. These effects were principally due to variations in the number of leaves per plant, caused by a higher tiller production in the treated plants and not by an increase in the number of leaves per tiller or by leaf size.
An application of sulphur or etephon at tillering increases grain yield by raising both the number of ears per plant and per plot, without modifying the number of grains per ear or 1000-grain weight. The similarity between the effects of sulphur and etephon may be due to the fact that sulphur absorbed by the leaves results in an increase in methyonine, the biological precursor or ethylene. The positive correlation between biomass at anthesis and number of ears over grain yield suggests that sink capacity and source activity are closely related and both simultaneously limited by the environment.  相似文献   
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