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Natural reforestation of European mountain landscapes raises major environmental and societal issues. With local stakeholders in the Pyrenees National Park area (France), we studied agricultural landscape colonisation by ash (Fraxinus excelsior) to enlighten its impacts on biodiversity and other landscape functions of importance for the valley socio-economics. The study comprised an integrated assessment of land-use and land-cover change (LUCC) since the 1950s, and a scenario analysis of alternative future policy. We combined knowledge and methods from landscape ecology, land change and agricultural sciences, and a set of coordinated field studies to capture interactions and feedback in the local landscape/land-use system. Our results elicited the hierarchically-nested relationships between social and ecological processes. Agricultural change played a preeminent role in the spatial and temporal patterns of LUCC. Landscape colonisation by ash at the parcel level of organisation was merely controlled by grassland management, and in fact depended on the farmer’s land management at the whole-farm level. LUCC patterns at the landscape level depended to a great extent on interactions between farm household behaviours and the spatial arrangement of landholdings within the landscape mosaic. Our results stressed the need to represent the local SES function at a fine scale to adequately capture scenarios of change in landscape functions. These findings orientated our modelling choices in the building an agent-based model for LUCC simulation (SMASH–Spatialized Multi-Agent System of landscape colonization by ASH). We discuss our method and results with reference to topical issues in interdisciplinary research into the sustainability of multifunctional landscapes.  相似文献   
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Genotypic differences in the translocation of temporarily stored 14C from the stem to the grains in spring wheat (Triticum aestivum L.)
In three field experiments with two spring wheat genotypes (Kolibri and breeding line 93117 ), changes in the total nonstructural carbohydrates (TNC) of the stem were observed after anthesis. Maximum values were measured in the third or fourth week following anthesis when stems contained 300 to 400 mg TNC. Thereafter TNC content declined up to maturity.
Flag leaves of individual shoots or all plants in micro-plots were labelled with 14C 5 days prior to anthesis, at anthesis or 5 days after anthesis to observe long term movements of assimilates during grain filling. After a chase period of two to three days, 60 to 80 % of total 14C recovered in the shoot was in the stem. From total l4C recovered two to three days after labelling, Kolibri had translocated 12.5 to 27.0 % into the grains by maturity whereas this portion was significantly higher for the breeding line 93117 (22.5 to 43.9 %). It was concluded that genotypes differ in the translocation of soluble carbohydrates from the stem to the grains. These differences were not related to parameters describing the 'source-sink' relationship, such as leaf area, grain number or grain size. However, the lower translocation rates of Kolibri coincided with a lower TNC concentration in the stem dry matter. This was due to a higher stem weight at anthesis, a longer period of stem elongation and a higher incorporation of assimilates into structural carbohydrates in non elongating stem parts after anthesis. It was therefore suggested that the accumulation of TNC in the stem and the remobilisation of these reserves for grain filling are determined partly by factors related to the carbohydrate metabolism in the stem.  相似文献   
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