Die Verjüngung gemischter Bergwälder—Praktische Konsequenzen aus 10jähriger Untersuchungsarbeit

Zusammenfassung Seit zehn Jahren betreibt der Lel-irstuhl für Waldbau and Forsternrichtung der Universit?t M?nchen intensive Verjüngungsstudien auf 25 Dauerversuchsfl?chen im Bergmischwald der ostbayerischen Kalkalpen. Am Bei-spiel der Aufnahmedaten yon fünf ausgew?hlten Versuchsfl?chen werden grundlegende Prinzipien des Ver-jüngungsprozesses im Bergmischwald aufgezeigt and daraus praktische Konsequznzen abgeleitet. Alle am Autbau der Altbest?nde des Bergmischwaldes beteiligten Baumarten bilden bislang trotz d utli-cher Waldschadenssvmptome reichlich Samen von hochwertiger Qualit?t aus. Damit aus diesem Verjüngungspotential Naturverjüngung entstchen kann, müssen die en tsprechenden überschirmungsverh?ltnisse lurch den Altbestand gegeben sein. Für die Dichte, Baumartenztusammenset zung and H?henentwicklung der Naturverjüngungspflanzen ist n?mlich die Uberschirrnung von ganz ent-scheidender Bedeutung. In einem sehr dichten Altbestand, in derv der überschirmungsgrad fiber 75% liegt , haben Verjüngungspflanzen Schwierigkeiten, sick zu etablieren. Eine Ieichte Auflichtung auf einen über-schirmungsgrad von 60 (der dann gegeben ist, wenn die rich unter dem Schirm entwickelnde Bodenvege cation Bedeckungsgrade zwischen 20 and 3C einstellt) erm?glicht den Naturverjüngungsflanzen ein überleben. Diese Uberschirmung kann durch einen schwachen Schirmhieb (Entnahme von etwa einem Vier-tel der Grundtl?che eines geschlossenen Bestandes) erreicht werden. Diese Eingriffsst?rke dürfte auch unter den Gesichtspunkten der Stabilit?it and des Zuwachsverlustes dill Altbestand zu vertreten sein. Vor einer st?rkeren Auflichtung des Altbestandes müsses genügend Naturverjüingungspflanztn ant Bo den vorhanden sein. Nur so is[ gew?hrleistet, daβ these einen Vorsprung vor der Bodenvegatation bekom Men. Die Pflanzung ist immer dann, wenn sich Schwierrigkeiten mit der Naturverjüngung ergeben, ein geeigne-tes Mittel, um einen neuen Bergmischwald zu begründen. Derzeit ist für das Entstehen tines neuen Bergmischwaldes - sei es durch Naturverjüngung odor lurch Pflanzung - Zaunschutz erforderlich. Summary In 1976, a research project entitled ’Natural Regeneration of Mixed Mountain Forests ’ was etablished in the Calcareous Alps of eastern Bavaria, by the Chair of Silviculture and Forest Management of the University Of Munich. Over the past ten years, the multi-variate aspects of regenaration have bean intensively investigated on 25 permanent research plots. The data from 5 selected plats were extracted and processed. from these data, it will be possible to present some fundamental principles of natural regenaration processes. From these prin-ciples, it is hoped that some practical applications can be derived. The mature stands of mixed mountain forests are still able to produce high quantities and good quality of seed, although distinct symptoms of forest decline are present. In order to utilize this potential for natural regeneration, an adequate canopy density is necessary. Canopy density is the crucial factor which regulates the density, species composition and height development of regeneration. In a very heavily, stocked mature stand with a canopy density of over 75 establishment of regenerations is difficult. When canopy density is reduced to, for example, 60% (this percentage can be estimated from a ground vegetation covering approximately 20–30 of the soil), the naturally regenerated, plants have a better opportunity to survive. A 60% canopy density can be achieved by light shelterwood cutting (removal of approximately 25% of basal area of a closed stand). This type of cutting can maintain the stability and prevent increment loss of forest stands. It is important that regeneration is established before opening up the stand, only, then these plants are capable to compete with the other ground vegetation. Planting is always appropriate when natural regeneration fails. Fencing is necessary for undisturbed development of natural and artificial regeneration.

---

---

Dieses Heft enth?lt waldbauliche Beitr?g zu den Forschungsschwerpunkten Bergwald, Waldschaden und Kiefer. Sie sind überwiegen als Vortr?ge auf der Hochschulwoche der Forstwissenschaftlichen Fakult?t München im Oktober 1987 gehalten worden. Ihre Autoren widmen sie Herrn Professor Dr. Peter Burschel zum 60. Geburtstag.

---

Auf der Forstlichen Hochschulwoche 1987 in München gehaltener Vortrag.     

中欧地区的森林培育及其启示

文中简述了中欧的林业发展史，特别对德国森林培育的现状和存在的问题进行了比较详细的分析。根据德国森林培育的历史和经验，提出了对中国林业建设的几点启示。     

Induction of de novo volatile terpene biosynthesis via cytosolic and plastidial pathways by methyl jasmonate in foliage of Vitis vinifera L

The terpene biosynthesis in leaves of Vitis vinifera L. cv. Morio Muskat was studied using methyl jasmonate to induce defensive responses in vivo. The experiments demonstrated the strong activation of the de novo biosynthesis of terpenoids via the octadecanoid-signaling cascade and release of the compounds to the gas phase. Feeding experiments with [5,5-(2)H(2)]-1-deoxy-d-xylulose and [5,5-(2)H(2)]mevalonic acid lactone allowed the investigation of the dynamic allocation of resources via the mevalonic acid and 1-deoxy-d-xylulose/2-C-methyl-d-erythritol 4-phosphate (DOXP/MEP) pathway under induced conditions and after treatment with the specific inhibitors mevastatin and fosmidomycin. The experiments reveal that monoterpenes are almost exclusively synthesized via the DOXP/MEP pathway, whereas sesquiterpenes are generated via both pathways at approximately equal rates. The biosynthesis of the homoterpene (E)-4,8-dimethyl-1,3,7-nonatriene was not affected by mevastatine or fosmidomycin.     

Conversion of citronellyl diphosphate and citronellyl beta-D-glucoside into rose oxide by Pelargonium graveolens

Deuterium-labeled citronellyl diphosphate and citronellyl beta-D-glucoside were fed to Pelargonium graveolens. Both precursors were converted into cis-/trans-rose oxide. Citronellyl diphosphate is more effectively converted into rose oxide. The phloem exudate of P. graveolens was analyzed after enzymatic cleavage of the glucosidated and phosphorylated volatiles. It could be shown that glucosidically bound citronellol is translocated in the plant. Phosphorylated citronellol could not be detected.     

Biogenetic studies in Syringa vulgaris L.: synthesis and bioconversion of deuterium-labeled precursors into lilac aldehydes and lilac alcohols

Syringa vulgaris L. inflorescences were fed with aqueous solutions of regioselectively deuterated compounds assumed to be precursors of lilac aldehyde and lilac alcohol, respectively. Volatiles were extracted by stir bar sorptive extraction (SBSE) and analyzed using enantioselective multidimensional gas chromatography/mass spectrometry (enantio-MDGC/MS); deuterium-labeled lilac aldehydes and lilac alcohols were separated from unlabeled stereoisomers on a fused silica capillary column, coated with heptakis(2,3-di-O-methyl-6-O-tert-butyldimethylsilyl)-beta-cyclodextrin (DIME-beta-CD) (30%) in SE 52 (70%), as the chiral stationary phase. Feeding experiments with [5,5-(2)H(2)]mevalonic acid lactone 22 and [5,5-(2)H(2)]deoxy-d-xylose 23 indicate that the novel mevalonate independent 1-deoxy-d-xylose 5-phosphate/2C-methyl-d-erythritol 4-phosphate pathway is the dominant metabolic route for biosynthesis in lilac flowers. Additionally, bioconversion of deuterium-labeled d(5)-(R/S)-linalool 3, d(6)-(R)-linalool 21, d(5)-(R/S)-8-hydroxylinalool 6, d(5)-(R/S)-8-oxolinalool 7, d(5)-lilac aldehydes 8-11 and d(5)-lilac alcohols 12-15 into lilac during in vivo feeding experiments was investigated and the metabolic pathway is discussed. Incubation of petals with an aqueous solution of deuterated d(5)-(R/S)-linalool 3 indicates an autonomic terpene biosynthesis of lilac flavor compounds in the flower petals of lilac.     

Comprehensive authentication of (E)-alpha(beta)-ionone from raspberries, using constant flow MDGC-C/P-IRMS and enantio-MDGC-MS

A new coupling system of GC-GC, connected via a Multi Column Switching Device MCS2 for measuring isotope ratios, is introduced. By means of several standard substances the precise and accurate measurement of isotopic values is proved. First applications concerning the authentication of raspberry aroma compounds are established. Consequently, the combination of constant flow multidimensional gas chromatography-combustion/pyrolysis-isotope ratio mass spectrometry (MDGC-C/P-IRMS) is applied to the authenticity assessment of (E)-alpha(beta)-ionone from six different raspberry cultivars. Furthermore, 12 commercially available raspberry products and samples of (E)-alpha(beta)-ionone, some declared to be natural, are investigated. delta(2)Eta(V)(-)(SMOW) and delta(13)C(V)(-)(PDB) values of (E)-alpha(beta)-ionone are determined, and characteristic authenticity ranges were concluded from raspberries by correlation of both delta(2)Eta(V)(-)(SMOW) and delta(13)C( V)(-)(PDB) values. The results are correlated with the determination of enantiomeric purities of (E)-alpha-ionone, using stir bar sorptive extraction enantio-multidimensional gas chromatography mass spectrometry (SBSE-enantio-MDGC-MS).     

Mechanism of wine lactone formation: demonstration of stereoselective cyclization and 1,3-hydride shift

The cyclization mechanism of (E)-2,6-dimethyl-6-hydroxyocta-2,7-dienoic acid to wine lactone under acidic aqueous conditions was investigated using the two stereoselectively deuterium-labeled precursors (2E,6R,7Z)-[8-2H]-2,6-dimethyl-6-hydroxyocta-2,7-dienoic acid and (2E,7E)-(+/-)-[8-2H]-2,6-dimethyl-6-hydroxyocta-2,7-dienoic acid. A detailed analysis of the generated wine lactone isomers by enantioselective multidimensional gas chromatography (MDGC)/ion trap tandem mass spectrometry demonstrates that the formation of wine lactone proceeds via a nonenzymatic stereoselective cationic cyclization cascade that includes a 1,3-hydride shift. Usually, such mechanisms are features of cyclization reactions that are catalyzed by terpene cyclases. This nonenzymatic conversion of an acyclic precursor to a bicyclic monoterpene under relevant cationic cyclization conditions has rarely been observed and confirms recent suggestions that the precursor itself can provide the chemical functionality required for specific steps in the cyclization cascade.     

Buchbesprechungen

Ohne Zusammenfassung     

Biosynthesis of monoterpenes and norisoprenoids in raspberry fruits (Rubus idaeus L.): the role of cytosolic mevalonate and plastidial methylerythritol phosphate pathway

The biosynthesis of the monoterpenes (-)-alpha-pinene, linalool, and the norisoprenoids alpha- and beta-ionone in raspberry fruits (rubus idaeus L.) was investigated by in vivo feeding experiments with [5,5-(2)H2]-mevalonic acid lactone and [5,5-(2)H2]-1-deoxy-D-xylulose. The volatile compounds were extracted by stirbar sorptive extraction and analyzed using thermal desorption-multidimensional gas chromatography-mass spectrometry (TD-enantio-MDGC-MS). The feeding experiments demonstrate that (-)-alpha-pinene and (S)-linalool are exclusively synthesized via the cytosolic mevalonic acid pathway. In contrast, (2)H-labeled (R)-(E)-alpha-ionone and (2)H-labeled (E)-beta-ionone are detectable after application of d2-1-deoxy-D-xylulose and d2-mevalonic acid lactone, respectively. However, (R)-linalool reveals no incorporation of either one of the fed precursors, even though this enantiomer is detectable in the fruit tissue.