Growth and yield models for uneven-sized forest stands in Finland

Uneven-sized forestry is gradually gaining popularity and acceptability also in the Nordic countries. This is because of the willingness of the public and some forest owners to avoid clear-fellings and pursue more near-nature forest structures. It has also been realized that the profitability of uneven-sized forestry may be competitive with even-aged forestry. In Finland, management of uneven-sized stands is hampered by the lack of information about the dynamics of such stands, and about the yield and profitability of uneven-sized forestry. This study developed models which allow managers to simulate the growth and yield of uneven-sized stands in Finland, making it possible to predict the yield and analyze the sustainability of different management options. The model set consists of individual-tree diameter increment, height and survival models, and a model for ingrowth. The modeling data consisted of two long-term field experiments of uneven-sized forest management, a set of temporary sample plots measured earlier for growth modeling purposes, and the sample plots of the third National Forest Inventory of Finland. The application area of the models covers all growing sites, all main tree species, and the whole surface area of Finland. According to the models, the sustainable harvest of a fertile (Oxalis-Myrtillus site) uneven-sized Norway spruce forest varies between 5.5 and 7 m³ ha⁻¹ a⁻¹ in Central Finland, depending on the length of the cutting cycle, stand density, and shape of the diameter distribution. It is profitable to harvest large diameter classes more heavily than small ones. Due to the large amount of data the models for diameter increment are highly significant and reliably show the growth level of trees in uneven-sized stands. The weakest models are the ingrowth models, which are based on a clearly smaller data set than the other models.     

Effects of three different concentrate proportions and rapeseed meal supplement to grass silage on animal performance of dairy-breed bulls with TMR feeding

A 3 × 2 factorial design with growing dairy-breed bulls was used to study the effects on animal performance of (1) proportion of concentrate (rolled barley) in the diet, and (2) inclusion of rapeseed meal (RSM) in the barley-based concentrate in a total mixed ration (TMR). The interactions between concentrate proportion and RSM supplement were also examined.Three feeding experiments comprised in total of 84 Finnish Ayrshire bulls and 6 Friesian bulls. The bulls were fed TMR ad libitum. The three concentrate proportions were 300 (L), 500 (M) and 700 (H) g/kg dry matter (DM), fed without RSM (RSM−) or with RSM (RSM+). Rapeseed meal was given so that the crude protein (CP) content of the concentrate was raised to 160 g/kg DM in the RSM+ diets. In the RSM− diets the CP content of the concentrate was 128 g/kg DM, so the CP content increased 25% with RSM supplementation. Increasing the proportion of concentrate led to a linear improvement in daily live weight gain (LWG) (P < 0.05), but there were no significant treatment differences in the DM intake (kg/d). Increasing the proportion of concentrate also led to significantly higher CP (P < 0.001) and phosphorus (P) (P < 0.001) supply and significantly improved DM and organic matter (OM) digestibility (P < 0.001). However, the digestibility of neutral detergent fibre (NDF) decreased (P < 0.001) as the proportion of concentrate increased. The feed conversion rate (kg DM/kg LWG) decreased significantly with increasing concentrate proportion (P < 0.001). Rapeseed meal supplement had no effect on animal performance, but the supply of CP (P < 0.01) and P (P < 0.001) was higher when RSM was included in the diet. The CP (P < 0.001) and NDF (P < 0.05) digestibilities were also higher for the RSM+ diets than for the RSM− diets. Because RSM at the concentration used did not affect animal performance, there is no reason to use RSM supplementation for finishing dairy bulls when there is good quality grass silage and barley-based concentrate in the TMR ration. This study also shows that there is a need to update the Finnish feeding recommendations for dairy-breed growing bulls, and extra calculations are needed for the energy and protein supply of growing dairy bulls.     

Forest management inspired by natural disturbance dynamics (DISTDYN) – a long-term research and development project in Finland

A large-scale and long-term research and development project Forest management inspired by natural disturbance dynamics (DISTDYN) was initiated in Finland in 2009. The aim is to examine how variation in spatial and temporal scales of felling affects forest structure and biota. Harvesting methods applied in this project include selection, gap, partial and clear felling with variable retention. Rehabilitation is also applied as dead wood is intentionally increased while harvesting. The motivation of the project lies in the growing interest in using natural disturbances as an inspiration of forest-ecosystem management and in the recognition that mid- to small-scale disturbances are a characteristic feature of unmanaged Fennoscandian forests. The effects of felling methods are studied at the stand scale. However, a unique aspect of the project is the focus on landscape-scale impacts of different harvesting alternatives by comparing ca. 150-ha landscape units with varying harvesting regimes. Overall, the project aims at creating a long-lasting multidisciplinary research infrastructure. It also addresses core issues of forest-ecosystem management, including ecological, timber-production, economic and socio-cultural performances of alternative harvesting practices.     

Optimal management of uneven-aged Norway spruce stands

An optimization model is specified to analyze forest management without any restrictions on the forest management system. The data on forest growth comes from unique field experiments and is used to estimate a nonlinear transition matrix or size-structured model for Norway spruce. The objective function includes detailed harvesting cost specifications and the optimization problem is solved in its most general dynamic form. In optimal uneven-aged management, stand density is shown to be dominated by limitations in natural regeneration. If the goal is volume maximization, even-aged management with artificial regeneration (and thinnings from above) is superior to uneven-aged management. After including regeneration and harvesting costs, the interest rate, and the price differential between saw timber and pulpwood, uneven-aged management becomes superior to even-aged management. However, in the short term the superiority is conditional on the initial stand state.     

Biodiversity policies in commercial boreal forests: Optimal design of subsidy and tax combinations

This paper examines the first-best instruments for biodiversity maintenance in commercial boreal forests when landowners behave either in Faustmannian or Hartmanian way. Using an extended Hartman model, we show that biodiversity conservation requires both prolonged rotation age and leaving retention trees. While the former promotes some old growth species, the latter create new structural elements of decaying and dead wood, which can sustain a variety of saprolyxic species. A fully synchronized combination of retention tree subsidy and tax instrument is needed both to lengthen the privately optimal rotation period and to provide an incentive to leave retention trees. Using Finnish data we illustrate empirically the sizes of instruments. When combined with a harvest tax, the retention tree subsidy is 1000 and 750 € in the Faustmann and in the Hartman model, respectively. When used with a timber subsidy or a site value tax, the retention tree subsidy is 1700 €/ha in both models. The harvest tax rate varies over the range 40–65% in the Faustmann model and 20–40% in the Hartman model, while timber subsidy is between 0.5–1.0% and site value tax is about 1.75%.     

Species Interactions in the Dynamics of Even- and Uneven-Aged Boreal Forests

Many boreal tree stands are neither clearly even-aged nor clearly uneven-aged. The stands may undergo a series of stages, during which an even-aged stand is transformed into two-storied mixed stand, and finally to multistoried or uneven-aged stand structure. The species composition often changes during the succession of stand stages. This study developed models for stand dynamics that can be used in different stand structures and species compositions. The model set consists of species-specific individual-tree diameter increment and survival models, and models for ingrowth. Separate models were developed for Scots pine, Norway spruce, and hardwood species. The models were used in a growth simulator, to give illustrative examples on species influences and stand dynamics. Methods to simulate residual variation around diameter increment and ingrowth models are also presented. The results suggest that mixed stands are more productive than one-species stands. Spruce in particular benefits from an admixture of other species. Mixed species improve diameter increment, decrease mortality, and increase ingrowth. Pine is a more beneficial admixture than birch. Simulations showed that uneven-aged management of spruce forests is sustainable and productive, and even-aged conifer stands growing on medium sites can be converted into uneven-aged mixed stands by a series of strong high thinnings.     

Crystallinity of wood and the size of cellulose crystallites in Norway spruce (<Emphasis Type="Italic">Picea abies</Emphasis>)

X-ray diffraction was used to study variations in the crystallinity of wood and the average thickness and length of the crystallites of cellulose as a function of the number of the year ring in Norway spruce [Picea abies (L.) Karst.]. The crystallinity increased from ring 4 to ring 10 from the pith and was constant after ring 10. The crystallinity of mature wood was about 30% ± 5%. The average thickness and average length of the crystallites were 3.2 ± 0.1nm and 28 ± 2nm, respectively; and no systematic variation of these values with the number of the year ring was observed. The mean microfibril angle decreased near the pith but was constant in the mature wood.     

Optimizing any-aged management of mixed boreal forest under residual basal area constraints

The current trend of forest management in many countries is reduced use of clear-felling and planting, and increased use of continuous cover management. In Finland, the new forest act of 2014 made all types of cuttings equally allowable on the condition that if the post-cutting residual stand basal area is too low, the stand must be regenerated within certain time frame. Forest landowner can freely choose between evenand uneven-aged management. This study developed a method for optimizing the timing and type of cuttings without the need to categorize the management system as either even-aged or uneven-aged. A management system that does not set any requirements on the sequence of post-cutting diameter distributions is called any-aged management. Planting or sowing was used when stand basal area fell below the required minimum basal area and the amount of advance regeneration was less than required in the regulations. When the cuttings of 200 stands managed earlier with even-aged silviculture were optimized with the developed system, final felling followed by artificial regeneration was selected for almost 50% of stands. Reduction of the minimum basal area limit greatly decreased the use of artificial regeneration but improved profitability, suggesting that the truly optimal management would be to use natural regeneration in financially mature stands. The optimal type of thinning was high thinning in 97–99 % of cases. It was calculated that the minimum basal area requirement reduced the mean net present value of the stands by 12–16 % when discount rate was 3–5 %.     

Survival of Atlantic salmon captured in and released from a commercial trap-net: Potential for selective harvesting of stocked salmon

The major wild Atlantic salmon stocks in the Baltic Sea began to recover in the late 1990s. This recovery has been partly due to strict regulations in the Gulf of Bothnia that effectively prevent salmon fisheries during the peak migration. About half of the migrating salmon, however, are reared fish that could be harvested. We simulated a limited trap-net fishery that selectively harvested reared salmon and released wild fish, and studied the survival and migration of the released salmon. We tagged and released 1970 salmon caught in the trap-nets along the coast in 2001 and 2002. The mean maximum capture and release induced mortality of salmon was 11%, ranging between 4% and 21% in different release groups by year, sea age and number of releases. The cumulative mortality for the total salmon population on their spawning migration in the Gulf of Bothnia was below 5%, and it would not increase considerably after the first capture and release events, provided fishing effort is not excessive and fish are handled properly. Survival of trap-net captured and released Baltic salmon appears high and their migration behavior is not altered due to this handling. Several preconditions, however, should be considered before selective fishing is introduced in the Gulf of Bothnia salmon fishery.     

Acceleration of N mineralization by release of enzymes and substrates from soil mineral particles with phosphates

A study was conducted to develop an improved method for measuring organic N (net) mineralization in which chemical extraction takes place in combination with suspension incubation in ammonia-absorbing membrane bottles. To obtain direct evidence of the extent to which extracted organic N is mineralizable, the extraction suspension was further incubated immediately after the extraction procedure with mild and selective extractants. In this ‘extraction incubation’ method, extraction continues during the incubation but only relatively easily mineralizable organic matter is released. Standard incubation is usually carried out in sealed N₂-flushed bottles. However, when phosphate or pyrophosphate soil suspensions are incubated, mineralization is much higher than in soil water suspensions. Further, accumulation of ammonia+(ammonium) and other gases, i.e. CO_2, can affect the reaction rate and final reaction equilibrium in the sealed incubation flask. It was to avoid these effects that the membrane method was developed. With this procedure, the flask is closed with an ammonia-absorbing membrane permeable to other gases. Water, phosphate and pyrophosphate suspensions were incubated at 37 °C in sealed bottles (SB), in sealed N₂ gas-flushed bottles (SBN₂), and in bottles with ammonia-trapping filters (MB). The maximum amount of released during 10 days' incubation was 133.0 mg kg⁻¹ in the water, 208.0 mg kg⁻¹ in the phosphate and 454.1 mg kg⁻¹ in the pyrophosphate suspension (soil total C content 6.2% and N 0.25%). During incubation in phosphate and pyrophosphate suspensions, the mobilization was nearly linear in membrane bottles. The variation between replicates was also smallest in these bottles. It was concluded that membrane bottles were best suited to incubation when mobilization reactions were accelerated with phosphate or pyrophosphate extractants. The method was easy to perform and gave results with good replicability.