基于地统计方法的山西省森林木本层生物量及碳密度空间分布格局研究

笔者利用经典统计和地统计分析方法,研究了山西省森林木本层生物量和碳密度的变异性及空间分布格局。结果表明,山西省森林乔木层和灌木层生物量平均值分别为60.26t/hm^2和3.22t/hm^2,木本层的碳密度平均值为31.42t/hm^2.灌木层生物量为强变异,乔木层生物量、木本层地上生物量和碳密度均为中等变异水平。乔木层生物量、灌木层生物量、地上生物量的最优半变异函数模型均为指数模型,碳密度为圆形模型;普通克里格差值的空间预测模型拟合效果较好。     

The contribution of competition to tree mortality in old-growth coniferous forests

Competition is a well-documented contributor to tree mortality in temperate forests, with numerous studies documenting a relationship between tree death and the competitive environment. Models frequently rely on competition as the only non-random mechanism affecting tree mortality. However, for mature forests, competition may cease to be the primary driver of mortality.We use a large, long-term dataset to study the importance of competition in determining tree mortality in old-growth forests on the western slope of the Sierra Nevada of California, U.S.A. We make use of the comparative spatial configuration of dead and live trees, changes in tree spatial pattern through time, and field assessments of contributors to an individual tree's death to quantify competitive effects.Competition was apparently a significant contributor to tree mortality in these forests. Trees that died tended to be in more competitive environments than trees that survived, and suppression frequently appeared as a factor contributing to mortality. On the other hand, based on spatial pattern analyses, only three of 14 plots demonstrated compelling evidence that competition was dominating mortality. Most of the rest of the plots fell within the expectation for random mortality, and three fit neither the random nor the competition model. These results suggest that while competition is often playing a significant role in tree mortality processes in these forests it only infrequently governs those processes. In addition, the field assessments indicated a substantial presence of biotic mortality agents in trees that died.While competition is almost certainly important, demographics in these forests cannot accurately be characterized without a better grasp of other mortality processes. In particular, we likely need a better understanding of biotic agents and their interactions with one another and with competition.     

Seasonal patterns of photochemical capacity and spring phenology reveal genetic differentiation among native Scots pine (Pinus sylvestris L.) populations in Scotland

Environment-driven genetic differentiation among populations is a common feature among forest trees, and an understanding of how populations have adapted to their home site conditions is essential for management and conservation practices. In Scotland, 84 native Scots pine (Pinus sylvestris L.) woodlands are recognised by the Forestry Commission and they occupy highly diverse environments from the maritime west coast to continental sites in eastern Scotland. However, it is not known whether adaptations to local environments along sharp temperature and rainfall gradients have occurred in different populations and as a result, the seed transfer guidelines of the species are based only on data from isozymes and monoterpenes. In this study of an outdoor common-garden trial, we used chlorophyll fluorescence to examine whether seedlings from 32 open-pollinated families and eight populations from sites experiencing contrasting annual temperature regimes differed in their response to variation in natural outdoor temperatures between September 2009 and May 2010. In addition, growth initiation in spring was recorded. Photochemical capacity at photosystem II F_v/F_m showed a distinct seasonal trend and remained at relatively high levels (∼0.7) until November. Following a period of over 2 weeks with temperatures below or close to 0 °C, F_v/F_m started decreasing towards its minimum values recorded in early March when population means varied between 0.35 and 0.45. By early May and along with rising temperatures, photochemical capacity had recovered to the same level as observed in early November. Populations were found to respond differently to the cold period starting in December. The largest drop in photochemical capacity was observed in seedlings from a low-altitude population located in the maritime western Scotland, while in seedlings from higher-altitude locations in the cooler eastern Scotland, the response was smaller. In March, the recovery of photochemical capacity was slowest in seedlings from the mildest and coolest sites. Evidence of adaptive genetic differentiation was also found in spring phenology. Initiation of shoot elongation and needle flush were earlier in families from higher altitudes (cooler areas), but population differences were not significant at the α = 0.05 level. These results suggest that adaptation to the spatially heterogeneous environment in Scotland has taken place in Scots pine and that in order to minimise the risk of planting maladapted seed stock, the patterns of environmental and adaptive genetic variation should be taken into account in the management of genetic resources in this species.     

Analysis of spatio-temporal changes in forest biomass in China

Forests play a central role in the global carbon cycle.China's forests have a high carbon sequestration potential owing to their wide distribution,young age and relatively low carbon density.Forest biomass is an essential variable for assessing carbon sequestration capacity,thus determining the spatio-temporal changes of forest biomass is critical to the national carbon budget and to contribute to sustainable forest management.Based on Chinese for-est inventory data (1999-2013),this study explored spatial patterns of forest biomass at a grid resolution of 1 km by applying a downscaling method and further analyzed spatio-temporal changes of biomass at different spatial scales.The main findings are:(1) the regression relationship between forest biomass and the associated influencing factors at a provincial scale can be applied to estimate biomass at a pixel scale by employing a downscaling method;(2) for-est biomass had a distinct spatial pattern with the greatest biomass occurring in the major mountain ranges;(3) forest biomass changes had a notable spatial distribution pattern;increase (i.e.,carbon sinks) occurred in east and southeast China,decreases (i.e.,carbon sources) were observed in the northeast to southwest,with the largest biomass losses in the Hengduan Mountains,Southern Hainan and Northern Da Hinggan Mountains;and,(4) forest vegetation functioned as a carbon sink during 1999-2013 with a net increase in biomass of 3.71 Pg.     

A mathematical model for generating land-use allocation alternatives for agroforestry systems

Agroforestry has been considered a viable land-use system particularly in the tropical regions. In developing or designing agroforestry systems, various concerns have to be addressed namely: (1) economic; (2) biological and physical; and (3) silvicultural and ecological considerations. This paper addresses these concerns from an interdisciplinary perspective. A land-use allocation model which ensures the simultaneous consideration of these concerns is proposed.     

扁刺蛾越冬幼虫空间分布型的研究

通过测定５个空间分布型指数和建立２个回归直线,发现扁刺蛾越冬幼虫空间分布型属聚集分布。另外,干基周围划分为３个层次,内层虫口数量与中、外两层的差异均达１％的显著水平。     

Using the Spectacled Bear as a Conservation Tool in the Condor Bioreserve,Ecuador

Abstract

Conservation organizations in the Condor Bioreserve (CBR) in northern Ecuador use the spectacled bear (Tremarctos ornatus) as a surrogate species to focus conservation efforts. However, the utility of single-species management has been questioned in recent scientific literature, making it appropriate to review the spectacled bear's efficacy as a conservation tool. The spectacled bear does not enjoy unqualified public support in CBR and surroundings, suggesting that it may not be appropriate as a flagship species. It may be more effective as a keystone species; its diet and migratory habits suggest that it may have a significant impact on plant community structure through fruit dispersal. However, more research is needed in this arena before drawing any conclusions. The spectacled bear is perhaps most appropriate as an umbrella species because of its large home range, which encompasses at least two ecosystem types: páramo and montane forest. Traditionally, the concept of the umbrella species has stressed minimum area requirements, but the spectacled bear may be a more effective umbrella species if its dispersal requirements and seasonal migration patterns are also addressed. Eco-Ciencia, an Ecuadorian research organization, recently constructed a habitat model for spectacled bear. They are uniquely positioned to address issues of habitat connectivity both within and between the protected areas of CBR. In doing so, they could influence future zoning decisions in and around CBR, an area that is experiencing rapid population growth. This could improve conservation efforts for spectacled bear and other species.     

Fine-scale species co-occurrence patterns in an old-growth temperate forest

The pattern of species co-occurrence is instrumental for understanding community assembly rules. In this paper, we analyzed the co-occurrence of tree species in a 25-ha old-growth temperate forest plot in Northeastern China. The analysis was conducted at seven scales from 5 m × 5 m up to 100 m × 100 m in order to determine the patterns of co-occurrence at different spatial scales. Our analyses were conducted for all species, species with larger abundances, species with larger sizes, and five phylogenetic-based species groups. Our results showed that at smaller scales, the co-occurrence patterns of all species, species with larger abundances, and species with larger sizes were significantly higher than expected by chance, suggesting that strong interspecies competition exists in the community. At larger scales, there was no significant difference compared to randomized matrices. The result indicated that plant assembly rules are only found at small spatial scales. However, when co-occurrence metrics were restricted to phylogenetic groups, we could not find any clear evidence of interspecific competition within these groups. In conclusion, we found that competition is an important assembly rule at small scales in governing tree communities of our temperate forest, although it is not the only process involved. The importance of other processes should also be taken into account to explain species co-occurrence patterns.     

Indexing of spatial patterns of trees using a mean of angles

The assessment of information relating to the spatial patterns of individual trees is becoming an increasingly important aspect of forest research and practical forestry. Numerous methods have been proposed over the years, with foresters preferring rapid methods able to provide statistically confident information. The Mean of Angles is one of the simplest of these methods. It is applicable for revealing complete spatial randomness (CSR) or the presence of clustering or regularity. We have tested the potential of the arithmetic mean of angles to serve as a practical measure of the degree of regularity or clustering. This study was conducted by applying random point sampling in various simulated theoretical point populations. The Mean of Angles method deserves consideration as a practical option for examining the spatial patterns of trees in forests. We suggest that the arithmetic mean of angles cannot be completely relied upon for indexing spatial patterns, but this deficiency can be overcome by examining the frequency distributions of angular measurements.     

Assessing changes in forest fragmentation following infestation using time series Landsat imagery

The current epidemic of mountain pine beetle (Dendroctonus ponderosae Hopkins) in British Columbia, Canada, has impacted an area of over 13 million hectares presenting a considerable challenge to provincial forest resource managers. Remote sensing technologies offer a highly effective tool to monitor this impact due to very large areas involved and its ability to detect dead and dying tree crowns. Conventionally, change detection procedures based upon spectral values have been applied; however, analysis of landscape pattern changes associated with long-time series change detection approaches present opportunities for the generation of unique and ecologically important information. This study is focussed on the detection and monitoring of the shape and area characteristics of lodgepole pine stands during mountain pine beetle infestation to quantify the progression of forest fragmentation and related loss of landscape connectivity. A set of landscape pattern indices were applied to a set of images consisting of six Landsat satellite images spanning the period from 1993 to 2006. Our results indicate that the impacts of the mountain pine beetle infestation on forest spatial pattern consist of an increase in the number of patches, an increase in forest patch shape complexity, a reduction in forest patch size, an increase in forest patch isolation, and a decrease in interspersion. These findings demonstrate the unique information available from long-time series satellite imagery combined with pattern analysis to better understand the combined effects of insect infestation and forest salvage and harvesting.