Patterns of Nitrogen Excretion by Embryonic Softshell Turtles (Trionyx spiniferus) Developing in Cleidoic Eggs

Embryonic softshell turtles develop inside eggs exchanging little, if any, liquid water with the nest environment. Despite the consequent restriction on availability of water to support development, embryos convert most of the ammonia released in catabolism of proteins into soluble urea rather than insoluble urate, and thereby commit a portion of their limited reserve of water to serve as solvent for metabolic wastes. This finding is contrary to expectations from prior studies of cleidoic eggs.