Weed biomass dynamics in planted fallow systems in the humid forest zone of southern Cameroon

Interest in planted fallow systems has focused on soil fertility improvement, neglecting other potential benefits of such systems. It is important to quantify other processes responsible for crop yield increases under planted fallows, such as weed control. The suppressive potential on weeds of Flemingia macrophylla [(Willd.) Merrill] and Pueraria phaseoloides (Roxb.) Benth, planted fallows was evaluated in field trials in three villages in southern Cameroon. In each village, experiments were set up in 4–5 year-old bush fallow dominated by Chromolaena odorata (L.) R. M. King & H. Rob. and 20 year-old secondary forest. Total aboveground biomass production of P. phaseoloides was 7.45 Mg ha⁻¹, 4.2 times higher than F. macrophylla (1.78 Mg ha⁻¹ ; P < 0.05). The high biomass of P. phaseoloides resulted in a significantly greater reduction in total weed biomass compared to Flemingia macrophylla in both wet and dry seasons. In the wet season (11 and 18 MAP), there were significant fallow system × land use and fallow system × village interactions for total weeds and broadleaf weeds. P. phaseoloides in bush (0.55 Mg ha⁻¹), and P. phaseoloides at Ngoumou (0.09 Mg ha⁻¹) had the lowest total weeds in the wet seasons. After the dry season, the lowest total weed mass was consistently recorded in P. phaseoloides while the highest was in the natural regrowth. The population of grasses was always higher in the F. macrophylla system than in P. phaseoloides system throughout the wet and dry seasons. Grass biomass in the P. phaseoloides-forest LUS was the least (0.01 Mg ha⁻¹), 58 times lower than in F. macrophylla-bush (0.58 Mg ha⁻¹). Biomass production of P. phaseoloides was highly significantly correlated with total weed biomass (r = −0.64; P = 0.004) while no relationship was found between biomass production of F. macrophylla and total weed biomass (r = −0.08, P = 0.747). It was concluded that P. phaseoloides was a suitable leguminous species for weed control. But for F. macrophylla, its low biomass production coupled with a compact plant architecture compromised it as an appropriate species for weed control in a planted fallow system.     

An anti-inflammatory and immunomodulatory polysaccharide from Orbignya phalerata

A polysaccharide, a glucan with mean M_r of 1.0×10⁶ (MP1), was isolated from the mesocarp of fruits of Orbignya phalerata. Chemical and spectroscopic studies indicated that MP1 has a highly branched glucan type structure composed of α-(1→4) linked -glucopyranose residues with (3→4), (4→6), and with (3→6) branching points. MP1 enhanced phagocytosis in vivo and exhibited anti-inflammatory activity.     

Influence of remnant trees on nutrients and fallow biomass in slash and burn agroecosystems in Guinea

Three tree species are traditionally conserved in the traditional slash and burn agricultural system practiced for the production of upland rice (Oryza sativa Linn.) in the Fouta Djallon region of Guinea, i.e. (Parinari excelsa [Sabine], Parkia biglobosa [Jacq.) Benth.], Erythrophleum guinensis [G. Don.]). Sampling a chronosequence of fallow sites indicated that extractable P; exchangeable K, Ca and Mg; cation exchange capacity (CEC) and pH all decreased over the fallow period with patterns that differed with soil depth. Soils under the remnant Parinari excelsa and Parkia biglobosa trees had higher concentrations of organic C; total N; extractable P; exchangeable K, Ca and Mg; total P and Ca; and CEC than did the open microsites. Extractable P; exchangeable K, Ca and Mg; total P and Ca; and CEC were greater under Parinari excelsa than under Erythrophleum guinensis. Intensive measurements of a single mature fallow site showed that the foliar nutrient concentration of the large trees was not an accurate index of which microsite had the greatest accumulation of biomass and nutrients in the aboveground fallow vegetation. The biomass and the total amounts of N, P, K, Ca and Mg in the fallow vegetation were significantly greater for the Parinari excelsa and Parkia biglobosa microsites than for the open microsites. We conclude that although the nutrient status of the soils decreased over the 8-year fallow period, the microsites under the large trees were generally more fertile than the open microsites. However, there were important differences in the effects of the different tree species. This revised version was published online in June 2006 with corrections to the Cover Date.     

Volunteer biomass production between multipurpose tree hedgerows after two years of fallow in southern Cameroon

The ability of multipurpose hedgerow tree species to out-compete undesired regrowth during fallow phases was examined. Biomass and spatial distribution of grass and broad leaf volunteers was measured after two years of fallow, in two alley cropping systems planted at six m interrow distance, at the Humid Forest Ecoregional Centre Research Station, Mbalmayo, southern Cameroon. The two experiments had been continuously cropped for five and six years previously. In the experiment cropped for six years, the presence of Senna spectabilis [(DC.) Irwin and Barnaby] hedgerows reduced the biomass of the volunteer regrowth from 9.2 to 4.3 Mg ha⁻¹. Tillage during the previous cropping phase increased the broad leaf biomass from 3.0 to 4.4 Mg ha⁻¹, reduced the biomass of grasses from 3.4 to 2.7 Mg ha⁻¹ but had no effect on the total volunteer biomass. Volunteer biomass was significantly lower within 1.5 m of the S. spectabilis hedgerows than at positions further away. In the experiment cropped for five years, S. spectabilis reduced the volunteer regrowth biomass significantly. Two other hedgerow species, Dactyladenia barteri [(Hook f ex Oliv.) Engl.] and Flemingia macrophylla [(Willd.) Merrill] had no effect on the total amount of volunteer regrowth but did reduce volunteer biomass within 0.5 m of the hedgerows. S. spectabilis caused a stronger reduction of volunteer biomass than D. barteri and F. macrophylla at almost all distances from the hedgerows. The competitiveness of D. barteri and F. macrophylla is insufficient and their growth habit is unsuitable to out-compete undesired species in this alley cropping system. This revised version was published online in June 2006 with corrections to the Cover Date.     

Shortened bush fallow rotations with relay-cropped Inga edulis and Desmodium ovalifolium in wet central Amazonian Peru

In the Palcazu Valley alluvial Inceptisols are relay-cropped with maize-cassavaplantain in rotation with 2–5 years of tree fallow. These lands, of limited extent, yet important for Yanesha Indian subsistence production, are being cropped even more intensively as population increases and land is converted to other uses. The relay-planting of the tree-thicket combination Inga edulis with Desmodium ovalifolium into the on-farm crop sequence was evaluated as a means to accelerate fallow recovery and thereby shorten fallow rotations. Three experiments with Inga/Desmodium planted with cassava-plantain and one with rice under different weeding regimes after a Desmodium fallow were conducted. Inga and Desmodium were not chopped back or pruned during these experiments. Desmodium/Inga suppressed herbaceous weeds from one year after planting. Desmodium/Inga accumulated more woody biomass than natural fallows. Cassava yields were unaffected by the presence of Desmodium/Inga, while plantain yields were greater under Desmodium/Inga compared to natural weeds. Desmodium/Inga, while promising for shortening fallow rotations, demonstrated potential difficulties: increased labor for establishment, tendency of Desmodium to weediness in later crop cycles, and suppression of the natural regeneration of trees and shrubs.     

Adoption of improved fallow technology for soil fertility management in Zambia: Empirical studies and emerging issues

In the subsistence-agricultural region of eastern Zambia, less than 10% of the households have adequate supply of maize (Zea mays L.), the staple food, throughout the year. A major constraint to increasing crop production in the region is poor fertility status of the soil. In order to address this problem, improved fallow has been introduced as a technology for improving soil fertility within a short span of two to three years. Farmers have been testing the technology and a number of empirical studies have been undertaken over the years to identify the factors influencing farmers' decision to adopt the technology. This paper presents a synthesis of the results of adoption studies and highlights generic issues on the adoption of improved fallows in Zambia. The synthesis indicates that farmers' decision on technology adoption does not have a simple directed relationship of some technological characteristics only, but constitutes a matrix of factors including household characteristics, community level factors, socioeconomic constraints and incentives that farmers face, access to information, local institutional arrangements and macro policies on agriculture. The adoption of improved fallows is not strictly speaking a binary choice problem but a continuous process in which farmers occupy a position along a continuum in the adoption path. Further, adoption of improved fallows may not take place in a policy vacuum but needs to be facilitated by appropriate and conducive policy and institutional incentives. Several questions and issues that require further study emerge from the synthesis. These include determination of the relative importance of the factors in the adoption matrix, identification of the conditions under which farmers use a combination of inputs and their profitability under changing price scenarios, exact definition to delineate between `non-adopters', `testers' and `adopters' of agroforestry technologies, and understanding the impact of cash crop farming in farmers' adoption decisions of improved fallows (where off farm opportunities exist). Further, there is a need to determine the inter-relationship between household poverty, labor availability and the adoption of improved fallows and, to evaluate a combination of policy interventions at both national and local level to promote the adoption of agroforestry-based soil fertility management. This revised version was published online in June 2006 with corrections to the Cover Date.     

Weed seedbank characteristics of arable fields under different fallow management systems in the humid tropical zone of southeastern Nigeria

The composition and pattern of weed flora in arable fields are determined by their seedbank structure; but the influence of fallow management practices on weed seedbank structure is presently unknown. The objective of this study was to investigate weed seedbank characteristics and weed population dynamics in arable fields in natural and planted-fallow systems. The study plots were at Mbaise, a densely populated area of southeastern Nigeria, where farmers regenerate their exhausted soils by maintaining planted fallows of Dactyladenia barteri (Hook. F. ex Oliv.) Prance & F. White, and at Umuahia, a less-densely populated area in the same region, where farmers depend on natural bush fallow for soil regeneration. The effect of three years of fallow on the weed flora of arable fields in the two fallow management systems differed remarkably. The first flush of weeds on fields that were cultivated after three years of planted D. barteri fallow (Mbaise) consisted of 80% broadleaf weeds, 7% grass weeds and 13% sedges. On the other hand, the first flush of weeds on the natural bush fallow fields (Umuahia) of the same fallow duration as the D. barteri fallow system consisted of 17% broadleaf weeds, 70% grasses and 13% sedges. Three years of planted fallow caused 36% decrease in weed seedbank at Mbaise relative to the cropped field while the same duration of natural bush fallow caused a 31% increase in weed seedbank at Umuahia. These results show that the planted D. barteri fallow system has a higher potential to reduce weed pressure in smallholder agriculture than the natural bush fallow system and may explain in part why farmers in this humid forest zone have adopted the practice. This revised version was published online in June 2006 with corrections to the Cover Date.     

Plant growth, biomass production and nutrient accumulation by slash/mulch agroforestry systems in tropical hillsides of Colombia

Planted fallow systems under ‘slash and mulch’ management were compared with natural fallow systems at two farms (BM1 and BM2) in the Colombian Andes. The BM1 site was relatively more fertile than the BM2 site. Planted fallow systems evaluated included Calliandra calothyrsus CIAT 20400 (CAL), Indigofera constricta (IND) or Tithonia diversifola (TTH). During each pruning event slashed biomass was weighed, surface-applied to the soil on the same plot and sub-samples taken for chemical analyses. While Indigofera trees consistently showed significantly greater (p < 0.05) plant height and collar diameter than Calliandra trees at both study sites, only collar diameter in Indigofera was significantly affected at all sampling times by differences between BM1 and BM2. After 27 months, TTH presented the greatest cumulative dry weight biomass (37 t ha^–1) and nutrient accumulation in biomass (417.5 kg N ha^–1, 85.3 kg P ha^–1, 928 kg K ha^–1, 299 kg Ca ha^–1 and 127.6 kg Mg ha^–1) among planted fallow systems studied at BM1. Leaf biomass was significantly greater (P < 0.05) for CAL than IND irrespective of site. However, CAL and IND biomass from other plant parts studied and nutrient accumulation were generally similar at BM1 and BM2. At both sites, NAT consistently presented the lowest biomass production and nutrient accumulation among fallow systems. Planted fallows using Calliandra and Indigofera trees had the additional benefit of producing considerable quantities of firewood for household use. This revised version was published online in June 2006 with corrections to the Cover Date.     

The potential of alley cropping in improvement of cultivation systems in the high rainfall areas of Zambia I. Chitemene and Fundikila

Three trials investigating the potential of alley cropping to improve the traditional systems of cultivation, chitemene and fundikila, in the Northern Province of Zambia are described. Flemingia congesta, Tephrosia vogelii, and Sesbania sesban, were grown in association with finger millet, groundnut, cowpea, and maize in various traditional cropping sequences. The indigenous species Tephrosia vogelii and Sesbania sesban were not able to withstand repeated pruning and the long following dry season, and were replaced with Calliandra calothyrsus, and Cassia spectabilis.Over a four year period, there was no benefit by alley cropping with any of the tree species on crop yields, and yields in alley crop treatments even dropped significantly below the control treatments in the fourth year, casting doubt on the potential of alley cropping for sustainable production. There was also no consistent effect on soil chemical characteristics in any of the trials. It was suggested that this lack of beneficial response to alley cropping was due to low tree biomass production low quality of prunings, and an inappropriate cropping sequence. There was no evidence that alley cropping contributed to enhanced nutrient recycling, despite substantial localised pools of soil nutrients, particularly in the chitemene, with which recycling could potentially occur.     

Tree-root systems and herbaceous species-characteristics under tree species introduced into grazing lands in subhumid Cameroon

The effect of tree species on the characteristics of the herbaceous stratum, during the first five years of a fallow, was evaluated in the North of Cameroon (average annual temperature 28.2 °C, total annual rainfall 1050 mm). Treatments included a natural grazed herbaceous fallow, a natural ungrazed herbaceous fallow and three planted tree fallows (Acacia polyacantha Willd. ssp. campylacantha (Hochst. ex A. Rich.), Senna siamea Lam. and Eucalyptus camaldulensis Dehnh.), which were protected against grazing. Because tree species influenced light interception in different ways, as well as having different root patterns, they had different effects on the herbaceous stratum in terms of species composition and biomass. The grazed herbaceous fallow maintained the greatest species richness. Protection against grazing or the introduction of tree species associated with the absence of grazing induced both a progressive evolution to a particular species composition. The ungrazed herbaceous fallow consisted mainly of Andropogon gayanus Kunth, which provided the greatest biomass (8 t dry matter ha^–1 at the end of the fallow period). E. camaldulensis provided little shade and the lowest fine root mass in the top layer allowing the growth of A. gayanus and thus a greater herbaceous biomass (3.5 t DM ha^–1) than that found under the other tree species. Under the heavy shade of A. polyacantha, the herbaceous stratum consisted mainly of annual Pennisetum spp. (2.2 t DM ha^–1) and showed the greatest N concentration (1.3%), probably due to N₂ fixation by the tree species. After the fourth year, despite the relatively open tree canopy, S. siamea, which showed the highest fine root mass, had a strong depressive effect on the herbaceous stratum. This revised version was published online in June 2006 with corrections to the Cover Date.     

Benefits and costs of intercropping reforestation in the Atlantic lowlands of Costa Rica.

Agroforestry options as a means of promoting reforestation were testedby establishing with 10 farmers simple comparisons between pure plantations andtimber trees with annual or perennial intercrops. Two year-old intercroppedtrees on-farm had 22–48% greater height and 24–38% greater diameterthan trees in pure plantations. The exception was Cordiaalliodora that did not respond significantly to intercropping withcassava (Manihot esculenta). Only intercrops with the mostvaluable crops on agricultural soils produced a return to labour above thedailywage rate. On non-agricultural soils, farmers only intercrop a small area oftheir plantations with subsistence crops to meet home consumption needs. In thecontext of Costa Rica intercropping appeared to have a limited potential tofinance the reforestation costs of farmers.     

Babassu palm in the agroforestry systems in Brazil's Mid-North region

Babassu palms (Orbignya spp.) cover nearly 200,000 km² in Brazil, providing cash income, fuel, fibre, edible oil and food to a large number of tenant farm households. Babassu is closely integrated within pastoral and shifting cultivation systems of Mid-North Brazil. In pastures, babassu provides shade for cattle, aids soil moisture retention, produces organic matter, generates supplementary farm income at little cost, and offers year-round employment. On the other hand, the persistence of juvenile palms reduces pasture grass productivity due to plant competition, and therefore there is a trend to eradicate babassu through clearcutting and understorey suppression. At moderate densities of less than 100 individuals per ha, mature babassu palms in cropland do not appear to harm crop productivity. In such cases, palms are thinned and leaves of the remaining ones are cut back, supplying fuel for the burn and nutrients to the soil. However, reduced fallow cycles due to pasture conversion threaten babassu as well as crop productivity.

---

Resumo Palmeiras de babaçu (Orbignya spp.) occorrem em quase 200.000 km² no Brazil, proporcionando renda em dinheiro, combustível, fibras, óleo e alimentos para cerca de quinhentas mil famílias de pequenos produtores rurais, a maioria arrendatários ou posseiros. O babaçu aparece integrado dentro de sistemas de produção pastorís e de agricultura migratória no Meio Norte do Brazil. Nas pastagens, o babaçu provê sombra para o gado, ajuda na retenção da umidade no solo, produz matéria orgânica, gera renda suplementar com investimento mínimo, e oferece condições para fixar a força de trabalho durante a entressafra dos produtos agrícolas. A presença de palmeiras juvenís, porém, reduz a produtividade dos pastos devido a competição, o que leva os criadores de gado a erradicar o babaçu, suprimindo tanto as palmeiras juvenís, como as palmeiras adultas.No que diz respeito à agricultura migratória, o babaçu adulto em densidades moderadas, ou seja, até 100 indivíduos por hectare, aparentemente não reduz a produtividade das culturas anuais. Nestes casos, parte das palmeiras são desbastadas e as folhas das restantes cortadas, proporcionando combustível para a queima e nutrientes para o solo. No entanto, a redução nos ciclos de descanso devido à conversão das terras agrícolas em pastagens ameaça a produtividade quer do babaçu quer das próprias culturas.

     

Nitrogen contribution of five leguminous trees and shrubs to alley cropped maize in Ibadan, Nigeria

There are abundant local legume trees and shrubs potentially suitable for alley cropping systems in the sub-Saharan Africa, which are yet to be studied. The nitrogen contribution of two years old Albizia lebbeck and S. corymbosato yield of maize grown in alley cropping was compared to that of Senna siamea, Gliricidia sepium and Leucaena leucocephala in four seasons at Ibadan. Maize shoot biomass and maize grain yield in A. lebbeck alley compared favourably with that in G. sepium and L. leucocephala. Maize biomass and grain yield in S. corymbosa alleys were the lowest. Within A. lebbeck, L. leucocpehala, and G. sepium alleys there were no significant differences in the maize yield in the alleys that received 0, 40 or 80 kg N/ha. Application of more than 40 kg N/ha in S. corymbosa alleys was not necessary as there was no significant increase in maize yield at the higher level of nitrogen. Maize yield and N uptake in A. lebbeck alleys were not significantly different from yield and N uptake in G. sepium, and L. leucocephala at the same fertilizer level. There was a significant correlation between hedgerow tree biomass and maize grain yield. At the end of twelve weeks after pruning application, the organic residues of the pruning applied in the alleys ranged from 5% in G. sepium and 44% in A. lebbeck in the first year compared with the original pruning applied which showed that the slow rate of A. lebbeck decomposition could have a beneficial effect on the soil. The maize N recovery from applied N fertilizer was low (10–22%). Percentage N recovery from the prunings was low in the non-N fixing trees (12–22%), while the recovery was high (49–59%) in A. lebbeck as well as in the other nitrogen fixing tree prunings. Thus A. lebbeck, apart from enhancing maize growth and grain yield like in L. leucocephala and G. sepium, had an added advantage because it remained longer as mulching material on the soil because of its slow rate of decomposition. It was able to survive pruning frequencies with no die-back. This indicates that A. lebbeck is a good potential candidate for alley cropping system in West Africa. S. corymbosa performed poorly compared with the other legume trees. Though it responded to N fertilizer showing a positive interaction between the hedgerow and fertilizer application, it had a high die back rate following pruning periods and termite attack.This revised version was published online in November 2005 with corrections to the Cover Date.     

Productivity of hedgerow shrubs and maize under alleycropping and block planting systems in semiarid Kenya

A field experiment was conducted for eight cropping seasons from 1988 to 1991 in semiarid Machakos, Kenya, to compare the productivity of shrubs and crops in intercropping (alleycropping) versus block (sole) planting systems. The study, conducted in a split-plot experiment with three replications, consisted of two tree species (Leucaena leucocephala andSenna siamea syn.Cassia siamea) in the main-plots and combinations of two planting systems (alleycropping and block planting of the shrubs and maize) with five different ratios of land allocation for the shrub and crop (1000, 1585, 2080, 2575, and 0100) in sub-plots. Thus, a sole maize and a sole tree were included in the subplot treatments. The spacing between hedgerows of shrubs in intercropping was 6.7, 5, and 4 m, respectively, corresponding to 15, 20, and 25% land allocation to shrubs. The trees were pruned to 0.5 m height four times a year. Intercropped hedgerows of senna and leucaena produced 10% and 24% more biomass than their respective block planting systems. On average, leucaena produced more biomass than senna. Maize alleycropped with leucaena yielded 16% less grain than sole-crop maize, whereas senna intercropping caused hardly any maize-yield reduction. Compared with the respective sole-crop systems, leucaena intercropping did not affect land equivalent ratio (LER), whereas LER increased by 28% with senna intercropping. The different tree:crop land occupancy ratios did not affect the production of either the component species or of the total system, except that LER declined with incrreased spacing between hedgerows. It is concluded that in semiarid highlands of Kenya, leucaena and crops should better be grown in sole blocks, not in alleycropping. In the case of slow-growing species such as senna, intercropping is worthwhile to consider only if the additional labour needed does not pose a serious problem for management, and the species has fodder value.     

Litterfall,litter decomposition and the use of mulch of four indigenous tree species in the Atlantic lowlands of Costa Rica

Litterfall, forest-floor litter biomass and nutrients, short-term litter decomposition and the effects of leaf mulches on initial growth of maize were studied for four indigenous tree species with agroforestry potential:Stryphnodendron microstachyum Poepp. et Endl.(S. excelsum), Vochysia ferruginea Mart,Vochysia guatemalensis Donn. Sm. (V. hondurensis) andHyeronima alchorneoides (O), growing in a young experimental plantation in the Atlantic humid lowlands of Costa Rica. Total annual leaf litterfall was higher inV. ferruginea plots, followed byS. microstachyum, V. guatemalensis andH. alchorneoides; all with values comparable to those reported for other tree species grown in agroforestry combinations in humid tropical regions. Forest-floor litter accumulation was highest underV. ferruginea andV. guatemalensis. Both litterfall and forest-floor litter material had similar patterns in nutrient concentrations: N was higher inS. microstachyum, Ca was higher inV. guatemalensis, K was higher inH. alchorneoides; Mg was higher inV. guatemalensis andH. alchorneoides; H. alchorneoides andV. guatemalensis had the highest P.V. ferruginea litter decomposed more slowly, whileS. microstachyum apparently decomposed faster than the other species. The twoVochysia species showed increases in N and P concentration in decomposing litter after seven weeks in the field,H. alchorneoides showed an increase in litter N and a decrease in litter P, andS. microstachyum showed a net decrease in both N and P over the same time period. The patterns found in the litter bag study were confirmed by results obtained in a tethered-leaves experiment.S. microstachyum andV. ferruginea litters lost more weight when mixed in a 11 proportion than either of them alone. Maize seedlings growing in plots mulched withS. microstachyum andH. alchorneoides leaves showed greatest initial growth, confirming patterns found in decomposition and nutrient release studies. The results show that these species could be used in agroforestry combinations with different advantages according to the specific objectives desired, whether these are soil protection, nutrient recycling, or enhancement of the growth of associated crops.     

Establishment of two indigenous timber species in dairy pastures in Costa Rica

A silvopastoral model that combines the production of pasture herbage with valuable native timber species has potential to simultaneously address the multiple goals of reforestation, conservation of native species and enterprise intensification. The objective of this study was to design, establish and monitor early growth of a silvopastoral experiment on a dairy farm in the north Atlantic zone of Costa Rica. Two indigenous timber species, Vochysia guatemalensis and Hyeronima alchorneoides were planted with and without the tropical pasture legume, Arachis pintoi in a split plot design, (2 × 2) factorial arrangement of treatments with four replications. After the first two years, V. guatemalensis was significantly taller (3.1 m) than H. alchorneoides (2.5 m). The mean root collar diameter for V. guatemalensis was significantly larger (6.5 cm) than H. alchorneoides (4.5 cm). Two-year establishment was acceptable for the tree component (83 to 85% survival) but poor for A. pintoi (2 to 8% of the sward). The most important pest affecting the establishment of the timber species was the leaf cutter ant, Atta cephalotes. An insect larvae, Cosmopterix sp., severely damaged 39% of the V. guatemalensis trees by repeatedly attacking their apical meristems. The two-year establishment data was insufficient to accurately predict future wood volume. A hypothetical economic analysis concluded that the silvopastoral system must average at least 1.2 m³ wood volume/paddock/year (20 m³/ha/year) throughout the first ten years of growth to assure a positive economic return from timber. The experiment is planned for a ten year period, which corresponds to the estimated rotation length for harvesting the timber species. This revised version was published online in June 2006 with corrections to the Cover Date.     

Leaf quality of selected hedgerow species at two canopy ages in the derived savanna zone of West Africa

Prunings in hedgerow intercropping systems are a potential source of organic matter and their quality is an important characteristic driving decomposition and nutrient release. To determine the potential range of residue characteristics and the impact of canopy age on those, selected characteristics of Leucaena leucocephala, Gliricidia sepium, and Senna siamea hedgerow leaf residues were determined for young (5–10 weeks) and old (39–47 weeks) hedgerow canopies in six sites, representative in terms of soil for the derived savanna zone of West-Africa. The N content of the Leucaena and Senna residues decreased with age (from 4.8 to 3.9% and from 4.1 to 3.0%, respectively, while the N content of the Gliricidia residues remained constant (3.9%). The P content of all species decreased with age (from 0.30 to 0.20%, on average). The lignin content of the Leucaena residues increased, and their polyphenol content decreased with age, while for the Senna residues only an increase in polyphenol content with canopy age was evident. Neither the lignin nor the polyphenol content of the Gliricidia leaf residues was markedly affected by canopy age. For all species, the P and ADF content were positively correlated with the ash and lignin content, respectively. The N content of the Leucaena and Senna residues was positively correlated with their P content while the lignin content of the same species was negatively correlated with the polyphenol content. Because of the low variation in certain residue characteristics between sites, the presented data and equations could potentially be used to reasonably estimate those characteristics if no other information is available. For other characteristics, such as polyphenol contents, measurements are preferable in view of the relatively high variation encountered.This revised version was published online in November 2005 with corrections to the Cover Date.     

Nitrogen fixation capacity in the component species of contour hedgerows: how important?

The choice of an appropriate hedgerow species is one of the most critical decisions in exploiting the value of a contour hedgerow system. The implications of hedgerow species with nitrogen (N)-fixation capacity on hedgerow-crop competition and crop productivity have been widely debated. We examined the agronomic significance of N-fixation by comparing the performance of species representing three classes of hedgerow vegetation: A nitrogen-fixing tree legumeGliricidia sepium), a non-nitrogen fixing tree (Senna spectabilis syn.Cassia spectabilis), and a forage grass (Pennisetum purpureum). The 4-year study investigated the hedgerow biomass and nutrient yields, and their relative effects on the performance of two annual crops commonly grown in alley farming systems, with emphasis on hedgerow-crop interference. The work was done on an Ultic Haplorthox (pH 4.8, organic C 1.9%, total N 0.18%).Senna produced 46% more pruning biomass on an annual basis than didGliricidia; N supplied to the alley crops was similar toGliricidia in the first year of observation, but 20–30% higher in the succeeding years. Upland rice and maize grain yields and total dry matter were unaffected by tree species, but the nitrogen-fixing tree exerted less competitive effects on the annual crops growing in adjacent rows. Grass hedgerows reduced maize yields 86% by the second year, indicating an unsustainable drawdown of nutrients and water. We conclude that hedgerow systems composed of a nitrogen-fixing tree did not exert significant advantages compared to a non-fixing tree species, and that factors other than N-fixation were more important determinants for the choice of hedgerow species.     

Yields of maize/cassava intercrops grown with hedgerows of three multipurpose trees on an acid Ultisol of Cameroon

Maize/cassava were intercropped between hedgerows of Senna spectabilis [(DC.) Irwin and Barneby], Flemingia macrophylla [(Willd.) Merrill] and Dactyladenia barteri [(Hook f ex Oliv.) Engl.] for five consecutive years on an Ultisol in southern Cameroon. Crop yields and hedgerow biomass production in the third to fifth year of cropping are reported. S. spectabilis produced more biomass than F. macrophylla and D. barteri in all years. Cumulative maize grain and cassava tuber yields were highest in F. macrophylla alley cropping, outyielding the no-tree control consistently by 42 to 67% (average 56%). Between hedgerows of D. barteri and S. spectabilis, crops yielded 17% and 16% more than the no-tree control, respectively. However, between S. spectabilis hedgerows, yields were highly variable between years (–15% to +35% compared to the no-tree control) and thus the system is at risk of failure. F. macrophylla is recommended for continuous alley cropping of maize/cassava intercrop. The use of D. barteri may require fallow phases for biomass accumulation followed by cropping phases with rigorous pruning. Although this may lead to lower cumulative yields, the products of the fallow phase, such as stakes and firewood, may provide some compensation.This revised version was published online in November 2005 with corrections to the Cover Date.     

Decomposition and nutrient release from leaves,twigs and roots of three alley-cropped tree legumes in central Togo

The decomposition of leaves, twigs and roots of two diameter classes (<1.5 mm, 1.5–5 mm) were examined in an alley cropping experiment withGliricidia sepium, Calliandra calothyrsus andSenna sianea in the subhumid savanna of Central Togo using the litterbag technique. The effect of the application of leaves and twigs as mulch or green manure was examined. Gliricidia showed the most rapid mass loss. For all species, leaves decomposed faster than roots. Twigs had the lowest decomposition rate except forCalliandra. The two diameter classes of roots decomposed differently in the three species:Gliricidia fine roots decomposed faster than its coarser root fraction, the coarse roots ofSenna decomposed faster than the fine roots.Termites influenced the mass loss of twigs and roots to varying extents for the different species. In the case of twigs this was markedly influenced by the mode of application: green manure showed more termite frass than mulch.Nutrient release resembled the mass loss patterns of the prunings except for K, which was leached independently from mass loss. The release of the different nutrients was in the order CaGliricidia andSenna seemed to be best for mulch and green manure production at our site.