Black locust (<Emphasis Type="Italic">Robinia pseudoacacia</Emphasis> L.) as a multi-purpose tree species in Hungary and Romania: a review

Black locust (Robinia pseudoacacia L.) was the first North-American tree species imported to Europe at the beginning of the seventeenth century. It is commonly planted worldwide because of its adaptability to environmental stresses, its valuable wood, easy propagation, frequent and abundant seed production, excellent coppicing, high seedling survival, and relatively high wood yield. In Europe, Romania and Hungary have the most highly-developed black locust growing techniques and experiences. As a result of increasing interest in black locust in many countries, this review aims to provide a comprehensive overview of state-of-the-art site requirements, propagation, improvement and management (including growth and yield as well as use in energy plantations).     

北京石质山地燃料型能源树种刺槐无性系筛选

刺槐因其抗性强、生长速度快、薪材产量和热值均较高而成为许多地区的重要燃料型能源树种.针对北京石质山地优良高产燃料型能源刺槐无性系的筛选进行了较为深入的研究,从造林成活率、林木萌芽物候期、两个采薪周期树高和地径生长、两个采薪周期产薪量等方面对鲁刺12个无性系、豫刺6个无性系以及当地刺槐的差异进行了比较.研究结果表明,豫8059号、豫8033号和豫8017号等是适应北京石质山地立地条件的优质高产无性系,豫8026号、豫8062号和鲁9号也有潜力.建议在华北石质山地通过试验推广所选出的豫8059号、豫8033号、豫8017号和鲁9号等刺槐无性系,并推广其优化栽培配套技术.     

匈牙利刺槐((Robinia pseudoacacia L.)林的经营研究(英文)

匈牙利刺槐((RobiniapseudoacaciaL.)是在17世纪初从北美引入到欧洲的第一个森林树种。在匈牙利,刺槐是非常重要的速生树种,为经济和生态目的,可以成功营造刺槐人工林。结合该树种的生长特点选择营林措施,可以最大限度的提高刺槐人工林的成活率和生产力。本文根据林分结构和森林产量长期实验林的研究成果,提出了刺槐林的抚育作业次序。制定好的营林计划和模式将会培育出效益刺槐林,并能使土地经营者更好地接受该树种。同时,刺槐也是一个非常有用的能源生产树种,本文展示了对这方面的研究成果。图2表1参8。     

The feature of color alteration of bleached oak,beech, and black locust surfaces during artificial xenon radiation

Abstract

In Europe, large quantities of oak wood are available and industrial partners showed interest in bleached oak as oak's reinterpretation in order to reintroduce it on the design furniture market. In the present work besides oak two further wood species beech and black locust were also examined in comparison. Bleaching of beech has a relative well-established technique and can be done with good results with H₂O₂, contrary to oak where bleaching with the same agent results in a spotty discoloration of the surface with high-greenish incrust. A second substance is suggested to be used, to achieve the same significant bleaching effect for oak and black locust as for beech. As bleached wood surfaces are also subject to color changes due to sun radiation, bleached samples of the three wood species were exposed to artificial xenon radiation and their behavior was investigated, with focus on the influence of the type of application. Expecting industrial-scale application the possible influence of application type on the measure of total color change was also investigated. The feature of color development was described as exponential function of time for all three wood species and all three application methods.     

匈牙利刺槐改良与经营(英文)

刺槐是 17世纪初由北美引入欧洲的第一批林木树种 .刺槐之所以在世界快速传播 ,归功于其对环境条件的广泛适应能力 ,易于培育的特性 ,结实频繁、结种量大 ,萌蘖力强 ,种苗保存率高而且速生高产 ,同时又是优良的蜜源植物 .事实上 ,很少有什么病虫对其造成伤害 .刺槐是匈牙利现有林分中最重要的成林树种之一 ,它覆盖 2 0 %的林地 .匈牙利的刺槐的木材产量占全国产量的 18% .鉴于许多国家对培育刺槐的兴趣越来越浓 ,在此将匈牙利的经验和研究结果整理成文 ,期望在刺槐课题方面有所裨益     

匈牙利刺槐（（Robinia pseudoacacia L.）林的经营研究

匈牙利刺槐（（Robinia pseudoacacia L.)是在17世纪初从北美引入到欧洲的第个森林树种。在匈牙利，刺槐是非常重要的速生树种，为经济和生态目的，可以成功营造刺槐人工林。结合该树种的生长特点选择营林措施，可以最大限度的提高刺槐人工林的成活率和生产力。本文根据林分结构和森林产量长期实验林的研究成果，提出了刺槐林的抚育作业次序。制定好的营林计划和模式将会培育出效益刺槐林，并能使土地经营者更好地接受该树种。同时，刺槐也是一个非常有用的能源生产树种，本文展示了对这方面的研究成果。图2表1参8。     

Breeding and cultivation of black locust, Robinia pseudoacacia, in Hungary

Black or common locust was introduced into Hungary between 1710 and 1720. Its present extent is 276 000 ha, 18.2% of the total forest area. Since its introduction, black locust has always been closely associated with farming.Selection breeding followed numerous steps. The best shipmast-type groves in the best stands were surveyed for superior trees which were grafted for clone tests.The National Agricultural Council for Variety Testing approved three selected or introduced locust cultivars in 1973 and five more in 1979. The registered cultivars and a further 13 candidates are grouped into three categories according to their proposed uses: sawlogs; poles and posts; and finally beekeeping and decorative planting.Some cultivars such as ‘Zalai’, ‘Kiskunsági’, ‘Császártöltési’, ‘Egylevelü’ and ‘Váti 46’ are suitable both for forestry and beekeeping.Vegetative propagation systems have been introduced for mass propagation of the improved cultivars. The planned production of 8 million plants in 1984 covers two-thirds of the demand.The selected cultivars demand intensive technologies, of which planting after deep-ploughing on better sites is the best. Silvicultural models have been worked out for producing sawlogs on better sites and poles and posts on medium sites.Interest in black locust is growing both in Europe and Asia. The area of man-made locust stands has increased in the last 20 years from 337 000 to 1 890 000 ha. These figures exclude China, where the tree is extensively used. In recent years the energy and forage aspects have become more interesting. South Korea leads in this respect, but various projects are also being undertaken in Hungary.     

Early evaluation of micropropagated black locust （Robinia pseudoacacia L.） clones in Hungary

In Hungary, black locust (Robinia pseudoacacia L.) is one of the most important exotic species, forming entire stands. Its importance is increasing in many other countries. As a result of a new selection program eight black locust clones have been improved for establishing clone trials and a seed orchard. In this study juvenile growth and stem quality of micropropagated clones of black locust were evaluated on a marginal site in central Hungary. At ten years old, the clones ’MB17D3/4’ and ’PV201E2/4’ appeared to be especially promising for high quality wood production under arid hydrological conditions. A tissue culture method can be considered a suitable tool of propagating superior individual trees and offers new prospects for the rapid cloning of selected genotypes.     

Wood properties of juvenile and mature heartwood in Robinia pseudoacacia L.

The aim of this study is to characterise the properties of juvenile and mature heartwood of black locust (Robinia pseudoacacia L.). Content, composition and the subcellular distribution of heartwood extractives were studied in 14 old-growth trees from forest sites in Germany and Hungary as well as in 16 younger trees of four clone types. Heartwood extractives (methanol and acetone extraction) were analysed by HPLC-chromatography. UV microspectrophotometry was used to topochemically localise the extractives in the cell walls. The natural durability of the juvenile and mature heartwood was analysed according to the European standard EN 350-1. Growth as well as chemical analyses showed that, based on extractives content, the formation of juvenile wood in black locust is restricted to the first 10–20 years of cambial growth. In mature heartwood, high contents of phenolic compounds and flavonoids were present, localised in high concentrations in the cell walls and cell lumen of axial parenchyma and vessels. In juvenile wood, the content of these extractives is significantly lower. Juvenile wood had a correspondingly lower resistance to decay by Coniophora puteana (brown rot fungus) and Coriolus versicolor (white rot fungus) than mature heartwood.     

黑木耳丰产技术研究

通过对黑木耳丰产栽培技术20a研究和实践，总结黑木耳丰产的3个关键问题是：菌种，耳木边材量，技术管理，优质菌种比劣质菌种可提高产量87．1％，边材量多的耳木比少的提高产量62．9％，所以必须选择优质菌种和边材量多的耳木,昌黑木耳丰产的保证，主要是发好菌，使菌种定植在耳木中，再经干湿交替管理，以保证优质菌种和耳木边材发挥最大经济效应。     

Establishment of a transgenic system in fast-growing black locust (Robinia pseudoacacia L.)

The AhDREB1 gene, cloned from Atriplex hortensis L., was transferred into black locust (Robinia pseudoacacia L.) by an Agrobacterium-mediated transformation. The results suggest that stems of black locust sub-cultured in vitro for 20 d are suitable for genetic transformation. The optimum concentrations of kanamycin and cefotaxime were 30 and 150 mg.L-1, respectively. Impor-tant factors affecting the transformation efficiency were studied by means of a L9(34) orthogonal design. An effective system for ge-netic transformation in black locust was developed as follows: the stems were pre-cultured for 2 d, immersed in the Agrobacterium solution (OD6oo = 0.7) with 10 mg'L-1 acetosyringone for 21 min and then co-cultured for 2 d. The selection pressures, changing from low to high, could improve transformation efficiency. The transgenic plants were identified by a PCR method. The PCR results indicated that the AhDREB1 gene had been integrated into the genome of black locust and two lines of the transgenic plants were obtained.     

Identification of selected internal wood characteristics in computed tomography images of black spruce: a comparison study

The feasibility of identifying internal wood characteristics in computed tomography (CT) images of black spruce was investigated using two promising classifiers: the maximum likelihood classifier (MLC) and the back propagation (BP) artificial neural network (ANN) classifier. Nine image features including one spectral feature (gray level values), a distance feature, and seven textural features were employed to develop the classifiers. The selected internal wood characteristics to be identified included heartwood, sapwood, bark, and knots. Twenty cross-sectional CT images of a black spruce log were randomly selected to develop the two classifiers. The results suggest that both classifiers produced high classification accuracy. Compared with the MLC classifier (80.9% overall accuracy), the BP ANN classifier had better classification performance (97.6% overall accuracy). Moreover, statistical analysis reveals that the heartwood of the black spruce log used in this study is the easiest to identify by either classifier compared with the other three log features. The results also suggest that the separability of one wood characteristic from the other wood characteristics in black spruce CT images is mainly related to moisture content.     

Effective rainfall seasons for interannual variation in δ13C and tree-ring width in early and late wood of Chinese pine and black locust on the Loess Plateau, China

We studied the importance of effective rainfall for interannual variation in water use efficiency (WUE) and tree-ring growth of Chinese pine (Pinus tabulaeformis Carr.) and black locust (Robinia pseudoacacia L.) by examining correlations of seasonal precipitation with annual values of stable carbon isotope ratio (δ¹³C) and tree-ring width in early and late wood. The correlations with precipitation were examined for each month and for periods of all possible lengths from 2 to 22 months starting from January of the previous year to October of the current year. The period with the highest correlation was adopted as the most effective rainfall season for interannual variations in WUE and tree-ring width. In early wood, precipitation during the dry season (October to May) before the growing season was negatively correlated with δ¹³C in pine trees and positively correlated with ring width in pine and locust trees. In late wood, rainfall during the growing season in the current year was negatively correlated with δ¹³C in pine and locust trees, and positively correlated with ring width in locust trees. Our results demonstrated the differences in the water use strategies of pine and locust trees. The δ¹³C in pines indicated higher WUE and more conservative water use than in locust trees. Precipitation during the dry season affected the interannual variation in WUE and tree-ring growth in pine and locust trees, indicating that rainfall during the dry season is important for carbon gain and tree-ring growth during the following growing season.     

沙地杨树刺槐人工混交林的改良土壤功能及养分互补关系

杨树刺槐混交林是我国北方地区沿河及滨海沙地上成功的固氮树种和非固氮树种人工混交林。本文在简单介绍混交林生长情况的基础上,主要对林地土壤养分改良及其种间Ｎ、Ｐ养分互补关系进行了阐述。指出杨树刺槐混交林生产力提高主要是因为林地土壤养分（特别是Ｎ素养分）的极大改善及树种间Ｎ、Ｐ养分的互补。混交林土壤Ｎ素养分水平提高的根本原因是林地土壤微生物活性高,加快了含Ｎ较高枯落物分解速度,增强了土壤氨化及硝化强度。而杨树刺槐种间的Ｎ、Ｐ养分互补则主要表现在混交林中杨树可将吸收入体内的Ｐ素通过根系接触转移给相邻的刺槐,而刺槐则可将固定的Ｎ素通过根系接触转移给杨树,而且Ｎ、Ｐ转移的强度很大。这种奇妙的树种间养分互补机制对杨树刺槐混交林幼林期生产力提高是有重大意义的。     

Black locust (Robinia pseudoacada L.) clonal seed orchards in Hungary

Black locust (Robinia pseudoacacia L.) is one of the most important stand-forming tree species in Hungary and its importance is increasing in many countries. The main aim of the discussed new selection programme is to identify black locust clones with good performance and good form for setting up clonal seed orchards. As a result of selection programme 16 new black locust clones have been improved. In spring 2002 a black locust seed orchard was established with the newly selected clones. About 40% of the plants can be considered to belong to the height growth rate class 1 and 2. Hungary was the first country where micropropagated black locust planting material was used for seed orchard establishment.     

Black locust (Robinia pseudoacacia L.) clonal seed orchards in Hungary

Black locust (Robinia pseudoacacia L.) is one of the most important stand-forming tree species in Hungary and its importance is increasing in many countries. The main aim of the discussed new selection programme is to identify black locust clones with good performance and good form for setting up clonal seed orchards. As a result of selection programme 16 new black locust clones have been improved. In spring 2002 a black locust seed orchard was established with the newly selected clones. About 40% of the plants can be considered to belong to the height growth rate class 1 and 2. Hungary was the first country where micropropagated black locust planting material was used for seed orchard establishment.     

Role of income diversification in protecting natural forests: evidence from rural households in forest margins of Sri Lanka

Ranchers need alternative livestock feeds when herbaceous forages become limiting in summer. Our objectives were to determine: (1) leaf and stem biomass components, (2) nutritive value [in vitro dry matter digestibility (IVDMD), total nonstructural carbohydrate (TNC), N, and N digestibility] of leaves for animal browse, (3) concentration of the secondary metabolites robinin and mimosine, and (4) in vitro leaf and bark toxicity for black locust (Robinia pseudoacacia L.) and mimosa (Albizia julibrissin Durz.), respectively, pollarded at 50 cm in Arkansas, USA. Black locust exceeded mimosa for every yield component (leaf mass tree⁻¹, leaves shoot⁻¹, shoots tree⁻¹, shoot mass tree⁻¹, stem basal area, and biomass tree⁻¹) except mass leaf⁻¹. Projected yields were 1,900 and 1,600 kg leaves ha⁻¹ for black locust and mimosa, respectively, assuming a population of 12,300 trees ha⁻¹. Mimosa leaves had greater IVDMD, TNC, and N digestibility than black locust. Mimosa leaves exceeded the nutritional N requirements of growing cattle (Bos taurus L.) and goats (Capra hircus L.), but protein supplementation would be needed for growing goats grazing black locust leaves. Tissue concentrations of secondary metabolites robinin and mimosine were below detectable limits in black locust and mimosa, respectively. The extract of black locust bark, but not leaves, was toxic to bioassayed African green monkey (Cercopithecus aethiops L.) cells. Either black locust or mimosa could provide moderate quantities of high quality, rotationally grazed forage for goats during summer months when herbaceous forage may in short supply.     

刺槐萌条与根系年龄不同组合对林分结构及生物量的影响

探讨了萌条年龄与根系年龄的不同组合，对刺槐薪炭林结构和生物量的影响。结果表明，未割柴林分的萌条密度、平均高度、平均地径、叶面积指数、生物量和生产量，均随年龄增长而增加。割柴后萌生的林分，萌条数量增长缓慢，其余指标均呈下降趋势。随萌条与根系年龄差距的加大，下降的趋势愈加显著。     

Effects of black locust on productivity and nitrogen nutrition of intercropped barley

Effects of black locust (Robinia pseudoacacia L.) on productivity and N nutrition of barley (Hordeum vulgare L.) were evaluated under various management regimes (2 soil types, 3 levels of N fertilizer, and 3 cropping systems — barley alone, and barley intercropped with trees pruned or unpruned).Intercropping did not affect productivity and N nutrition of barley in 1988 when trees were small. However, there was a significant yield decline in 1989 as the trees grew bigger. On average, productivity of the sole crop was 8% higher in both soil types. Pruning and mulching moderated the yield reduction compared with the unpruned treatment. Competition for soil moisture was considered a major constraint. Nonetheless, the overall productivity (barley+black locust) from the intercropped treatments was 53% higher than sole cropping.In 1989 and 1990, intercropped barley had significantly higher grain and straw N concentrations (%). In 1989, for example, grain N content was 11% higher than in the sole crop. Removal of trees in 1990 resulted in significant increase in productivity and N content of subsequent barley crop relative to continuous sole cropping. From N nutrition viewpoint, barley from previously intercropped treatments showed superior quality and it had, on average, 23% higher grain N content than the sole crop. This was attributed to N₂-fixation and N return by black locust. It was estimated that black locust contributed about 36 kg N ha^–1 to the system.This study underscores the role black locust is potentially capable of playing in the development of sustainable and low-input agricultural systems in temperate regions. Nonetheless, the study also illustrates the importance of the below and above-ground interactions that occur in intercropped systems and the need for further research in this area.     

Black locust (Robinia pseudoacacia L.) clonal seed orchards in Hungary

Black locust (Robinia pseudoacacia L.) is one of the most important stand-forming tree species in Hungary and its importance is increasing in many countries. The main aim of the discussed new selection programme is to identify black locust clones with good performance and good form for setting up clonal seed orchards. As a result of selection programme 16 new black locust clones have been improved. In spring 2002 a black locust seed orchard was established with the newly selected clones. About 40% of the plants can be considered to belong to the height growth rate class 1 and 2. Hungary was the first country where micropropagated black locust planting material was used for seed orchard establishment.