Effect of tree species on carbon stocks in forest floor and mineral soil and implications for soil carbon inventories

Forest soil organic carbon (SOC) and forest floor carbon (FFC) stocks are highly variable. The sampling effort required to assess SOC and FFC stocks is therefore large, resulting in limited sampling and poor estimates of the size, spatial distribution, and changes in SOC and FFC stocks in many countries. Forest SOC and FFC stocks are influenced by tree species. Therefore, quantification of the effect of tree species on carbon stocks combined with spatial information on tree species distribution could improve insight into the spatial distribution of forest carbon stocks.We present a study on the effect of tree species on FFC and SOC stock for a forest in the Netherlands and evaluate how this information could be used for inventory improvement. We assessed FFC and SOC stocks in stands of beech (Fagus sylvatica), Douglas fir (Pseudotsuga menziesii), Scots pine (Pinus sylvestris), oak (Quercus robur) and larch (Larix kaempferi).FFC and SOC stocks differed between a number of species. FFC stocks varied between 11.1 Mg C ha⁻¹ (beech) and 29.6 Mg C ha⁻¹ (larch). SOC stocks varied between 53.3 Mg C ha⁻¹ (beech) and 97.1 Mg C ha⁻¹ (larch). At managed locations, carbon stocks were lower than at unmanaged locations. The Dutch carbon inventory currently overestimates FFC stocks. Differences in carbon stocks between conifer and broadleaf forests were significant enough to consider them relevant for the Dutch system for carbon inventory.     

A novel approach to optimize management strategies for carbon stored in both forests and wood products

We present a new approach to maximize carbon (C) storage in both forest and wood products using optimization within a forest management model (Remsoft Spatial Planning System). This method was used to evaluate four alternative objective functions, to maximize: (a) volume harvested, (b) wood product C storage, (c) forest C storage, and (d) C storage in the forest and products, over 300 years for a 30,000 ha hypothetical forest in New Brunswick, Canada. Effects of three initial forest age-structures and a range of product substitution rates were tested. Results showed that in many cases, C storage in product pools (especially in landfills) plus on-site forest C was equivalent to forest C storage resulting from reduced harvest. In other words, accounting for only forest, and not products and landfill C, underestimates true forest contributions to C sequestration, and may result in spurious C maximization strategies. The scenario to maximize harvest resulted in mean harvest for years 1–200 of 3.16 m³ ha⁻¹ yr⁻¹ and total C sequestration of 0.126 t ha⁻¹ yr⁻¹, versus 0.98 m³ ha⁻¹ yr⁻¹ and 0.228 t ha⁻¹ yr⁻¹ for a scenario to maximize forest C. When maximizing total (forest + products) C, mean harvest and total C storage for years 1–200 was 173% and 5% higher, respectively, than when maximizing forest C; and 218% and 6% higher, respectively, when maximizing substitution benefits (0.25 t of avoided C emissions per m³ of lumber used) in addition to total C. Initial forest age-structure affected harvest in years 1–50 < 34% among the four alternative management objective scenarios, and resulted in mean C sequestration rates of 0.31, 0.10, and −0.14 t ha⁻¹ yr⁻¹ when maximizing total C storage for young, even-aged, and old forests, respectively. Our results reinforce the importance of including products in forest-sector C budgets, and demonstrate how including product C in management can maximize forest contributions toward reduced atmospheric CO₂ at operational scales.     

Ecosystem carbon storage and partitioning in a tropical seasonal forest in Southwestern China

Tropical forests play an important role in the global carbon cycle. Despite an increasing number of studies have addressed carbon storage in tropical forests, the regional variation in such storage remains poorly understood. Uncertainty about how much carbon is stored in tropical forests is an important limitation for regional-scale estimates of carbon fluxes and improving these estimates requires extensive field studies of both above- and belowground stocks. In order to assess the carbon pools of a tropical seasonal forest in Asia, total ecosystem carbon storage was investigated in Xishuangbanna, SW China. Averaged across three 1 ha plots, the total carbon stock of the forest ecosystem was 303 t C ha⁻¹. Living tree carbon stocks (both above- and belowground) ranged from 163 to 258 t C ha⁻¹. The aboveground biomass C pool is comparable to the Dipterocarp forests in Sumatra but lower than those in Malaysia. The variation of C storage in the tree layer among different plots was mainly due to different densities of large trees (DBH > 70 cm). The contributions of the shrub layer, herb layer, woody lianas, and fine litter each accounted for 1–2 t C ha⁻¹ to the total carbon stock. The mineral soil C pools (top 100 cm) ranged from 84 to 102 t C ha⁻¹ and the C in woody debris from 5.6 to 12.5 t C ha⁻¹, representing the second and third largest C component in this ecosystem. Our results reveal that a high percentage (70%) of C is stored in biomass and less in soil in this tropical seasonal forest. This study provides an accurate estimate of the carbon pool and the partitioning of C among major components in tropical seasonal rain forest of northern tropical Asia. Results from this study will enhance our ability to evaluate the role of these forests in regional C cycles and have great implications for conservation planning.     

Carbon stock potentials of woodlands in north western lowlands of Ethiopia

ABSTRACT

This paper examined the potential of dry north western woodlands of Ethiopia (Adi Goshu, Lemlem Terara, and Gemed) for carbon stocks. Allometry equations were used to determine the aboveground, belowground, and dead woods biomasses; litter and herbaceous biomasses were determined using direct harvesting method. The result showed the estimated mean carbon stocks of the aboveground, belowground, and the dead wood biomass for the Untapped Boswellia Papyrifera Woodland (UW) in Lemlem Terara site were significantly higher (P < 0.05) than that of the Adi Goshu site. In the Gemed site, the mean Herb Biomass Carbon (HBC) stock was 1.2 Mg ha^?1, which is significantly highest (P < 0.05) than the other two study sites (Lemlem Terara, 0.42 Mg ha^?1 and Adi Goshu, 0.45 Mg ha^?1) for the Tapped Boswellia Papyrifera Woodland (TW). In UW, the mean soil carbon stock of the Lemlem Terara site (58.19 Mg ha^?1) was significantly (P < 0.05) higher than that of Adi Goshu (33.61 Mg ha^?1). In the case of the total carbon stocks in UW stratum, for the Adi Goshu site, the carbon stock was estimated to be about 55.26 Mg ha^?1 while 96.74 Mg ha^?1 for Lemlem Terara. Therefore, Carbon stock in different carbon pools (aboveground and belowground biomass, dead wood, litter, herbaceous biomass, and soil) has a potential to decrease the rate of enrichment of atmospheric concentration of carbon dioxide.     

Uncertainty of 21st century growing stocks and GHG balance of forests in British Columbia, Canada resulting from potential climate change impacts on ecosystem processes

Over the coming decades, climate change will increasingly affect forest ecosystem processes, but the future magnitude and direction of these responses is uncertain. We designed 12 scenarios combining possible changes in tree growth rates, decay rates, and area burned by wildfire with forecasts of future harvest to quantify the uncertainty of future (2010-2080), timber growing stock, ecosystem C stock, and greenhouse gas (GHG) balance for 67 million ha of forest in British Columbia, Canada. Each scenario was simulated 100 times with the Carbon Budget Model of the Canadian Forest Sector (CBM-CFS3). Depending on the scenario, timber growing stock over the entire land-base may increase by 14% or decrease by 9% by 2080 (a range of 2.8 billion m³), relative to 2010. However, timber growing stock available for harvest was forecast to decline in all scenarios by 26-62% relative to 2010 (a range of 1.2 billion m³). Forests were an annual GHG source in 2010 due to an ongoing insect outbreak. If half of the C in harvested wood was assumed to be immediately emitted, then 0-95% of simulations returned to annual net sinks by 2040, depending on scenario, and the cumulative (2010-2080) GHG balance ranged from a sink of −4.5 Pg CO₂e (−67 Mg CO₂e ha⁻¹) for the most optimistic scenario, to a source of 4.5 Pg CO₂e (67 Mg CO₂e ha⁻¹) for the most pessimistic. The difference in total ecosystem carbon stocks between the most optimistic and pessimistic scenarios in 2080 was 2.4 Pg C (36 Mg C ha⁻¹), an average difference of 126 Tg CO₂e yr⁻¹ (2 Mg CO₂e yr⁻¹ ha⁻¹) over the 70-year simulation period, approximately double the total reported anthropogenic GHG emissions in British Columbia in 2008. Forests risk having reduced growing stock and being GHG sources under many foreseeable scenarios, thus providing further feedback to climate change. These results indicate the need for continued monitoring of forest responses to climatic and global change, the development of mitigation and adaptation strategies by forest managers, and global efforts to minimize climate change impacts on forests.     

Changes in carbon stock following soil scarification of non-wooded stands in Hokkaido,northern Japan

To restore non-wooded stands dominated by dwarf bamboo species (Sasa kurilensis or S. senanensis) into forests, mechanical soil scarification has been applied in northern Japan since the 1960s. The treatment is followed both by natural regeneration and artificial planting. In this study, we quantified the total carbon stock (plants plus 0.3 m depth of soil) of these stands over 35-year age-sequences. The natural regeneration stands were gradually dominated by Betula ermanii. The carbon stock increased linearly to 215.1 ± 35.2 Mg C ha⁻¹ for a 37-year-old stand formerly dominated by S. kurilensis, and 181.1 ± 29.8 Mg C ha⁻¹ for a 34-year-old stand formerly dominated by S. senanensis. The latter was similar to that of a Picea glehnii plantation, formerly dominated by S. senanensis, with comparable stand age (160.3 ± 6.7 Mg C ha⁻¹ for 35-year-old stands). Although the carbon stock in plants quickly offset the untreated level, that in the soil remained depressed even in the older stands. This resulted in small differences in carbon stock of these stands with untreated dwarf bamboo stands. We conclude that natural regeneration following scarification could be a prime option for carbon sink management in the region. However, we should take a long rotation period (i.e., >50 years) to ensure a carbon sink state. A potential of further improvements of the practice, including that reduce intensity of soil disturbance, was presented.     

Impact of bark beetle (Ips typographus L.) disturbance on timber production and carbon sequestration in different management strategies under climate change

The likely environmental changes throughout the next century have the potential to strongly alter forest disturbance regimes which may heavily affect forest functions as well as forest management. Forest stands already poorly adapted to current environmental conditions, such as secondary Norway spruce (Picea abies (L.) Karst.) forests outside their natural range, are expected to be particularly prone to such risks. By means of a simulation study, a secondary Norway spruce forest management unit in Austria was studied under conditions of climatic change with regard to effects of bark beetle disturbance on timber production and carbon sequestration over a time period of 100 years. The modified patch model PICUS v1.41, including a submodule of bark beetle-induced tree mortality, was employed to assess four alternative management strategies: (a) Norway spruce age-class forestry, (b) Norway spruce continuous cover forestry, (c) conversion to mixed species stands, and (d) no management. Two sets of simulations were investigated, one without the consideration of biotic disturbances, the other including possible bark beetle damages. Simulations were conducted for a de-trended baseline climate (1961–1990) as well as for two transient climate change scenarios featuring a distinct increase in temperature. The main objectives were to: (i) estimate the effects of bark beetle damage on timber production and carbon (C) sequestration under climate change; (ii) assess the effects of disregarding bark beetle disturbance in the analysis.Results indicated a strong increase in bark beetle damage under climate change scenarios (up to +219% in terms of timber volume losses) compared to the baseline climate scenario. Furthermore, distinct differences were revealed between the studied management strategies, pointing at considerably lower amounts of salvage in the conversion strategy. In terms of C storage, increased biotic disturbances under climate change reduced C storage in the actively managed strategies (up to −41.0 tC ha⁻¹) over the 100-year simulation period, whereas in the unmanaged control variant some scenarios even resulted in increased C sequestration due to a stand density effect.Comparing the simulation series with and without bark beetle disturbances the main findings were: (i) forest C storage was higher in all actively managed strategies under climate change, when biotic disturbances were disregarded (up to +31.6 tC ha⁻¹ over 100 years); and (ii) in the undisturbed, unmanaged variant C sequestration was lower compared to the simulations with bark beetle disturbance (up to −69.9 tC ha⁻¹ over 100 years). The study highlights the importance of including the full range of ecosystem-specific disturbances by isolating the effect of one important agent on timber production and C sequestration.     

Deadwood in New Zealand's indigenous forests

We present the results of a systematic, unbiased national survey of deadwood volume and biomass in New Zealand's remaining indigenous forests based on an 8-km grid of 894 permanent plots. New Zealand's old growth evergreen temperate forests are largely comprised of long-lived, slow-growing tree species typically growing in cool, humid conditions; collectively these conditions are thought to promote accumulation of high deadwood stocks. We estimated deadwood biomass and volume in New Zealand's forests and compared these stocks with published values from other broadleaved evergreen temperate forests. Mean deadwood biomass in New Zealand was 54 Mg ha⁻¹ but ranged across plots from 0 to 550 Mg ha⁻¹. Mean deadwood volume was 158 m³ ha⁻¹ and ranged across plots from 0 to 1890 m³ ha⁻¹. Fallen logs accounted for 63% of total deadwood volume and 65% of total deadwood biomass, with standing dead trees being the remainder. Each piece of deadwood was classified into one of three broad decay classes and >40% of deadwood was fallen logs of the intermediate decay class. Deadwood biomass and volume varied 1.8- and 1.9-fold, respectively, among forest types and was greatest in broadleaved forests, dominated by Weinmannia racemosa (Cunoniaceae), Metrosideros umbellata (Myrtaceae) and Metrosideros robusta, and broadleaf-Nothofagus (Nothofagaceae) forests supporting the large tree species Nothofagus fusca. Deadwood biomass and volume were least in broadleaf-conifer admixtures. We used structural equation models to determine whether deadwood biomass could be predicted from climate and environment (vapor pressure deficit, elevation and slope), live tree biomass, forest composition (captured by two ordination axes), wood density of live trees, and tree size (a proxy for stand age). The model that best fit the data retained only vapor pressure deficit, live tree biomass and the first ordination axis as predictors of deadwood biomass. However, this model predicted just 2.4% of the variation in deadwood biomass, suggesting that additional factors not captured by this dataset, such as disturbance dynamics, may control deadwood abundance. Comparisons with other temperate and tropical forests did not support the hypothesis that New Zealand's cool temperate rainforests support higher than expected biomass or volume of deadwood.     

Short- and long-term effects of thinning and prescribed fire on carbon stocks in ponderosa pine stands in northern Arizona

Euro-American logging practices, intensive grazing, and fire suppression have increased the amount of carbon that is stored in ponderosa pine (Pinus ponderosa Dougl. Ex Laws) forests in the southwestern United States. Current stand conditions leave these forests prone to high-intensity wildfire, which releases a pulse of carbon emissions and shifts carbon storage from live trees to standing dead trees and woody debris. Thinning and prescribed burning are commonly used to reduce the risk of intense wildfire, but also reduce on-site carbon stocks and release carbon to the atmosphere. This study quantified the impact of thinning on the carbon budgets of five ponderosa pine stands in northern Arizona, including the fossil fuels consumed during logging operations. We used the pre- and post-treatment data on carbon stocks and the Fire and Fuels Extension to the Forest Vegetation Simulator (FEE-FVS) to simulate the long-term effects of intense wildfire, thinning, and repeated prescribed burning on stand carbon storage.The mean total pre-treatment carbon stock, including above-ground live and dead trees, below-ground live and dead trees, and surface fuels across five sites was 74.58 Mg C ha⁻¹ and the post-treatment mean was 50.65 Mg C ha⁻¹ in the first post-treatment year. The mean total carbon release from slash burning, fossil fuels, and logs removed was 21.92 Mg C ha⁻¹. FEE-FVS simulations showed that thinning increased the mean canopy base height, decreased the mean crown bulk density, and increased the mean crowning index, and thus reduced the risk of high-intensity wildfire at all sites. Untreated stands that incurred wildfire once within the next 100 years or once within the next 50 years had greater mean net carbon storage after 100 years compared to treated stands that experienced prescribed fire every 10 years or every 20 years. Treated stands released greater amounts of carbon overall due to repeated prescribed fires, slash burning, and 100% of harvested logs being counted as carbon emissions because they were used for short-lived products. However, after 100 years treated stands stored more carbon in live trees and less carbon in dead trees and surface fuels than untreated stands burned by intense wildfire. The long-term net carbon storage of treated stands was similar or greater than untreated wildfire-burned stands only when a distinction was made between carbon stored in live and dead trees, carbon in logs was stored in long-lived products, and energy in logging slash substituted for fossil fuels.     

Carbon sequestration in the growing stock of trees in Finland under different cutting and climate scenarios

In this work the aim was to determine how carbon sequestration in the growing stock of trees in Finland is dependent on the forest management and increased production potential due to climate change. This was analysed for the period 2003–2053 using forest inventory data and the forestry model MELA. Four combinations of two climate change and two management scenarios were studied: current (CU) and gradually warming (CC) climate and forest management strategies corresponding to different rates of utilisation of the cutting potential, namely maximum sustainable removal (Sust) or maximum net present value (NPV) of wood production (Max). In this analysis of Finland, the initial amount of carbon in the growing stock was 765 Mt (2,802 Tg CO₂). At the end of the simulation, the carbon in the growing stock of trees in Finland had increased to 894 Mt (3,275 Tg CO₂) under CUSust, 906 Mt (3,321 Tg CO₂) under CUMax, 1,060 Mt (3,885 Tg CO₂) under CCSust and 1,026 Mt (3,758 Tg CO₂) under CCMax. The results show that future development of carbon in the growing stock is not only dependent on climate change scenarios but also on forest management. For example, maximising the NPV of wood production without sustainability constraints results, over the short term, in a large amount of wood obtained in regeneration cuttings and a consequent decrease in the amount of carbon in growing stock. Over the longer term, this decrease in the carbon of growing stock in regenerated forests is compensated by the subsequent increase in fast-growing young forests. By comparison, no drastic short-term decrease in carbon stock was found in the Sust scenarios; only minor decreases were observed.     

Methane and nitrous oxide emissions from mature forest stands in the boreal forest, Saskatchewan, Canada

Despite the spatial significance of Canada's boreal forest, there is very little known about CH₄ and N₂O emissions from non-peatlands within it. The primary objective of this project was to study the atmosphere–soil exchange of CH₄ and N₂O at three sites in the boreal forest of central Saskatchewan. In the summers of 2006 and 2007, CH₄ and N₂O emissions were measured along transects in three different mature forest stands (aspen, black spruce and jack pine) using a sealed chamber method. At the aspen site, the gross rates of mineralization and nitrification, and the relative contribution of nitrification and denitrification to N₂O emissions, were also measured using the ¹⁵N isotope dilution technique. Results indicated that the jack pine and black spruce sites were slight sinks of CH₄ (−0.123 g CH₄–C m⁻² yr⁻¹and −0.017 g CH₄–C m⁻² yr⁻¹ respectively in 2006 and −0.095 g CH₄–C m⁻² yr⁻¹and 0.045 g CH₄–C m⁻² yr⁻¹ respectively in 2007), whereas the aspen site was a net source (4.40 g CH₄–C m⁻² yr⁻¹ in 2006 and 19.60 g CH₄–C m⁻² yr⁻¹ in 2007). The high CH₄ emissions at the aspen site occurred at depressions that were water-filled due to above-average precipitation levels in 2005–2007. All three sites had very low cumulative N₂O emissions, ranging from −0.002 to 0.014 g N₂O–N m⁻² yr⁻¹ in both years. The ¹⁵N results indicated that N cycling at the aspen site was very conservative, allowing little N to escape the system as N₂O; the emissions that did occur were due primarily to a nitrification-related process.     

EU-wide maps of growing stock and above-ground biomass in forests based on remote sensing and field measurements

The overall objective of this study was to combine national forest inventory data and remotely sensed data to produce pan-European maps on growing stock and above-ground woody biomass for the two species groups “broadleaves” and “conifers”. An automatic up-scaling approach making use of satellite remote sensing data and field measurement data was applied for EU-wide mapping of growing stock and above-ground biomass in forests. The approach is based on sampling and allows the direct combination of data with different measurement units such as forest inventory plot data and satellite remote sensing data. For the classification, data from the Moderate Resolution Imaging Spectroradiometer (MODIS) were used. Comprehensive field measurement data from national forest inventories for 98,979 locations from 16 countries were used for which tree species and growing stock estimates were available. The classification results were evaluated by comparison with regional estimates derived independently from the classification from national forest inventories. The validation at the regional level shows a high correlation between the classification results and the field based estimates with correlation coefficient r = 0.96 for coniferous, r = 0.94 for broadleaved and r = 0.97 for total growing stock per hectare. The mean absolute error of the estimations is 25 m³/ha for coniferous, 20 m³/ha for broadleaved and 25 m³/ha for total growing stock per hectare. Biomass conversion and expansion factors were applied to convert the growing stock classification results to carbon stock in above-ground biomass. As results of the classification, coniferous and broadleaved growing stock as well as carbon stock of the above-ground biomass is mapped on a wall-to-wall basis with a spatial resolution of 500 m × 500 m per grid cell. The mapped area is 5 million km², of which 2 million km² are forests, and covers the whole European Union, the EFTA countries, the Balkans, Belarus, the Ukraine, Moldova, Armenia, Azerbaijan, Georgia and Turkey.     

Biomass burning in Brazil's Amazonian “arc of deforestation”: Burning efficiency and charcoal formation in a fire after mechanized clearing at Feliz Natal, Mato Grosso

Estimates of greenhouse-gas emissions from deforestation are highly uncertain because of high variability in key parameters and because of the limited number of studies providing field measurements of these parameters. One such parameter is burning efficiency, which determines how much of the original forest's aboveground carbon stock will be released in the burn, as well as how much will later be released by decay and how much will remain as charcoal. In this paper we examined the fate of biomass from a semideciduous tropical forest in the “arc of deforestation,” where clearing activity is concentrated along the southern edge of the Amazon forest. We estimated carbon content, charcoal formation and burning efficiency by direct measurements (cutting and weighing) and by line-intersect sampling (LIS) done along the axis of each plot before and after burning of felled vegetation. The total aboveground dry biomass found here (219.3 Mg ha⁻¹) is lower than the values found in studies that have been done in other parts of the Amazon region. Values for burning efficiency (65%) and charcoal formation (6.0%, or 5.98 Mg C ha⁻¹) were much higher than those found in past studies in tropical areas. The percentage of trunk biomass lost in burning (49%) was substantially higher than has been found in previous studies. This difference may be explained by the concentration of more stems in the smaller diameter classes and the low humidity of the fuel (the dry season was unusually long in 2007, the year of the burn). This study provides the first measurements of forest burning parameters for a group of forest types that is now undergoing rapid deforestation. The burning parameters estimated here indicate substantially higher burning efficiency than has been found in other Amazonian forest types. Quantification of burning efficiency is critical to estimates of trace-gas emissions from deforestation.     

Sustainable management regimes for Europe's forests — a projection with EFISCEN until 2050

In Europe, forest policy discussions are moving towards a European Union-wide strategy. This will further intensify the relations between European countries in the field of forests and forest management. European-wide forest planning and decision-making require that policy makers have insight into the long-term development of European forests under alternative regimes. The European Forest Information Scenario Model (EFISCEN) was used to make projections of the development of the European forests under four scenarios: (1) business as usual; (2) EFISCEN European timber trend studies (ETTS); (3) maximum sustainable production; and (4) multifunctional management. The simulations were carried out for 30 countries individually, i.e. harmonized scenarios were run, but the special circumstances and demands that play a role in each country were taken into account. The simulations covered 139.2 million ha increasing to 143 million ha in 2050. The initial year varied per country, but was mostly in the region of 1990. The average age of European forests was 57 years in 1990 with a mean growing stock of 142 m³ ha⁻¹. The results showed that the future European total fellings may vary between a stable amount of 400 million m³ year⁻¹ in the ‘business as usual’ scenario to 647 million m³ year⁻¹ in the ‘maximum sustainable production’ scenario. The other two scenarios incorporated a 9% gradual increase in fellings over the first 30 years of the simulation period (i.e. 0.3% year⁻¹). The average growing stock will rise to approximately 250 m³ ha⁻¹ in 2050, with the exception being the ‘maximum sustainable production’ scenario, in which the growing remains at approximately 137 m³ ha⁻¹. The average net annual increment remains at approximately 5 m³ ha⁻¹ year⁻¹ throughout the simulation period, almost irrespective of the scenario. In the multifunctional scenario, special attention was paid to nature values by increasing the area of strict reserves from 4 million ha in 1990 to 12.3 million ha in 2050 (8.6% of the total forest area). The assumed increase in fellings of 0.3% year⁻¹ appeared possible in combination with this area of reserves. The simulations showed that growing stock development and increment development differed very much for each country separately per scenario. Therefore, the results show a strong need for maintaining the national diversity that constitutes European forestry within harmonized European-wide forest management strategies. In this article, we address what the consequences of each scenario are for wood production, biodiversity, and environmental functions of the forest. The results provide policy makers with a challenge on whether to intervene in the ongoing trend of build-up of growing stock and whether to choose for biodiversity, for increased use of domestically produced wood products, or a combination of these, but spatially separated.     

Short-term effects of forest restoration management on non-symbiotic nitrogen-fixation in western Montana

Forest restoration treatments involving selection harvest and prescribed fire have been applied throughout the Rocky Mountain West with only a limited understanding of how these treatments influence plant community composition and soil processes. Forest restoration treatments, especially those involving fire, have the potential to reduce N capital on site. Unfortunately there has been only limited effort to investigate the effects of forest restoration treatments on forest ecosystem N inputs, especially free living N-fixation in soil and woody residues. Recent studies have highlighted the potential for decaying woody roots to serve as hot spots for N-fixation. The fire and fire surrogates (FFS) study site at Lubrecht Experimental Forest in Western Montana provided a unique opportunity to investigate the effect of restoration treatments on N-fixation. We set out to examine how prescribed fire, selection harvest, and a combination of both influence free living N-fixing bacteria that colonize decomposing woody roots, mineral soil, and soil crusts. Soil, root, and soil crust samples were collected from replicated treatment plots in August 2005 and soil samples were recollected in May 2006 just following snowmelt. Acetylene reduction assays were run on all samples, and extractable inorganic N and potentially mineralizable N (PMN) were measured in mineral soil. While restoration treatments caused an increase in dead roots associated with stumps and fire killed trees, N-fixation rates were nearly non-existent in these root systems. Nitrogen-fixation rates were not significantly influenced by treatments in decomposing woody roots or in mineral soil, but were slightly greater (P < 0.10) in soil crusts when the control stand was compared to treated plots. Nitrogen-fixation rates were also greater in mineral soil than in roots. Soil collected in August exhibited greater rates of N-fixation than soil collected in May which we attributed to higher moisture and an increase in available N following spring thaw. Average rates of free living N-fixation across the treatment plots at Lubrecht were low (0.26 kg N ha⁻¹ year⁻¹), but over time we estimate that these sources, along with the sparse population of symbiotic N-fixing plants and wet N deposition, would replenish soil N lost through fire or harvesting in approximately 40–100 years.     

Decomposition of Pinus radiata coarse woody debris in New Zealand

The decomposition of Pinus radiata (D. Don) stems, coarse woody roots and stumps was studied in Tarawera forest, Bay of Plenty region, North Island, New Zealand. The study examined the residues from two thinning events with 6 and 11 years of decay. Changes in the mass of stems, and density of roots and stumps were used to estimate the decay rate constants using a single exponential model. The decay rate of stems was not significantly related to DBH and averaged 0.1374 year⁻¹ (22 years for 95% mass loss). The decay rate of coarse woody roots was not significantly different to stem decay and averaged 0.1571 year⁻¹ (19 years for 95% mass loss). A large range in stump decay rates was measured and a significantly lower decay constant was observed for stumps (0.1101 year⁻¹, 27 years for 95% mass loss), possibly due to the stumps being kept alive after felling through root grafting and a resistance to decay due to the presence of resin. The concentration of C remaining in stems and stumps increased with mass loss from 52% to 55% C after 11 years of decay. The C concentration in coarse woody roots initially increased but then declined near to the original level of 50% after 11 years of decay. Nitrogen concentrations increased substantially in all components with decay.     

Production and quality of senesced and green litterfall in a pine–oak forest in central-northwest Mexico

Litterfall is an important ecological process in forest ecosystems, influencing the transfer of organic matter, carbon (C), nitrogen (N), phosphorous (P) and other nutrients from vegetation to the soil. We examined the production of different litterfall fractions as well as nutrient content and nutrient inputs by senesced and green leaf-litter in a semiarid forest from central Mexico. From September 2006 to August 2007, monthly litter sampling was carried out in monospecific and mixed stands of Quercus potosina and Pinus cembroides. Litterfall displayed a marked bimodal pattern with the largest annual amount (5993 ± 655 kg ha⁻¹ yr⁻¹) recorded in mixed stands, followed by Q. potosina (4869 ± 510 kg ha⁻¹ yr⁻¹), and P. cembroides (3023 ± 337 kg ha⁻¹ yr⁻¹). Leaves constituted the largest fraction of total litterfall reaching almost 60%, while small branches contributed with 20–30%. Overall, N content in leaf-litter was higher while lignin content was significantly lower for Q. potosina than for P. cembroides. Thus, greater litter quality together with higher litter production caused the largest C, N and P inputs to forest soils to occur in monospecific Q. potosina stands. Green leaf fall displayed significantly lower lignin:N and C:N ratios in Q. potosina than P. cembroides suggesting faster decomposition and nutrient return rates by the former. Although we recorded only two green leaf fall events, they accounted for 18% and 11% of the total N and P input, respectively, from leaf-litter during the study period. Apart, from the large spatiotemporal heterogeneity introduced by differences in litter quantity and quality of evergreen, deciduous and mixed stands, green litterfall appears to represent a much more important mechanism of nutrient input to semiarid forest ecosystems than previously considered.     

Diversity and spatio-temporal dynamics of dead wood in a temperate near-natural beech forest (<Emphasis Type="Italic">Fagus sylvatica</Emphasis>)

The diversity, spatial patterns and temporal dynamics of dead wood were examined within the near-natural beech forests (Fagus sylvatica) of Serrahn (North-eastern Germany). Data were collected in an 8 ha sample plot and in two permanent plots (0.36 and 0.25 ha) that had been established at the end of the 1960s. The mean volume of dead wood was 94 m³ ha⁻¹, amounting to 14% of the total volume of all trees. The dead wood displayed a large variation in dead wood type, tree size and decay class. Standing dead wood accounted for about one-third of the total dead wood volume. The densities of standing dead trees were about 10% of the densities of the living trees over a wide range of diameters. The overall spatial distribution of dead trees exhibited a random pattern. Among the different dead wood types, standing entire dead trees and uprooted trees deviated from this pattern by displaying a significantly aggregated pattern. In the permanent plots a high mortality of overstorey trees was observed (1.3% year⁻¹) and the average amount of dead wood increased greatly from 2.9 to 111.6 m³ ha⁻¹ over the 35-year observation period. The near-natural beech forests of Serrahn have experienced a long period of low human interference. Nevertheless, our results suggest that the structure and dynamics of dead wood are strongly affected by the last major disturbance events that took place at the end of the Middle Ages. Information about the forest history is therefore a basic requirement when interpreting the results obtained in near-natural forests.     

Carbon Stock in Community Managed Hill Sal (Shorea robusta) Forests of Central Nepal

Community forests of developing countries are eligible to participate in the Reducing Emissions from Deforestation and Forest Degradation (REDD+) scheme. For this, estimation of carbon stock and the sequestration is essential. The carbon stock in the living biomass of nine community managed Shorea robusta forests of the mid hill regions of central Nepal (managed for 4–29 yr) were estimated. The carbon stock of trees and shrubs was estimated using an allometric equation while the biomass of herbaceous vegetation was estimated by the harvest method. The carbon stock in the living biomass of the studied forests ranged from 70–183 Mg ha^?1(mean: 120 Mg ha^?1) and it increased with increasing soil organic carbon. However, the carbon stock did not vary significantly with species richness and litter cover. The biomass and carbon stock in the forests managed for >20 yr were significantly higher than in the forests managed for < 20 yr. The carbon stock increased with the management duration (p < .05) with sequestration rate of 2.6 Mg C ha^?1 yr^?1. The local management has had positive effects on the carbon stock of the forests and thus the community forests have been acting as a sink of the atmospheric CO₂. Therefore, the community managed forests of Nepal are eligible to participate in the REDD+ scheme.