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1.

Key message

Selective thinning is a more viable method for beech coppice conversion to high forest when compared with thinning from below as it enhances tree growth, reduces mortality of the remaining trees, and allows to obtain stands with a higher mechanical stability and larger crowns.

Context

Beech forests in North-East Italy have been largely managed as coppice. Due to socio-economic changes, a large conversion to high forests program started in the second half of the past century.

Aims

A long-term experiment testing the effects on tree growth and stability of two different conversion methods (thinning from below—method A; selective thinning—method B) was implemented.

Methods

Both silvicultural treatments started in 1979 with a first thinning followed by a second one in 1997. All trees were periodically measured in order to assess mortality, stability, and growth during the period 1979–2010. In 2010, an assessment of stem quality and crown size was also performed.

Results

Both methods were economically viable, but method B acted with a higher intensity both in 1979 and in 1997, thus making the harvest more profitable for the owners. Moreover, method B enhanced tree growth, especially in the period after the first thinning, reduced mortality, and allowed to obtain stands with a higher mechanical stability and with larger crowns.

Conclusion

It would be possible to adopt some of the criteria prescribed with method B in future thinnings over the large areas actually managed with method A, as prescribed by the law.
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2.

Key message

Segregating stands and logs based on internal wood properties is likely to lead to improvements in value for forest and mill owners, but some situations were found where no segregation was the best alternative. Where segregation was the best alternative, segregating logs at the landing, or stands based on pre-harvest inventory assessments, led to the greatest value improvements.

Context

The benefits of segregating stands, stems and logs based on wood properties are not clear due to the high variability of wood properties, poor market signals for wood with superior properties and poor understanding of the costs across the value chain.

Aims

The aim of this study was to determine if the benefits of segregating stands and logs outweighed the additional costs.

Methods

A techno-economic model (SEGMOD) was constructed that allowed comparisons of segregation at different approaches in the supply chain. The model was populated with Pinus radiata (D.Don) stand, cost and price data from companies operating in four forestry regions of New Zealand. A total of 255 segregation scenarios were modelled, which included variations in segregation approach, stand type, stand location, terrain type, market focus and market horizon.

Results

Segregating logs based on internal wood properties led to improvements in stumpage and mill door value for most of the scenario sets evaluated. The No Segregation option was found (infrequently) to be best in unpruned stands. Segregating logs based on pre-harvest inventory assessments or at the landing would appear to be the best approach.

Conclusion

The economic benefits of segregating stands and logs for forest and mill owners outweighed the additional costs in most of the scenarios evaluated.
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3.

Key message

Silver fir transplantations along elevational gradients revealed a high diversity but no local adaptation. Populations displayed similar abilities to adapt to new environments including those due to climate change.

Context

The sustainability of forest stands depends on the ability of species and communities to adapt by combining plasticity and genetic evolution. Although well-documented at the scale of species distributions, the variability and adaptation of forest tree genetic resources are less understood at the short-distance scale.

Aims

We analysed the effects of genetic and environmental factors on the local-scale phenotypic diversity of traits related to adaptation in Abies alba. We also sought to highlight local adaptation, revealing past selection.

Methods

Six adaptive traits related to growth, phenology and survival were measured on seedlings from 57 half-sib families collected from 15 provenances and planted in a nine-site reciprocal transplant experiment distributed along three elevational gradients.

Results

Most part of the phenotypic variability was attributed to the environmental factors. Provenances and families had also significant effects on seedling performances, but the genetic variability was mostly attributed to the families. No pattern of local adaptation was observed, except in the presence of lateral branches in the driest garden.

Conclusion

The absence of local adaptation suggests a similar ability of all silver fir populations to develop in the various environments. This result provides favourable conditions for coping with the ongoing climate change without exotic resources enrichment.
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4.

Key message

We generate flexible management rules for black pine stands, adaptable to alternative stand management situations and entailing thinnings, final-felling, and salvage cuts, based on the results on 270 stand level optimizations.

Context

Forest management instructions often rely on the anticipated prediction of the stand development, which poses a challenge on variable economic and environmental conditions. Instead, an alternative approach to better adapt forest management decisions to changing conditions is defining flexible rules based on thresholds that trigger management operations.

Aims

This article develops rules for the adaptive management of P. nigra stands in Catalonia (Spain) addressing the risk of fire and post-fire forest management.

Methods

The stochastic version of the simulation-optimization system RODAL was used to optimize the management of forest stands in three sites under different fire probability levels. A total of 270 optimizations were done varying site fertility, fire probability, and economic factors. The results of the optimizations were used as the basis of flexible forest management rules for adaptive stand management.

Results

The developed management rules defined the basal area limit for thinning, the thinning intensity, the mean tree diameter at which regeneration cuttings should start, and the basal area below which a salvage cutting should be done. Fire risk was not a significant predictor of the models for thinning and final cutting rules.

Conclusion

The presented rules provide a flexible tool for forest management during the stand development and under changing conditions when the management objective is to maximize economic profitability of timber production.
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5.

Key message

Optimal management of cork oak forest stands was analyzed for different site indices and cork growth rates. Optimal debarking intervals varied during the rotation and were sometimes shorter or longer than the officially recommended range of 9–14 years.

Context

Quercus suber L. is one of the most important multipurpose tree species in the Mediterranean area. Its main product is cork, appreciated for its elasticity, impermeability, and thermal insulation properties. Cork oaks are debarked at constant intervals, which vary from 9 to 14 years depending on the area. However, since the growth rate of cork is not constant during the rotation, it may be optimal to use variable debarking intervals.

Aims

This study optimized the debarking and cutting schedules of Quercus suber stands and analyzed the influence of economic and stand-related factors on optimal management.

Methods

The study employed a simulation system where the existing growth and yield models for Quercus suber were used with a non-linear derivative-free optimization algorithm. Discount rates and cork prices were tested as economic factors and cork growth rate and site productivity as stand-related factors.

Results

The optimal debarking interval varied during the rotation. Increasing cork growth rate increased the optimal number of debarkings and shortened their interval. Decreasing discount rate increased the optimal number of debarkings during rotation while decreasing cork price decreased the number of debarkings.

Conclusion

The profitability of the management of cork oak stands depends on site fertility and stand density; management is not profitable on poor sites or at high discount rates. This study is the first that simultaneously optimizes the cutting and debarking schedule of cork oak stands, allowing the debarking interval to vary.
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6.

Key message

Carbon stock density was quite similar in planted vs natural forest of Masson’s pine ( Pinus massoniana Lamb.) in China across three ages (7, 15, and 50 years). The stock in the standing trees was larger in planted than in natural forests, but this difference was compensated by larger stocks in the soil and the debris of natural forests.

Context

Most studies on the carbon stocks are focused on management strategies to maximize carbon stocks. We still lack data comparing planted vs natural conifer forests.

Aims

We compared carbon storage in the different compartment (vegetation, soil, debris) along a chronosequence of Masson’s pine plantations vs natural forests.

Methods

We investigated 58 Masson’s pine (Pinus massoniana Lamb.) forest stands (20 m?×?50 m plots), that differ in stand management (planted and natural forests) and age (young, middle-aged, and mature ages) and then calculated the carbon stock densities of vegetation biomass (tree, shrub, and herb), debris, and soil.

Results

The carbon stock densities in the planted and natural Masson’s pine forest ecosystems ranged from 78 to 210 Mg ha?1 and from 97 to 177 Mg ha?1 respectively. The carbon stock densities in the vegetation were significantly greater in planted forests than in natural forests. A lower carbon stock density in debris and soil alleviated the increase of biomass carbon stock densities in planted vs natural forests, leading to similar carbon stock densities at ecosystem level. The carbon stock densities in the vegetation increased with age, whereas those of debris and soil remained stable.

Conclusions

Planted forests of Masson’s pine sequester similiar amounts of carbon at ecosystem level to those in natural forests, reinforcing the idea that planted pine forests can contribute to the mitigation of greenhouse gas emission.
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7.

Key message

LiDAR data (low-density data, 0.5 pulses m ?2 ) represent an excellent management resource as they can be used to estimate forest stand characteristics in short-rotation willow coppice (SRWC) with reasonable accuracy. The technology is also a useful, practical tool for carrying out inventories in these types of stands.

Context

This study evaluated the use of very low-density airborne LiDAR (light detection and ranging) data (0.5 pulses m?2), which can be accessed free of charge, in an SRWC established in degraded mining land.

Aims

This work aimed to determine the utility of low-density LiDAR data for estimating main forest structural attributes and biomass productivity and for comparing the estimates with field measurements carried out in an SRWC planted in marginal land.

Methods

The SRWC was established following a randomized complete block design with three clones, planted at two densities and with three fertilization levels. Use of parametric (multiple regression) and non-parametric (classification and regression trees, CART) fitting techniques yielded models with good predictive power and reliability. Both fitting methods were used for comprehensive analysis of the data and provide complementary information.

Results

The results of multiple regression analysis indicated close relationships (Rfit 2 = 0.63–0.97) between LiDAR-derived metrics and the field measured data for the variables studied (H, D20, D130, FW, and DW). High R 2 values were obtained for models fitted using the CART technique (R 2 = 0.73–0.94).

Conclusion

Low-density LiDAR data can be used to model structural attributes and biomass yield in SRWC with reasonable accuracy. The models developed can be used to improve and optimize follow-up decisions about the management of these crops.
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8.

Key message

Static site indices determined from stands’ top height are derived from different forest inventory sources with height and age information and thus enable comparisons and modeling of a species’ productivity encompassing large environmental gradients.

Context

Estimating forest site productivity under changing climate requires models that cover a wide range of site conditions. To exploit different inventory sources, we need harmonized measures and procedures for the productive potential. Static site indices (SI) appear to be a good choice.

Aims

We propose a method to derive static site indices for different inventory designs and apply it to six tree species of the German and French National Forest Inventory (NFI). For Norway spruce and European beech, the climate dependency of SI is modeled in order to estimate trends in productivity due to climate change.

Methods

Height and age measures are determined from the top diameters of a species at a given site. The SI is determined for a reference age of 100 years.

Results

The top height proves as a stable height measure that can be derived harmoniously from German and French NFI. The boundaries of the age-height frame are well described by the Chapman-Richards function. For spruce and beech, generalized additive models of the SI against simple climate variables lead to stable and plausible model behavior.

Conclusion

The introduced methodology permits a harmonized quantification of forest site productivity by static site indices. Predicting productivity in dependence on climate illustrates the benefits of combined datasets.
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9.

Key message

Mixing sessile oak and Scots pine in central France to reduce intraspecific competition for water resources did not improve the ability of these two species to withstand severe drought during the summer.

Context

In order to reduce the impact of increasingly extreme droughts on forests, managers must adapt their practices to future climate conditions. Maintaining a greater diversity of tree species in temperate forest ecosystems is one of the recommended options.

Aims

We addressed how interactions between sessile oak and Scots pine in mixed forests in central France affect their functional response to drought.

Methods

We characterized the carbon isotope composition (δ13C) in the tree growth rings formed during wet (2001, 2007) or dry (2003, 2004) summers for each of the two species growing both in pure and in mixed stands in order to compare the effect of stand composition on variations in carbon isotope discrimination (Δ13C) among contrasted years.

Results

The severe drought in 2003 induced a strong decrease in Δ13C for all trees and in all stands as compared to 2001. This decrease was greater in pine than in oak. There was no significant difference between pure and mixed stands in the response of either species to drought.

Conclusion

Mixing sessile oak and Scots pine in stands in central France does not improve the ability of either species to withstand severe drought during the summer.
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10.

Key message

Trees with otherwise equal dimensions have different leaf areas if they are located in different stand types. While leaf area of European larch is affected by mixture proportion, leaf area of Norway spruce is affected by stand density.

Context

Leaf area is a key parameter for evaluating growth efficiency of trees, and therefore needs to be measured as consistently and accurately as possible. This is even more important when comparing monospecific and mixed stands.

Aims

The aim of the study is to find combinations of parameters and allometric relationships that can be used to estimate accurately the leaf area of individual trees.

Methods

Allometries of the measured leaf area of 194 trees in 12 stands were analysed in order to find variables affecting leaf area. Existing functions from the literature were validated. Finally, models were fitted to find the most appropriate method for estimating leaf area of mixed and monospecific stands of Norway spruce and European larch.

Results

Allometric relationships of leaf area to other measurable characteristics of trees vary in different stand types. Besides individual tree dimensions such as diameter and crown surface area, leaf area of Norway spruce is related to stand density, whereas the leaf area of European larch is dependent on the admixture of Norway spruce in the stand.

Conclusion

In contrast to models for estimating individual tree leaf area of Norway spruce, models for leaf area of European larch have to consider mixture proportions in order to correctly interpret the growth efficiency of mixed stands.
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11.

Key message

Self-thinning lines are species- and climate-specific, and they should be used when assessing the capacity of different forest stands to increase biomass/carbon storage.

Context

The capacity of forests to store carbon can help to mitigate the effects of atmospheric CO2 rise and climate change. The self-thinning relationship (average size measure ~ stand density) has been used to identify the potential capacity of biomass storage at a given density and to evaluate the effect of stand management on stored carbon. Here, a study that shows how the self-thinning line varies with species and climate is presented.

Aims

Our main objective is thus testing whether species identity and climate affect the self-thinning line and therefore the potential amount of carbon stored in living biomass.

Methods

The Ecological and Forest Inventory of Catalonia was used to calculate the self-thinning lines of four common coniferous species in Catalonia, NE Iberian Peninsula (Pinus halepensis, Pinus nigra, Pinus sylvestris and Pinus uncinata). Quadratic mean diameter at breast height was chosen as the average size measure. The self-thinning lines were used to predict the potential diameter at a given density and study the effect of environmental variability.

Results

Species-specific self-thinning lines were obtained. The self-thinning exponent was consistent with the predicted values of ?3/2 and ?4/3 for mass-based scaling for all species except P. sylvestris. Species identity and climatic variability within species affected self-thinning line parameters.

Conclusion

Self-thinning lines are species-specific and are affected by climatic conditions. These relationships can be used to refine predictions of the capacity of different forest stands to increase biomass/carbon storage.
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12.

Key message

From 1973 to 1991, Polish SO 2 emissions above 3250 Gg/year resulted in a decline of fir Abies alba Mill. After stresses connected with these emissions, five main diameter at breast height (DBH) structural types were described. This type of heterogeneous forest structure is supposed to increase forest resistance and resilience to abiotic, biotic and anthropogenic disturbances.

Context

The analyses of forest structure are important under the current scenario of global change, since heterogeneous structures allow for better responses to disturbances. Forests with more complex structures should present greater vitality.

Aims

The main hypotheses were as follows: (1) the temporal changes of atmospheric SO2 emissions caused (a) the abrupt changes in the tree DBH radial increment and (b) the death of fir trees; and (2) atmospheric SO2 emissions and related fir decline and recovery processes ultimately result in the development of stands characterised by diverse DBH structures.

Methods

Radial growth trends of 49 and 215 fir trees in the older and younger generations, respectively, and 85 dead fir trees were evaluated. Using data collected in 32 stands, the DBH structural types were identified, and the shapes of these types were illustrated.

Results

The structural diversification of forest patches may influence forest resistance and resilience to disturbances; five main structural types were identified: ML1 and ML2 represent DBH distributions of multi-layered stands, and OS, TS1 and TS2 represent DBH distributions of one- and two-storied stands.

Conclusion

Structural diversity of forests was a response to SO2 emissions; fir trees had the ability to increase their radial growth, although there were still high SO2 emissions.
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13.

Key message

Short-rotation forestry using eucalyptus in degraded oak forests in the semi-arid area of NW Morocco can be a useful strategy to avoid further degradation and carbon loss from this ecosystem, but it might be constrained by nutrient and water supply in the long term.

Context

Land degradation and deforestation of natural forests are serious issues worldwide, potentially leading to altered land use and carbon storage capacity.

Aims

Our objectives were to investigate if short-rotation plantations can restore carbon pools of degraded soils, without altering soil fertility.

Methods

Carbon and nutrient pools in above- and below-ground biomass and soils were assessed using stand inventories, harvested biomass values, allometric relationships and selective sampling for chemical analyses.

Results

Carbon pools in the total ecosystem were low in the degraded land and in croplands (6–13 Mg ha?1) and high in forests (66–94 in eucalyptus plantations; 86–126 in native forests). The soil nutrient status of eucalyptus stands was intermediate between degraded land and native forests and increased over time after eucalyptus introduction. All harvest scenarios for eucalyptus are likely to impoverish the soil but, for the moment, the soil nutrient status has not been affected.

Conclusion

Afforestation of degraded land with eucalyptus can be a useful restoration tool relative to carbon storage and soil fertility, provided that non-intensive forestry is applied.
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14.

Key message

Stand density has a positive effect on C, K and Mg concentration in needle litterfall and a negative one on C, N, Ca, K, Mg, P, S, Zn, and Cu release from needle litter. Consequently, forest management practices such as thinning decrease nutrient concentration in needle litterfall and accelerate nutrient release from decomposing needles in Pinus halepensis plantations in Spain.

Context

Silvicultural practices usually include stand density reduction resulting in changes in litterfall and litter decomposition rates. Little is known about the effect on nutrient concentrations in litterfall and nutrient release during decomposition even when this is the main path of nutrient return to soils.

Aims

The aims of the study are to evaluate the seasonal pattern of nutrient concentration in litterfall, to study how nutrients are released from needle litterfall during decomposition, and to assess whether local basal area of the stand affects nutrient concentration of litterfall and nutrient release during litter decomposition.

Methods

Eight plots were established on each of four stands covering the widest range in local basal area. A littertrap and 15 litterbags were placed on each plot. Periodically, needle litterfall and litter contained in the litterbags were analyzed for C, N, Ca, K, Mg, P, S, Fe, Cu, Mn, and Zn.

Results

Local basal area had a positive effect on C, K, and Mg concentration in needle litterfall and a negative effect on the release of all the nutrients studied but Fe and Mn during the first 2 years of litter decomposition.

Conclusion

Density management of stands has an impact on nutrient cycling, reducing nutrient concentration in needle litterfall, and accelerating nutrient release during decomposition.
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15.

Key message

Soil texture and temperature-related variables were the variables that most contributed to Nothofagus antarctica forest height in southern Patagonia. This information may be useful for improving forest management, for instance related to the establishment of silvopastoral systems or selection of suitable sites for forest reforestation in southern Patagonia.

Context

Changes in forest productivity result from a combination of climate, topography, and soil properties.

Aims

The relative importance of edaphic and climatic variables as drivers of productivity in Nothofagus antarctica forests of southern Patagonia, Argentina, was evaluated.

Methods

A total of 48 mature stands of N. antarctica were selected. For each study site, we measured the height of three mature dominant trees, as an indicator of productivity. Seven soil, five spatial, and 19 climatic features were determined and related to forest productivity. Through partial least squares regression analyses, we obtained a model that was an effective predictor of height of mature dominant trees in the regional data set presented here.

Results

The four variables that most contributed to the predictive power of the model were altitude, temperature annual range, soil texture, and temperature seasonality.

Conclusion

The information gathered in this study suggested that the incidence of the soil and temperature-related variables on the height of dominant trees, at the regionally evaluated scale, was higher than the effect of water-related variables.
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16.

Key message

The invasion of moso bamboo ( Phyllostachys edulis (Carriere) J. Houzeau) into neighboring Cryptomeria japonica (L. f.) D. Don plantations significantly altered soil P status and dynamics. This alteration in phosphorus dynamics must be considered when assessing the ecological consequence of moso bamboo invasion in subtropical China.

Context

Moso bamboo is a native species that commonly invades into adjacent forests in Asia. Such invasions may significantly alter soil chemical characteristics because moso bamboo has very different traits compared with the tree species it displaces. However, few studies have investigated the effects of moso bamboo invasion on soil phosphorus (P) dynamics.

Aims

The objective of this study was to investigate the effects of moso bamboo invasion on soil P dynamics. Specifically, we quantified soil total P, available P, acid phosphatase activity (APA), and microbial biomass P (MBP) in moso bamboo-invaded coniferous stands and compared them to uninvaded stands and pure moso bamboo stands.

Methods

We compared seasonal dynamics of soil P (e.g., total P, available P, APA, and MBP) over a 24-month period among three stand types at Lushan mountain in subtropical China: Cryptomeria japonica plantation (CR), Cryptomeria japonica plantation invaded by Phyllostachys edulis (PH-CR), and Phyllostachys edulis stand (PH).

Results

Total soil P concentration was significantly lower in PH-CR than in CR and PH stands, but soil available P concentration was significantly lower in CR and PH stands. Soil APA was significantly higher in PH-CR than in CR and PH stands. Similarly, soil MBP concentration was higher in PH-CR than in CR and PH stands. Also, soil total P, available P, APA, and MBP concentrations displayed seasonal fluctuations in PH-CR, but remained relatively stable in CR and PH stands during the 2 years.

Conclusion

The invasion of moso bamboo into adjacent C. japonica stands significantly increased soil available P, acid phosphatase activity, and microbial biomass phosphorus, but decreased soil total P. The implication of these changes to ecosystem composition, structure, and function must be explicitly considered in managing moso bamboo invasion in subtropical China.
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17.

Key message

Bark thickness was shown to vary between regions, stands, and trees. Bark thickness prediction equations of different model complexity can be suggested depending on the purpose of application. Site and stand conditions, which influence variation of growth rate to a large extent, seem to have a strong influence on bark thickness, with better site quality leading to smaller relative bark thickness.

Context

For many applications in forestry and forest science, local or regional species-specific bark thickness equations are used to estimate inside-bark diameters with outside-bark diameter measurements.

Aims

The objectives of this study were (1) to assess variation in bark thickness due to tree and stand factors in two Norway spruce (Picea abies (L.) Karst) datasets from different time periods, (2) to compare and evaluate alternative established model forms for estimating bark thickness, and (3) to assess spatial variation in bark thickness to estimate the effects of environmental factors on bark thickness.

Methods

Different bark thickness models were chosen from the literature and compared for their predictive quality for new measurements and a dataset from the 1970s. Mixed-effect modelling was applied to account for the hierarchical data structure, and generalized additive mixed models were used to analyse spatial effects and the influence of climatic factors, such as precipitation and temperature.

Results

A strong positional autocorrelation of bark thickness within trees and within plots could be shown. Bark thickness was smaller in the new data compared to the measurements from the 1970s. The variation between stands could not be explained by the tested environmental factors, but tree age had a strong positive effect on bark thickness.

Conclusion

In the study region, the variation of site productivity and individual growth rate seem to have a strong influence on bark thickness, whereas no significant effect of large scale climatic factors could be found.
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18.

Key message

Non-stochastic portfolio optimization of forest stands provides a good alternative to stochastic mean-variance optimization when available statistical data is incomplete. The suggested approach has a theoretical background in the areas of robust optimization, continuous multicriteria decision-making, and fuzzy theory. Resulting robust portfolios only show slight economic losses compared to the efficient frontier of a stochastic optimization.

Context

Economic optimization addressing diversification in mixed uneven-aged forest stands is a useful tool for forest planners.

Aims

The study aims to compare two approaches for optimizing rotation age cohort portfolios under risk. Rotation age cohorts emerge from age-based regeneration-harvesting operations simulated for two tree species: Picea abies and Fagus sylvatica.

Methods

The first optimization approach is a stochastic mean-variance approach. The second is a non-stochastic optimization approach, which has rarely been applied to optimize tree species composition and the distribution of harvested timber over many periods. It aims at relatively good solutions, even if the deviation from the initially assumed return is very high. The objective function for both approaches is sensitive to the selection of various harvesting periods for different parts of the stand. For the stochastic approach, the objective function maximizes the annuitized net present value (economic return) for specific levels of risk by allocating area proportions to harvesting periods and tree species. In the non-stochastic approach, the allocation of area proportions instead minimizes the maximum deviation from the greatest possible economic return among many uncertainty scenarios (non-stochastic approach).

Results

Portfolios from both approaches were diverse in rotation age cohorts. The non-stochastic portfolios were more diverse when compared with portfolios from the efficient frontier, which showed the same standard deviation. However, P. abies clearly dominated the non-stochastic portfolios, while stochastic portfolios also integrated beech to a greater extent, but only in very low risk portfolios. The economic losses of the non-stochastic portfolios compared to the efficient frontier of the mean-variance approach lay between 1 and 3% only for different levels of accepted risk.

Conclusion

The non-stochastic portfolio optimization over a large uncertainty space is so far uncommon in forest science, yet provides a viable alternative to stochastic optimization, particularly when available data is scarce. However, further research should consider ecological effects, such as increased resistance against hazards of conifers in mixed stands.
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19.

Key message

The diversity of forest management systems and the contrasted competition level treatments applied make the experimental networks of the GIS Coop, a nationwide testing program in the field of emerging forestry topics within the framework of the ongoing global changes.

Context

To understand the dynamics of forest management systems and build adapted growth models for new forestry practices, long-term experiment networks remain more crucial than ever.

Aims

Two principles are at the basis of the experimental design of the networks of the Scientific Interest Group Cooperative for data on forest tree and stand growth (GIS Coop): contrasted and extreme silvicultural treatments in diverse pedoclimatic contexts.

Methods

Various forest management systems are under study: regular and even-aged stands of Douglas fir, sessile and pedunculate oaks, Maritime and Laricio pines, mixed stands of sessile oak, European silver fir, and Douglas fir combined with other species. Highly contrasted stand density regimes, from open growth to self-thinning, are formalized quantitatively.

Results

One hundred and eighty-five sites representing a total of 1206 plots have been set up in the last 20 years, where trees are measured regularly (every 3 to 10 years). The major outputs of these networks for research and management are the calibration/validation of growth and yield models and the drawing up of forest management guides.

Conclusion

The GIS Coop adapts its networks so that they can contribute to develop growth models that explicitly integrate pedoclimatic factors and thus also contribute to research on the sustainability of ecosystems under environmental and socio-economic changes.
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20.

Key message

In tree communities, tree size inequality reduces productivity and interacts with tree shade tolerance to modulate stand productivity, with a higher productivity in stands where shade-intolerant species dominate shade-tolerant species in size.

Context

Positive diversity–productivity relationships have been reported in different plant communities, including tree communities. These effects may be strongly related to both structural diversity and functional diversity, but also to their interactions if there is a non-random distribution of species functional characteristics among canopy layers.

Aims

We explore the relative effects on forest productivity of tree species diversity, tree size inequality, and species shade tolerance diversity, as well as the effect of the distribution of tree shade tolerance in the canopy.

Methods

We used 11,054 mixed-species forest plots from the French Forest Inventory (IGN) distributed throughout France (2006–2011). We analyzed the effects of species richness, shade tolerance diversity, and height inequality on forest plot productivity, represented by basal area annual increment over a period of 5 years, while controlling for first-order structure characteristics (basal area and quadratic mean diameter) and environmental factors (soil water budget and sum of growing degree days). Using the covariance between tree height and shade tolerance in mixed species canopies, we also explored the effect of the distribution of species’ shade tolerance among canopy layers.

Results

The results showed a positive effect of species richness (effect size, 0.02) and a negative effect of height inequality (??0.05) on mixed-forest productivity. We also showed that a negative covariance between shade tolerance and height (e.g., higher proportion of shade-tolerant species in lower height classes) increased productivity (0.01). Shade tolerance diversity did not affect productivity.

Conclusion

In tree communities, as shown previously in monospecific forest stands, tree size inequality reduces productivity. This effect is modulated by the distribution of shade tolerance among canopy layers. Previous studies on species diversity effect have generally overlooked the importance of the size structure and the size hierarchy of functional characteristics. These effects are, however, crucial and deserve to be explored in greater detail.
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