生物结皮对荒漠草本植物群落结构的影响

通过人工去除生物结皮试验,分析古尔班通古特沙漠生物结皮对常见荒漠草本植物群落结构的影响。根据人为去除生物结皮的试验结果,结合生物结皮对种子萌发、幼苗存活以及土壤养分和水分等影响,综合分析生物结皮对草本植物多样性的影响。结果表明:尖喙牻牛儿苗(Erodium oxyrrhynchum)、条叶庭荠(Alyssum linifolium)、琉苞菊(Hyalea pulchella)、荒漠庭荠(Alyssum desertorum)及旱麦草(Eremopyrum triticeum)的生物量总和占总生物量的90%以上,是该沙漠的优势草本植物。去除生物结皮后,草本植物的植被高度、物种丰富度和多样性指数都显著提高;但是,植被盖度、密度和群落生物量却显著降低。说明生物结皮的存在改变了古尔班通古特沙漠南缘荒漠草本植物的群落结构,植被盖度和群落生物量提高,这对于防风固沙、稳定沙面具有重要的生态意义。     

黄土高原生物结皮对土壤水分入渗的影响

利用圆盘入渗仪测量6种处理（无生物结皮、生物结皮、长芒草、长芒草＋生物结皮、柠条、柠条＋生物结皮）下水分的入渗特征,并进行对比分析。结果表明：①无论有、无植被,生物结皮的存在都会阻碍水分的入渗,无生物结皮的稳定入渗速率和累积入渗量均大于有生物结皮;植被的存在,加剧了生物结皮的阻水性。②Kostiakov模型对无结皮、长...     

古尔班通古特沙漠地衣结皮对放牧踩踏干扰的小尺度响应

古尔班通古特沙漠地衣结皮对放牧踩踏干扰及其所引起的生境异质性具有重要的指示作用。研究表明：① 放牧踩踏干扰主要集中于结皮层（0~5 cm）,随机调查的样方中,90%的样方干扰率均低于30%,生物土壤结皮(BSCs)破损尚处于较安全的范围;② 放牧踩踏干扰降低了BSCs总盖度,不同类型的结皮对践踏干扰的反应具差异性,其中,真菌-藻类共生形成的地衣结皮与干扰率呈极显著负相关（P<0.01）,干扰率>30%的样方中,抗风蚀能力较强的苔藓结皮、地衣结皮盖度均显著低于未受干扰样方,地衣共生体的破坏导致真菌数量显著增加（P<0.05）;③ 有机质、土壤容重等土壤理化指标均与干扰率呈负相关,干扰率>30%的样方中各指标均低于未受干扰样方。其中,土壤容重是对放牧干扰比较敏感的指标,与地衣结皮盖度亦呈极显著正相关（P<0.01）,与流沙盖度呈显著负相关（P<0.05）。     

荒漠结皮对土壤水分状况的影响

本研究针对鄂尔多斯沙地生物结皮进行调查 ,利用人工喷水模拟降雨分析结皮对土壤入渗性能的影响。研究表明 :生物结皮能显著地降低土壤水分的入渗速率。在去掉结皮时 ,表层下 5 cm处的土壤含水量在 1 0分钟时即开始显著增加 ,而有结皮时 ,此处的含水量则在 1 5分钟时才开始显著增加。利用圆盘入渗仪测定有结皮和无结皮条件下的土壤饱和导水率表明 :固定沙丘间地有生物结皮的土壤饱和导水率范围是 :2 9.1 0 - 82 .2 1 mm/ h;半固定沙丘有微弱结皮时饱和导水率为 1 43.5 4 - 2 30 .2 5 ;去掉结皮后土壤的饱和导水率可显著上升数倍 ,无结皮的流沙的饱和导水率最高。     

沙坡头地区生物结皮的水文物理特点及其环境意义

包兰铁路中甘段,大规模生物防护体系取得了明显的生态环境效益。随着时间的展延环境诸要素发生变化,特别生物结皮的形成和发育,制约着生态防护体系的演变和发展。生物结皮是人工生态防护体系建立后,外来尘埃动态沉积,一年生植被的侵入,植物凋落物腐解,在微生物及其环境条件综合作用下的新生成物。生物结皮作为一新的环境因子介入人工生态防护体系之中,使沙面固结稳定,增强了抗风蚀的能力。同时它的形成和发育,增大了降水的无效蒸发,使降雨入渗浅,沙地水分恶化,导致深根性植物衰退,浅根性的半灌木和草本植物得到发展,使人工体系向半人工半自然体系变化和演变。     

毛乌素沙地南缘沙区水分入渗特征及其影响因素

水分入渗是土壤水分循环的重要环节。为探明毛乌素沙地生物结皮覆盖沙区水分入渗特征及其影响因素,采用双环法测定不同类型地表(裸沙、浅灰色藻类结皮、黑褐色藻类结皮和藓类结皮,依次用BS、LC、DC和MC表示)水分入渗速率及其土壤理化性质。结果表明:1与BS相比,LC、DC和MC样地初始入渗率分别减少了37.7%、59.2%和73.6%,稳定前平均入渗率分别减少了37.5%、62.2%和81.3%,稳定入渗率分别减少了13.6%、67.4%和78.9%,累计入渗量分别减少了25.8%、61.3%和78.6%。2不同类型地表达到稳定入渗所需的时间在3~8 min;相关性分析结果显示,水分入渗速率与砂粒和土壤容重呈正相关关系,与粉粒、黏粒、结皮厚度、结皮抗剪强度、结皮层容重和有机碳含量呈负相关关系。3主成分分析结果表明,黏粒、结皮抗剪强度和土壤容重是影响该地区水分入渗速率的主要因子。因此,生物结皮在沙丘表面的形成和发育过程中,通过改变土壤的理化性质,减缓了该地区水分入渗速率,增加了降水的径流流失风险。     

黄土地与沙地生物结皮的发育特征及其生态功能异同

基于课题组已有研究成果,选取陕北水蚀风蚀交错区内气候条件相同但土壤质地迥异的试验区,探讨黄土地和沙地生物结皮发育特征及其生态效应的异同。结果表明:(1)苔藓结皮是2种土地生物结皮的重要类型,其中,黄土地的优势藓种为尖叶对齿藓[Didymodon constrictus(Mitt.)Saito.]、真藓(Bryum argenteum Hedw.)、狭网真藓(B.algovicum Sendt.);沙地的优势藓种为黄色真藓(B.pallescens Scheich.)、弯叶真藓(B.recurvulum Mitt.)、银叶真藓(B.argenteum Hedw.)。沙地乔灌植物下生物结皮盖度(77.5%)、厚度(11.8 mm)及容重(1.9 g·cm~(-3))均高于黄土地生物结皮,而黄土地多年生草本植物下生物结皮抗剪强度(26.5 k Pa)高于沙地生物结皮,总体上,沙地生物结皮发育的更好。(2)黄土地的入渗增幅和0~200 cm剖面的平均土壤含水率增幅均显著高于沙地(P0.05),且在旱季和雨季其0~200 cm剖面的平均土壤含水率增幅分别比沙地高1.4%和1.9%。(3)两地生物结皮均表现出了较好的减蚀作用,其减蚀效率分别为81.0%和90.6%。     

黄土地表生物结皮对土壤贮水性能及水分入渗特征的影响

采用双环法对山西省偏关县3种结皮覆盖下(苔藓藻结皮、藻结皮和薄层藻结皮)土壤的贮水性能和渗透特征进行了对比研究.结果表明:不同结皮覆盖下土壤的贮水能力受结皮厚度和孔隙度状况的影响较大,0～10 cm土层饱和贮水量为502.69～525.80 t/hm2 ,滞留贮水量为169.71～198.29 t/hm2;初渗速率的变化范围为5. 19～11.10 mm/min,无结皮最高,苔藓藻结皮最低;稳渗速率变化范围为1.6 7～2.67 mm/min.采用的3种入渗模型(Kostiakov模型、Horton模型和Ph ilip模型)中Horton模型的拟合值更接近于实测值, 决定系数R2在0.98～0.99,更适用于描述本研究区具有生物结皮土壤的入渗特征.     

土壤微生物在生物结皮形成中的作用及生态学意义

在荒漠地区的固定、半固定沙地和流动沙地上,微生物、藻类植物及地衣、苔藓植物能参与并形成生物结皮,其在干旱和半干旱荒漠地区具有重要生态意义。其中土壤微生物因其极强的适应性和生存能力,在生物结皮形成各阶段都起着积极的作用。土壤微生物的生长和活性与土壤酶活性的变化具有良好的相关性,它的组成、分布和数量因沙丘类型和沙土层深度的不同而有所变化,也受到季节和植被的影响。土壤微生物的生理代谢活动及数量变化可以改变沙土表面理化性质,在土壤结皮的形成、植物营养的转化过程中发挥积极作用,为植物生长创造了良好的条件。     

几种工程措施对黄土区陡峭边坡植被盖度的影响及其机理

陡削以兰州新区的一段陡峭边坡作为试验地点,通过分析在植被重建初期经人工整地后形成的4种微地形(大圆形坑、小圆形坑、条形坑和原状坡面)的植被盖度特征及环境因子(土壤硬度和水分、地表和空气温度、坡位、无纺布)对植被盖度的影响,探寻能有效提高植被盖度的几种工程改造措施。研究表明:①铺设了无纺布坡面的植被盖度显著大于对照坡面;陡峭边坡不同坡位的植被盖度:下坡位>中坡位>上坡位(P<0.05),这与自然坡面植被盖度分布规律一致。②3种微地形的植被盖度均显著大于陡峭坡面,其中条形坑的土壤水分和植被盖度最大,最大盖度维持的时间最长。③微地形和坡面的植被盖度均与土壤水分呈显著正相关,与空气温度、地表温度均呈显著负相关;微地形的土壤硬度(3kg·cm^-2)适宜植物生长,大于该值会对植被生长产生抑制作用;铺设无纺布有利于促进上坡位的植被生长,使坡面植被分布更均匀。研究结果表明,在陡峭边坡坡度、坡位和温度等无法改变的情况下,选择对压实的坡面进行整地翻耕处理,增加长条坑的数量并铺设无纺布等有效的工程改造措施,能显著增加植被盖度,促进植被恢复。     

土壤吸附作用对土壤水分有效性的影响

围绕土壤结皮生物是否可以对夜间土壤吸附水输入产生响应展开研究。假定-1.5 MPa作为土壤水分有效性的临界值,通过观测降水后清晨和午后0~3 cm土壤含水量,分析土壤水分的有效性和空间异质性;并利用连续记录气象数据,建立表层土壤含水量与水汽压饱和差数量关系。结果表明:土壤吸附作用提高了土壤水分对土壤结皮生物的有效性,并延长了土壤结皮生物对降水脉冲的响应时段。在古尔班通古特沙漠东南部的不同部位,凝结吸附作用对沙面水分条件影响很大(特别是干旱季节),水分条件由好到坏依次为:沙垄西坡面、沙垄垄顶、沙垄东坡面和垄间低地。     

古尔班通古特沙漠生物结皮影响下土壤水分的日变化

对新疆古尔班通古特沙漠生物结皮影响下土壤水分的日变化特征进行了定量研究,结果表明:①无论是结皮覆盖区还是无结皮覆盖区,其土壤含水量随深度的变化均表现出明显的层次性,其中以距地表 20 cm处的土壤含水量最高.②结皮覆盖区距地表5 cm和10 cm处的土壤含水量显著高于无结皮覆盖区(LSD检验, p ＜0.05);而45 cm和60 cm处的土壤含水量则表现为结皮覆盖区极显著低于无结皮区(LSD检验,p＜0.05),说明生物结皮具有较强的保持土壤表层水分的能力.③结皮覆盖区与无结皮覆盖区土壤水分的日变化特征明显不同于距地表5 cm处,而其他各土层的变化趋势则较为一致.无结皮覆盖区距地表5 cm处土壤水分的日变化呈先上升后下降的趋势,而结皮覆盖区的变化则恰恰相反.     

Soil hydraulic conductivity as affected by vegetation restoration age on the Loess Plateau,China

The Loess Plateau of China has experienced extensive vegetation restoration in the past several decades,which leads to great changes in soil properties such as soil bulk,porosity,and organic matter with the vegetation restoration age.And these soil properties have great effect on the soil infiltration and soil hydraulic conductivity.However,the potential changes in soil hydraulic conductivity caused by vegetation restoration age have not been well understood.This study was conducted to investigate the changes in soil hydraulic conductivity under five grasslands with different vegetation restoration ages(3,10,18,28 and 37 years)compared to a slope farmland,and further to identify the factors responsible for these changes on the Loess Plateau of China.At each site,accumulative infiltration amount and soil hydraulic conductivity were determined using a disc permeameter with a water supply pressure of –20 mm.Soil properties were measured for analyzing their potential factors influencing soil hydraulic conductivity.The results showed that the soil bulk had no significant changes over the initial 20 years of restoration(P0.05);the total porosity,capillary porosity and field capacity decreased significantly in the grass land with 28 and 37 restoration ages compared to the slope farmland;accumulative infiltration amount and soil hydraulic conductivity were significantly enhanced after 18 years of vegetation restoration.However,accumulative infiltration amount and soil hydraulic conductivity fluctuated over the initial 10 years of restoration.The increase in soil hydraulic conductivity with vegetation restoration was closely related to the changes in soil texture and structure.Soil sand and clay contents were the most influential factors on soil hydraulic conductivity,followed by bulk density,soil porosity,root density and crust thickness.The Pearson correlation coefficients indicated that the soil hydraulic conductivity was affected by multiply factors.These results are helpful to understand the changes in hydrological and erosion processes response to vegetation succession on the Loess Plateau.     

Microbiotic Crust Influence on Unsaturated Hydraulic Conductivity

Microbiotic crusts occur extensively in rangeland soils . Developed by filaments of cyanobacteria and algae , and thalli of lichen and moss entanglement of soil particles , they create a physical discontinuity in the surface profile with greater concen trations of clay , silt , and potentially hydrophobic organic matter. These conditions potentially contribute to variability in soil hydrology of arid land and should be considered in the development of hydrologic and erosion models. However, there is limited manipulative research examining the functional relationships between soil and microbiotic crusts . We investigated the influence of cyanobacterial-dominated microbiotic crust on measured hydraulic conductivity (K) in a sandy loam soil at a southeastern Utah site . Using a tension infiltrometer , we determined K under three surface treatments: undisturbed , chemically killed (representing dead microphytes within the crust), and removed (scalped) microbiotic crusts. We applied treatments to spatially interspersed intact surface soils within shrub interspaces . Microbiotic crusts at this site and in this stage of successional development had no discernible influence on K. This finding supports results from research conducted in a variety of soils from sandy to silt dominated with a range of microbiotic development . Because this research was site and time specific , and because the role of microbiotic crusts in the environment continues to be debated , additional research is warranted to deter mine how stage of development of microbiotic crust influences soil hydrology .     

Effect of soil management on soil erosion on sloping farmland during crop growth stages under a large-scale rainfall simulation experiment

Soil erosion on farmland is a critical environmental issue and the main source of sediment in the Yellow River, China. Thus, great efforts have been made to reduce runoff and soil loss by restoring vegetation on abandoned farmland. However, few studies have investigated runoff and soil loss from sloping farmland during crop growth season. The objective of this study was to investigate the effects of soil management on runoff and soil loss on sloping farmland during crop growth season. We tested different soybean growth stages (i.e., seedling stage (R1), initial blossoming stage (R2), full flowering stage (R3), pod bearing stage (R4), and initial filling stage (R5)) and soil management practice (one plot applied hoeing tillage (HT) before each rainfall event, whereas the other received no treatment (NH)) by applying simulated rainfall at an intensity of 80 mm/h. Results showed that runoff and soil loss both decreased and infiltration amount increased in successive soybean growth stages under both treatments. Compared with NH plot, there was less runoff and higher infiltration amount from HT plot. However, soil loss from HT plot was larger than that from NH plot in R1-R3, but lower in R4 and R5. In the early growth stages, hoeing tillage was effective for reducing runoff and enhancing rainfall infiltration. By contrast, hoeing tillage enhanced soil and water conservation during the late growth stages. The total soil loss from HT plot (509.0 g/m²) was 11.1% higher than that from NH plot (457.9 g/m²) in R1-R5. However, the infiltration amount from HT plot (313.9 mm) was 18.4% higher than that from NH plot (265.0 mm) and the total runoff volume from HT plot was 49.7% less than that from NH plot. These results indicated that crop vegetation can also act as a type of vegetation cover and play an important role on sloping farmland. Thus, adopting rational soil management in crop planting on sloping farmland can effectively reduce runoff and soil loss, as well as maximize rainwater infiltration during crop growth period.     

一株葡萄孢属(Botrytis)真菌结皮效果的研究

从陕北毛乌素沙地生物结皮的下层中筛选得到一株真菌,经鉴定,表明该菌株属于葡萄孢属(Botry-tis)。它能在生长繁殖过程中分泌出大量胞外黏多糖,室内摇瓶振荡培养3 d黏度可达9 860 mPa.s,产量达19.246g/L,通过采用薄层层析法对多糖的组分的测定,初步表明该多糖主要由D-甘露糖和D-半乳糖组成。通过结皮试验,结果表明该株真菌菌剂喷洒于流沙表面后,能够形成约8.2 mm厚的一层具有粘结沙粒、保持水分的生物结皮层,同时具有明显的减缓土壤中水分蒸发的效果。表明该菌株在荒漠化治理、恢复和保护生态环境方面具有重要的应用潜力。     

不同沙丘部位和不同结皮类型对土壤种子库的影响

采用萌发法对古尔班通古特沙漠沙垄不同部位生物结皮类型的种子库进行研究,发现不同沙丘部位和结皮类型种子分布特征有一定的规律性:藻类结皮的种子数量、丰富度和多样性均显著高于地衣和苔藓结皮。生物结皮从藻类到地衣和苔藓的演替,有降低种子数量和种子多样性的现象。在不同沙丘部位,沙丘顶部流沙的种子数量和多样性较低,与丘顶流沙相比沙丘底部的藻类结皮显示出较高的丰富度和多样性,但沙丘底部的地衣和苔藓结皮与丘顶流沙相比未表现出明显的差异。背风坡中部的藻类结皮种子多样性较高,显著高于迎风坡中部的藻类结皮。     

坡耕地暴雨结皮对作物生长发育影响的实验研究

采用盆栽和小区试验的方法,经3年试验发现,坡耕地土壤结皮对作物的出苗、生长、产量和生物量等均有不同程度的影响。小麦出苗率、苗高、最大苗高、产量和生物量分别降低了22.06％、4.60％、6.86％、1.16-9.84％和0.74-6.98％;结皮虽对玉米出苗影响不大,但延长了出苗期,使产量和生物量降低了5.66—9.17％和7.17—11.25％,而且随结皮厚度、强度和地表坡度的增加,其影响更为明显。因此,通过增加地面覆盖和雨后人工耕锄以消除结皮,增加产量和保持水土。