Sward management, lamina turnover and tiller population density in continuously stocked Lolium perenne-dominated swards

Measurements of rates of growth and senescence of leaf lamina per tiller and of changes in tiller population densities were made in three experiments designed to investigate the influence of sward slate on leaf turnover and net production under continuous stocking. In each experiment initially uniform swards were fenced to provide four plots on which animal numbers were adjusted twice weekly to give a series of swards maintained as nearly as possible in a steady state with respect to sward surface height (range 1.1–6.4 cm) and herbage mass (range 440–2690 kg OM ha^?1). Two experiments were carried out in July–September on vegetative swards and one in May–June on a reproductive sward. Measurements were begun 3–7 weeks after treatments were started and were repeated weekly during 3–4 week measurement periods. In all three experiments the rate of lamina growth per tiller increased linearly with an increase in sward surface height and herbage mass. In the two experiments conducted in July–August this relationship was partially offset by a linear increase in the rate of senescence per tiller but net production per tiller also increased linearly in relation to sward height and mass. In the experiment conducted in May–June the rates of growth and senescence per tiller increased in parallel so that net production per tiller showed no relationship with sward condition. Tiller population densities in the July–August experiments were highest in swards maintained between 2 and 3 cm surface height and declined in swards maintained above and below this height. In the experiment in May–June tiller numbers were similar in all treatments prior to the summer solstice but diverged in a manner similar to the other experiments later in the year. The rate of lamina growth per unit area increased in a manner consistent with an asymptotic relationship and the rate of senescence increased linearly with increasing sward height and mass in all three experiments. Net production per unit area was reduced on swards below about 2.5 cm in height but was insensitive to variation in sward surface height between 2.5 and 6.0 cm (approximately 1000–2500 kg OM ha^?1 herbage mass). The effectiveness of adjustments in tiller numbers and production per tiller and of changes in the balance between growth and senescence as mechanisms of sward homeostatis, together with their implications in the scope for manipulating herbage production by grazing management are discussed briefly.     

Herbage intake and liveweight gain of bulls and steers continuously stocked at fixed sward heights during autumn and spring

Two experiments were conducted to examine the relationship between sward surface height, herbage intake and liveweight gain in beef cattle grazed on pasture. In Experiment 1, two 'animal types' (18 Charolais × Angus steers and 18 Friesian bulls) were continuously grazed for 22 days during the late autumn on replicated swards maintained at sward surface heights of 6, 10 and 15 cm. Herbage intakes, assessed from the faecal concentration of chromium delivered from an intraruminal controlled release capsule and the in vitro digestibility of hand-plucked herbage samples, were curvilinearly related to sward height (r = 0·76, p <0·0·01). Average liveweight gains were 0·02, 0·61 and 1·31 kg d^-1 ( P <0·05) and increased linearly ( r = 0·84, P <0·001) with sward height. The maintenance organic matter intakes of the steers and bulls, with initial mean (± s.e.) live weights of 225 ± 15 kg and 172 ± 15 kg respectively, were estimated to be 3·6 and 3·5 kg d^-1 respectively. In Experiment 2 (spring) 36 cattle, including 35 of those used in Experiment 1, were reallocated to sward heights of 5, 10 and 15 cm using the same design as for Experiment 1. Average liveweight gains were 0·94, 1·57 and 1·68kg d^-1 ( p 0·05) and were curvilinearly related to sward height ( r = 0·093, p <0·05). Maintenance intakes could not be reliably extimated for the cattle in Experiment 2 because few animals had liveweight gains close to zero. These trials confirm that liveweight gain in continuously grazed finishing steers and bulls increases with sward surface height to maximum of 8–10cm with spring ryegrass/white clover pastures while, in autumn, swards of 12–15cm height are required to achieve maximum performance.     

A comparison of spectral reflectance and sward surface height measurements to estimate herbage mass and leaf area index in continuously stocked ryegrass pastures

Measurements of sward surface height and of the ratio of light at 660 and 730 nm reflected from a pasture canopy were correlated with measurements of leaf area index (LAI) and herbage mass of two ryegrass dominant swards. Both pastures were continuously stocked by sheep to maintain a range of sward heights from 2 to 6 cm corresponding approximately to LAI 2 to 5.
Sward height appeared to be linearly related to both LAI and herbage mass, whereas 660/730 reflectance displayed a non-linear relationship with both parameters. The accuracy of prediction by the two methods over the range LAI 1 to 3 or herbage mass 700 to 1800 kg DM ha^-1 was very similar. However, reflectance measurements could not be used above about LAI 3–4 and to this extent were less useful. Nevertheless, reflectance measurements have a potential advantage in their ability to sample large areas of pasture very quickly.     

The effect of surface height of swards continuously stocked with cattle on herbage production and clover content over four years

The long-term influence of sward height from April to July (Phase 1) and from July to early October (Phase 2) on total herbage and white clover production was measured over four years (1988–1991) as herbage accumulation. A subsidiary experiment to determine the influence of leaf area index (LAI) on gross canopy photosynthesis was conducted to aid interpretation of growth from herbage accumulation data. Target sward heights in 0·5 ha plots in two blocks were 5,7 or 9 cm in Phase 1 and 7 or 9 cm in Phase 2, although mean actual heights per phase were slightly higher. Net herbage accumulation (NHA) was measured within mobile exclosure areas over successive two-week intervals. Gross photosynthesis was measured in circular turves removed from the trial area representing a range of LAIs with an at least reasonable clover content. Despite wide differences in mean sward height and herbage mass, NHA and net clover accumulation for a given phase were not generally affected by treatments. Positive effects of grazing at 5 cm in Phase 1 on NHA and clover accumulation later in the year, and of grazing at 7 cm in Phase 2 on NHA in the following spring were sometimes apparent. Gross canopy photosynthesis (g CO₂ m^?2 h^?1) at 1500 μE m^?2 s^?1 and 18–21°C was linearly related to LAI described by 1·003 + 1·165 LAI over the LAI range 0·7 to 4·5. Total herbage and clover growth, interpreted from NHA by a previously described model, was predicted to be marginally lower in shorter swards. Similarity in NHA and clover accumulation between treatments was considered to be because of lower senescene and decomposition, and a higher proportion of new tissue being assigned to lamina growth, despite lower LAI and gross photosynthesis in the shorter swards. It was concluded that stocking intensity in swards continuously stocked with cattle did not have a strong influence on net total herbage and clover growth.     

Effects of sward surface height on the performance of continuously stocked spring-calving beef cows and their calves

The response of continuously stocked spring-calving beef cows and calves to a range of sward surface heights was determined. Mean sward heights of 4-5, 60, 70, 91 and 11-0 cm (treatments A to E respectively) were maintained from mid-May to mid-August (Period 1) and treatments C, D and E continued until the end of September (Period 2), during which time mean sward heights were 5-4, 7-8 and 9-2 cm respectively. Hereford x Friesian or White Shorthorn-x Galloway cows with their Charolais-cross calves were used, with 9, 6, 6, 5 and 9 cows on treatments A to E respectively. During Period 1 the mean live weight gains of cows were-0-52,-005,0-32,075 and 0 40 (s.e.d. 0167) kg d⁻¹ on treatments A to E respectively and during Period 2.-0-31, 057 and 059 (s.e.d. 0153) kg d⁻¹ for treatments C, D and E respectively with maximum cow liveweight gain at sward heights of 8 to 10 cm. Calf liveweight gains were 0 88,091, 098, 1 04 and 1-06 (s.e.d. 0068) kg d⁻¹ in Period 1 for treatments A to E respectively and 098, 1-22 and 1 35 (s.e.d. 0067) kg d⁻¹ in Period 2 for treatments C, D and E. The percentage of area infrequently grazed was generally less than 20% on treatments A to D, but on treatment E it was more than 40% for 6 weeks after turnout and thereafter remained between 20 and 40%. Maximum calf liveweight gain per hectare was achieved on the shorter swards, but maximum total liveweight gain of cows and calves per hectare occurred on treatment D. It is concluded that for maximum cow and calf performance on continuously stocked pastures, sward height should be maintained at no more than 8 cm in spring and early summer and then increased to 9 to 10 cm later in the grazing season.     

Herbage losses from tiller pulling in a continuously grazed perennial ryegrass sward

Tiller pulling was studied in a perennial ryegrass sward that was continuously grazed by cattle. The treatments included severe (sward height after grazing 25 mm), medium (50 mm) or lenient (75 mm) grazing from turn-out in April to 1 June, followed by grazing to 50 mm in the remainder of the season. Tiller pulling was confined to the midsummer-autumn period of the grazing season. The losses were most severe in swards that had been leniently grazed to a mean height of 75 mm during the spring and least severe in swards grazed to a height of 25 mm. The lenient grazing treatment allowed both the true stem development and aerial tillering whereas in the more tightly grazed swards true stem development was significantly less and aerial tillering was virtually absent. The pulled organic matter in the lenient treatment was equivalent to about 5·3% of the total harvested yield. In the severely grazed swards, pulling losses were equivalent to about 1·7% of the total harvested yield. A high rate of turnover of the pulled herbage was found in all the treatments with between 69 and 78% of the freshly pulled herbage disappearing within a week of being pulled. Tiller pulling was found to have no effect on either the subsequent autumn-winter tiller density or yields of cuts taken in the following year.     

Quantifying the influence of sward height, concentrate level and initial milk yield on the milk production and grazing behaviour of continuously stocked dairy cows

Two factorial design experiments were carried out in the spring of 1994 and 1995, each of 6 weeks, to quantify the effects of sward height (SH), concentrate level (CL) and initial milk yield (IMY) on milk production and grazing behaviour of continuously stocked dairy cows. In Experiment 1, forty‐five Holstein Friesian cows were in five groups with initial milk yields of 16·9, 21·1, 28·0, 31·5 and 35·5 kg d^–1, grazed sward heights were 3–5, 5–7 and 7–9 cm (LSH, MSH and HSH respectively), and concentrates were fed at rates of 0, 3 and 6 kg d^–1. In Experiment 2, 48 cows were in two groups with IMY of 21·3 and 35·5 kg d^–1, grazed sward heights were 3–5 and 7–9 cm (LSH and HSH), and concentrates were fed at 0 and 6 kg d^–1 and ad libitum. Multiple regression models were used to quantify the effects of the three variables on milk yield persistency (MYP), estimated herbage dry‐matter (DM) intake (HDMI), grazing time (GT) and rate of DM intake (RI). The partial regression coefficients showed that increased SH led to increased MYP (Experiment 1 P < 0·001, Experiment 2 P < 0·05), increased HDMI (P < 0·01, P < 0·01), increased GT (P < 0·001, P < 0·05) and increased RI (P < 0·001, P < 0·05). Increasing CL led to increased MYP (NS, P < 0·001), decreased HDMI (P < 0·001, P < 0·001), decreased GT (NS, P < 0·001) and decreased RI (P < 0·001, P < 0·001). Higher IMY level of cows decreased MYP (P < 0·001, P < 0·001), increased HDMI (P < 0·001, P < 0·001), increased GT (P < 0·001, P < 0·05) and increased RI (P < 0·05, P < 0·01). The models were highly significant (P < 0·001), and accounted for 0·48–0·87 of the total variance. The partial regression coefficients quantified the extent to which GT and RI by cows respond positively to higher IMY, and negatively to increased CL, but respond differently (GT declines in response to a higher RI) with increasing SH.     

GrazeIn: a model of herbage intake and milk production for grazing dairy cows. 2. Prediction of intake under rotational and continuously stocked grazing management

Decision support tools to help dairy farmers gain confidence in grazing management need to be able to predict performance of grazing animals with easy‐to‐obtain variables on farm. This paper, the second of a series of three, describes the GrazeIn model predicting herbage intake for grazing dairy cows. The model of voluntary intake described in the first paper is adapted to grazing situations taking account of sward characteristics and grazing management, which can potentially affect intake compared to indoor feeding. Rotational and continuously stocked grazing systems are considered separately. Specific effects of grazing management on intake were quantified from an extensive literature review, including the effect of daily herbage allowance and pre‐grazing herbage mass in rotational grazing systems, sward surface height in continuously stocked grazing systems, and daily time at pasture in both grazing systems. The model, based on iterative procedures, estimates many interactions between cows, supplements, sward characteristics and grazing management. The sensitivity of the prediction of herbage intake to sward and management characteristics, as well as the robustness of the simulations and an external validation of the GrazeIn model with an independent data set, is presented in a third paper.     

Changes in the structure and physiology of a perennial ryegrass sward when released from a continuous stocking management: implications for the use of exclusion cages in continuously stocked swards

A perennial ryegrass sward was managed by continuous stocking with sheep (April–September) for 4 successive years after sowing. The sward was grazed to maintain a leaf (lamina) area index (LAI) close to 1.0. Areas of the sward were released from grazing on three occasions: once during summer in the third year after sowing, and twice during spring and summer in the fourth year after sowing. There were marked changes in the structure and physiology of the continuously stocked sward following release from grazing. After several successive years of continuous stocking, the sward comprised a large population of small tillers and the small LAI resulted in consistently low rates of photosynthesis. Following release from grazing, photosynthesis increased markedly as the LAI increased but this change was associated with the loss of a large proportion of the population of tillers. There were seasonal differences in the pattern of changes in photosynthesis and tiller numbers following release from grazing which were not apparent under continuous stocking. The changes in the structure and physiology of the sward following release from grazing suggest that the net accumulation of herbage in areas of sward from which the animals are excluded, for instance using cages, may be an unreliable estimate of production under continuous stocking.     

The effects of autumn closing date on sward leaf area index and herbage mass during the winter period

The provision of grass for early spring grazing in Ireland is critical for spring calving grass‐based milk production systems. This experiment investigated the effect of a range of autumn closing dates (CD), on herbage mass (kg DM ha^?1), leaf area index (LAI) and tiller density (m^?2) during winter and early spring. Thirty‐six grazing paddocks, closed from 23 September to 1 December 2007, were grouped to create five mean CD treatments – 29 September, 13 October, 27 October, 10 November, 24 November. Herbage mass, tiller density and LAI were measured every 3 weeks from 28 November 2007 to 20 February 2008; additionally, herbage mass was measured prior to initial spring grazing and tiller density was measured intermittently until September 2008. Delaying CD until November significantly (P < 0·05) reduced herbage mass (by approximately 500 kg DM ha^?1) and LAI (by approximately 0·86 units) in mid‐February. On average, 35% of herbage mass present on swards on 20 February was grown between 28 November and 30 January. LAI was positively correlated with herbage mass (R² = 0·78). Herbage mass increased by approximately 1000 kg DM ha^?1 as spring grazing was delayed from February to April. Tiller density increased from November to February, although it did fluctuate, and it was greatest in April (9930 m^?2). This experiment concludes that in the south of Ireland adequate herbage mass for grazing in early spring can be achieved by delaying closing to early mid‐October; swards required for grazing after mid‐March can be closed during November.     

Impact of spatial heterogeneity of plant species on herbage productivity,herbage quality and ewe and lamb performance of continuously stocked,perennial ryegrass–white clover swards

The benefits of white clover (Trifolium repens L.) in pastures are widely recognized. However, white clover is perceived as being unreliable due to its typically low content and spatial and temporal variability in mixed (grass‐legume) pastures. One solution to increase the clover proportion and quality of herbage available to grazing animals may be to spatially separate clover from grass within the same field. In a field experiment, perennial ryegrass (Lolium perenne L.) and white clover were sown as a mixture and compared with alternating strips of ryegrass and clover (at 1·5 and 3 m widths), or in adjacent monocultures (strips of 18 m width within a 36‐m‐wide field). Pastures were stocked by ewes and lambs for three 10‐month grazing periods. Over the 3 years of the experiment, spatial separation of grass and clover, compared with a grass–clover mixture, increased clover herbage production, although its proportion in the sward declined through time (0·49–0·54 vs 0·34 in the mixture in the first year, 0·28–0·33 vs 0·15 in the second year and 0·03–0·18 vs 0·01 in the third year). Total herbage production in the growing season in the spatially separated treatments decreased from 11384 kg DM ha^?1 in the first year to 8150 kg DM ha^?1 in the third year. Crude protein concentration of clover and grass components in the 18‐m adjacent monoculture treatment was greater than the mixture treatment for both clover (310 vs 280 g kg^?1 DM) and grass (200 vs 180 g kg^?1 DM). There was no clear benefit in liveweight gain beyond the first year in response to spatially separating grass and clover into monocultures within the same field.     

Effects of grazing by lambs in autumn on a red clover-perennial ryegrass sward

In 1976, 1977 and 1978 a red clover-perennial ryegrass sward was cut twice for silage, and in the autumn of 1976 and 1977 it was either grazed at low and high stocking rates, i.e. seventeen and thirty-four lambs per ha respectively, or was cut with a forage harvester. The effects of grazing on yield in the following year were examined. Herbage growth in the grazing period was slow and did not differ significantly between the treatments. The yield of silage dry matter taken in May and July was highest in ungrazed plots (9·8 and 8·1 t ha^-1 in 1977 and 1978 respectively) and lowest in plots stocked at the high rate (5·9 and 5·7 t ha^-1 in 1977 and 1978 respectively). The aftermath yield for grazing in 1977 was slightly but significantly greater on treatments grazed in the previous year compared with ungrazed treatments. Red clover content decreased markedly during grazing, the high stocking rate treatment containing 2·4% clover and the ungrazed treatment 57·3% clover. The high stocking rate treatment also had the lowest red clover content in the first silage cut. Red clover content in grazed plots increased to a level similar to that in ungrazed plots by the start of the grazing period in the subsequent harvest year. Animal performance was higher at the low than at the high stocking rate but herbage consumption per head did not differ significantly between the two grazing treatments. Possible reasons for the adverse effect of grazing on the red clover are defoliation and treading. It is concluded that such experiments can form the basis of an economic assessment of red clover and help the farmer decide whether or not he should integrate the crop into his system.     

Evaluation of a sward sampling technique for estimating herbage intake by grazing sheep

A sampling procedure for estimating herbage intake by the ‘Difference’ method over short grazing periods using an electrically powered sheep-shearing head to cut long narrow sample strips, was compared with several indirect animal techniques on single-variety grass swards. The method provided intake estimates in good general agreement with those obtained by animal methods but discrepancies arose when used on a semi-prostrate variety due, probably, to incomplete recovery of sampled herbage cut from post-grazing residues. It was concluded that the method might be usefully adopted for the preliminary assessment of the intake characteristics of herbage varieties in routine evaluation programmes, provided that care is taken to ensure the efficient collection of herbage sampled post-grazing and that a reduction in the land requirement can be achieved without undue loss of accuracy or precision. Confirmation of results should be sought using animal methods when possible.     

The effects of sward characteristics and supplement type on the herbage intake and milk production of summer-calving cows

High (H; 27 350 m^?1:) or low (L. 13 300 m-¹) tiller density perennial ryegrass swards were created in the mid- and late grazing season by imposing different sward heights in the spring. Summer-calving cows then grazed these swards from 6 June to 2 September 1992 and were offered 5 kg fresh weight hd^?1 d^?1 of either a barley (S) or a molassed sugar beet pulp (F) based supplement. The factorial combination of sward and supplement types resulted in four experimental swards being grazed by thirteen Holstein/Friesian cows each. Supplement F contained more crude fibre (110 vs. 58 g kg^?1) and less metabolizable energy [12–5 vs. 13–2 MJ kg^?1 dry matter (DM)] than supplement S. Herbage on the H sward contained more metabolizable energy (11–9 vs. 104 MJ kg^?1 DM) and crude protein (232 vs. 205 g kg^?1 DM), had fewer rejected areas f 16–5 vs. 26–9%) and a higher live-dead tiller ratio (4–6 vs. 2–1) than that on the L sward. Sward, but not supplement type, significantly affected the intake of grazed herbage (P<0–001). On average, the herbage intakes of cows grazing II swards were higher than for L swards (14–5 vs. 11 6kg DM d^?1) and those of cows on the S and F supplements were 12–6 and 13–5 kg DM d^?1 respectively. Averaged over the grazing period, sward and supplement had no significant effects on milk yield, milk composition or yield of constituents. When expressed on an average weekly basis, cows grazing an L sward and offered the F supplement on occasions had significantly lower milk yields and higher milk fat contents (P<005) than those grazing an H sward and offered the S supplement. There were no significant effects on cow live weight or condition score change. The results suggested that grazing swards with a high density of live tillers increased herbage intakes and on occasions milk yield, relative to low density swards. However, small increases in energy intake from sward and supplement effects were used primarily to ameliorate liveweight loss.     

Effect of early season sward management on sward quality and lamb liveweight gain during autumn

In two experiments (1990 and 1991) perennial ryegrass (cv. Talbot) pasture was subjected to six different management treatments by adjusting stock numbers to achieve set sward heights for set periods between turn-out in early April and weaning in early July. The main objective of the experiments was to measure the effect of differences in sward management up to July on subsequent sward type, and the effect of sward type on the performance of weaned lambs between mid-July and early October when all swards were grazed at a sward height of 7 cm. In Experiment 1 swards were grazed at 7 cm initially and then reduced to 3 cm at various dates and maintained at 3 cm until early July. In Experiment 2 swards were grazed initially at 7 cm, and this was reduced to 3 cm and then allowed to return to 7 cm at various dates up to early July. Grazing at a sward height of 7 cm up to weaning, in early July, gave mean twin lamb growth rates of 310 g d^?1, while at a tower sward surface height of 3 cm lamb growth rate was reduced to 206 g d^?1 The density of seed heads produced from July was strongly influenced by earlier management. Grazing at 7 cm up to July gave a high density of seed heads (213m^?2), and this reduced (126 m^?2) by grazing to 3 cm. Seed heads were almost eliminated (17 m^?2) by grazing at 7 cm until late June followed by grazing down to 3 cm in early July. Reducing sward surface height from 7 cm to 3 cm at an earlier date tended to increase the development of seed heads from early July. Where swards were grazed from 7 cm to 3 cm and then allowed to return to 7 cm, the later in the pre-weaning period this took place the greater the reduction in seed head development. Greater seed head density was associated with a significant decrease in the nitrogen content of the herbage on offer in Experiment 2, but an increase in modified acid detergent (MAD) fibre content was not significant. Despite the differences in the sward type on offer, the effect on weaned lamb growth rate was small. Lamb liveweight gain during autumn was significantly reduced in Experiment 2 following earlier grazing at 7 cm compared with 3 cm, but liveweight gain differences in Experiment 1 were not significant.     

Effects of perennial ryegrass (Lolium perenne L.) cultivars on herbage production, nutritional quality and herbage intake of grazing dairy cows

Four perennial ryegrass (Lolium perenne L.) cultivars were compared for differences in herbage production, nutritive value and herbage intake of dry matter (DM) during the summers of 2002 and 2003. Two paddocks were sown with pure stands of four cultivars in a randomized block design with three replicates. Each plot was subdivided into fourteen subplots (22 m × 6 m) which were grazed by one cow during 24 h. Twelve lactating dairy cows were assigned to one cultivar for a period of 2 weeks in a 4 × 4 Latin square experimental design; the experiment lasted 8 weeks in each year. Sward structure (sward surface height, DM yield, green leaf mass, bulk density and tiller density) and morphological characteristics were measured. The ash, neutral‐detergent fibre, acid‐detergent lignin, crude protein and water‐soluble carbohydrate concentrations, and in vitro digestibility of the herbage were measured. The sward was also examined for infestation by crown rust (Puccinia coronata f. sp. lolii). Herbage intake of dairy cows was estimated using the n‐alkane technique. Cultivar differences for all sward structural characteristics were found except for bulk density and tiller density in 2003. Cultivars differed for proportions of pseudostem, stem (in 2003 only) and dead material. The chemical composition of the herbage was different among cultivars, with the water‐soluble carbohydrate concentration showing large variation (>0·35). Cultivars differed in susceptibility to crown rust. Herbage intake differed among cultivars in 2002 (>2 kg DM) but not in 2003. Herbage intake was positively associated with sward height, DM yield and green leaf mass. Canopy morphology did not affect herbage intake. Crown rust affected herbage intake negatively. It was concluded that options for breeders to select for higher intake were limited. High‐yielding cultivars and cultivars highly resistant to crown rust were positively related with a high herbage intake.     

Effect of grazing severity on perennial ryegrass herbage production and sward structural characteristics throughout an entire grazing season

The objective of this study, which was part of a larger grazing‐systems experiment, was to investigate the cumulative impact of three levels of grazing intensity on sward production, utilization and structural characteristics. Pastures were grazed by rotational stocking with Holstein–Friesian dairy cows from 10 February to 18 November 2009. Target post‐grazing heights were 4·5 to 5 cm (high; H), 4 to 4·5 cm (intermediate; I) and 3·5 to 4 cm (low; L). Detailed sward measurement were undertaken on 0·08 of each farmlet area. There were no significant treatment differences in herbage accumulated or in herbage harvested [mean 11·3 and 11·2 t dry matter (DM) ha^?1 respectively]. Above the 3·5 cm horizon, H, I and L swards had 0·56, 0·62 and 0·67 of DM as leaf and 0·30, 0·23 and 0·21 of DM as stem respectively. As grazing severity increased, tiller density of grass species other than perennial ryegrass (PRG) decreased (from 3,350 to 2,780 and to 1771 tillers m^?2 for H, I and L paddocks respectively) and the rejected area decreased (from 0·27 to 0·20 and to 0·10 for H, I and L paddocks respectively). These results indicate the importance of grazing management practice on sward structure and quality and endorse the concept of increased grazing severity as a strategy to maintain high‐quality grass throughout the grazing season. The findings are presented in the context of the need for intensive dairy production systems to provide greater quantities of high‐quality pasture over an extended grazing season, in response to policy changes with the abolition of EU milk quotas.     

The effect of the nitrification inhibitor dicyandiamide (DCD) on spring and annual herbage production in urine patches when applied in late summer or early autumn

A cut plot experiment was undertaken at two sites in Ireland, one a free‐draining acid brown earth at Moorepark (MPK) and the other a fine loam soil with imperfect drainage at Johnstown Castle (JC). The effect of applying the nitrification inhibitor dicyandiamide (DCD) at 10 kg ha^?1 in July, August and September or not applying DCD to plots receiving synthetic urine or zero urine on spring and annual herbage production was examined. In the experiment, each site received 350 kg nitrogen (N) fertilizer ha^?1 year^?1. The application of DCD in August at a rate of 10 kg ha^?1 significantly increased spring and annual herbage production by 14 and 15%, respectively, at MPK, when applied following urine application in year 1. There was no effect of DCD applied in year 1 on herbage production at JC. The application of DCD in August resulted in lower soil total oxidized N (TON) content up to sampling day 56 post‐urine application, at MPK in year 1, retaining higher N content in the soil. There was no effect of DCD on any of the parameters measured in year 2 at MPK or at JC. Urine application did not increase spring herbage production at either site. Urine application significantly increased annual herbage production at MPK only in year 1. Urine application increased annual herbage N uptake, herbage crude protein (CP) content and soil mineral N at both sites in both years.     

Effect of nitrogen fertilization on herbage production of tall fescue swards continuously grazed by sheep. 2. Consumption and efficiency of herbage utilization

The effect of a reduced level of nitrogen fertilizer application upon the herbage consumption of a continuously grazed sward of tall fescue has been investigated during two successive years. Two contrasting cultivars were used and two levels of leaf area index were maintained on the swards in order to obtain a wide range of sward structure. The herbage growth, herbage senescence and herbage consumption were measured directly on labelled tillers. It was possible to estimate the different components of the defoliation process (frequency and intensity) at the level of individual tillers of each leaf category. The results show that the effect of reduction in N fertilizer on herbage consumption rate is proportionally more important than that on herbage growth rate. In consequence, the efficiency of herbage utilization in continuously grazed conditions is reduced when nitrogen nutrition becomes limiting for herbage growth. The data suggest that this phenomenon is not directly attributable to nitrogen deficiency but is an indirect effect owing to the reduction of the stocking density at the low N level, which leads to a lower frequency of defoliation of individual tillers.     

Effects of date of autumn closing and timing of winter grazing on herbage production in winter and spring

Growth of grass herbage in Ireland is highly seasonal with little or no net growth from November to February. As a result, feed demand exceeds grass supply during late autumn, winter and early spring. At low stocking rates [≤2 livestock units (LU) ha^?1], there is potential to defer some of the herbage grown in autumn to support winter grazing. This study examined the effects of four autumn‐closing dates and four winter‐grazing dates in successive years on the accumulation of herbage mass and on tiller density in winter and subsequent herbage production at two sites in Ireland, one in the south and one in the north‐east. Closing swards from grazing in early and mid‐September (north‐east and south of Ireland respectively) provided swards with >2 t DM ha^?1 and a proportion of green leaf >0·65–0·70 of the herbage mass above 4 cm, with a crude protein (CP) concentration of >230 g kg^?1 DM and dry matter digestibility (DMD) of >0·700. The effects of autumn‐closing date and winter‐grazing date on herbage production in the subsequent year varied between the two sites. There was no significant effect of autumn‐closing date in the north‐eastern site whereas in the south earlier autumn closing reduced the herbage mass in late March by up to 0·34 t DM ha^?1 and delaying winter grazing reduced the herbage mass in late March by up to 0·85 t DM ha^?1. The effects of later grazing dates in winter on herbage mass continued into the summer at the southern site, reducing the herbage mass for the period from late March to July by up to 2 t DM ha^?1. The effects of imposing treatments in successive years did not follow a consistent pattern and year‐to‐year variation was most likely linked to meteorological conditions.