Variation in vegetation composition,biomass production,and carbon storage in ridge top forests of high mountains of Garhwal Himalaya

Abstract

The present study was aimed to anticipate how forest composition, regeneration, biomass production, and carbon storage vary in the ridge top forests of the high mountains of Garhwal Himalaya. For this purpose five major forest types—(a) Pinus wallichiana, (b) Quercus semecarpifolia, (c) Cedrus deodara, (d) Abies spectabilis, and (e) Betula utilis mixed forests—were selected on different ridge tops in the Bhagirathi Catchment Area of the Uttarkashi District of Garhwal Himalaya. The highest species richness (10 species) and stand density (804 ± 184.5 stems ha^?1) were recorded in Abies spectabilis forests, whereas lowest species richness (4 species) and species density (428 ± 144.7 stems ha^?1) were found in Quercus semecarpifolia forests. The total basal cover (TBC) values were maximum (91.1 ± 24.4 m² ha^?1) in Cedrus deodara forests and minimum (26.5 ± 11.7 m² ha^?1) in Pinus wallichiana forests. The highest total biomass density (TBD) (464.2 ± 152.5 Mg ha^?1) and total carbon density (TCD; 208.9 ± 68.6 Mg C ha^?1) values were recorded for Cedrus deodara forests; however, lowest TBD (283.4 ± 74.8 Mg ha^?1) and TCD (127.5 ± 33.7 Mg C ha^?1) values for Quercus semecarpifolia forests. Our study suggests that Abies spectabilis-dominated forests should be encouraged for biodiversity enrichment and reducing carbon emissions on ridge top forests of high mountains.     

Aboveground biomass and carbon stock in the largest sacred grove of Manipur,Northeast India

Aboveground biomass and carbon stock in the largest sacred grove of Manipur was estimated for trees with diameter [10 cm at 1.37 m height.The aboveground biomass,carbon stock,tree density and basal area of the sacred grove ranged from 962.94 to 1130.79 Mg ha~(-1),481.47 to 565.40 Mg ha~(-1) C,1240 to 1320 stem ha~(-1) and79.43 to 90.64 m~2 ha~(-1),respectively.Trees in diameter class of 30–40 cm contributed the highest proportion of aboveground biomass(22.50–33.73%).The aboveground biomass and carbon stock in research area were higher than reported for many tropical and temperate forests,suggesting a role of spiritual forest conservation for carbon sink management.     

Carbon sequestration in Chir-Pine （Pinus roxburghii Sarg.） forests under various disturbance levels in Kumaun Central Himalaya

We studied variations in tree biomass and carbon sequestration rates of Chir Pine(Pinus roxburghii. Sarg.) forest in three categories of forest disturbance, protected, moderately disturbed, and highly disturbed. In the first year, total biomass was 14.7 t?ha-1 in highly disturbed site, 94.46 t?ha-1 in moderately disturbed forest, and 112.0 t?ha-1 in protected forest. The soil organic carbon in the top 20 cm of soil ranged from 0.63 to 1.2%. The total rate of carbon sequestration was 0.60(t/ha)·a-1on the highly disturbed site, 1.03(t/ha)·a-1 on the moderately disturbed site, and 4.3(t/ha)·a-1 on the protected site.     

Carbon storage in biomass,litter, and soil of different native and introduced fast-growing tree plantations in the South Caspian Sea

Replantation of degraded forest using rapidgrowth trees can play a significant role in global carbon budget by storing large quantities of carbon in live biomass,forest floor,and soil organic matter.We assessed the potential of 20-year old stands of three rapid-growth tree species,including Alnus subcordata,Populus deltoides and Taxodium distichum,for carbon(C) storage at ecosystem level.In September 2013,48 replicate plots(16 m × 16 m) in 8 stands of three plantations were established.36 trees were felled down and fresh biomass of different components was weighed in the field.Biomass equations were fitted using data based on the 36 felled trees.The biomass of understory vegetation and litter were measured by harvesting all the components.The C fraction of understory,litter,and soil were measured.The ecosystem C storage was as follows: A.subcordata(626.5 Mg ha~(-1)) [ P.deltoides(542.9Mg ha~(-1)) [ T.distichum(486.8 Mg ha~(-1))(P \ 0.001),of which78.1–87.4% was in the soil.P.deltoides plantation reached the highest tree biomass(206.6 Mg ha~(-1)),followed by A.subcordata(134.5 Mg ha~(-1)) and T.distichum(123.3 Mg ha~(-1)).The highest soil C was stored in theplantation of A.subcordata(555.5 Mg ha~(-1)).The C storage and sequestration of the plantations after 20 years were considerable(25–30 Mg ha~(-1) year~(-1)) and broadleaves species had higher potential.Native species had a higher soil C storage while the potential of introduced species for live biomass production was higher.     

Forest structure and carbon dynamics of an intact lowland mixed dipterocarp forest in Brunei Darussalam

Tropical forests play a critical role in mitigating climate change because they account for large amount o terrestrial carbon storage and productivity.However,there are many uncertainties associated with the estimation o carbon dynamics.We estimated forest structure and carbon dynamics along a slope(17.3°–42.8°)and to assess the relations between forest structures,carbon dynamics,and slopes in an intact lowland mixed dipterocarp forest,in Kuala Belalong,Brunei Darussalam.Living biomass,basa area,stand density,crown properties,and tree family composition were measured for forest structure.Growth rate,litter production,and litter decomposition rates were also measured for carbon dynamics.The crown form index and the crown position index were used to assess crown properties,which we categorized into five stages,from very poor to perfect.The living biomass,basal area and stand density were 261.5–940.7 Mg ha~(-1),43.6–63.6 m~2ha~(-1)and 6,675–8400 tree ha~(-1),respectively.The average crown form and position index were 4,which means that the crown are mostly symmetrical and sufficiently exposed for photosynthesis.The mean biomass growth rate,litter production,litter decomposition rate were estimated as11.9,11.6 Mg ha~(-1)a~(-1),and 7.2 g a~(-1),respectively.Biomass growth rate was significantly correlated with living biomass,basal area,and crown form.Crown form appeared to strongly influence living biomass,basal area and biomass growth rate in terms of light acquisition.However,basal area,stand density,crown properties,and biomass growth rate did not vary by slope or tree family composition.The results indicate that carbon accumulation by tree growth in an intact lowland mixed dipterocarp forest depends on crown properties.Absence of any effect of tree family composition on carbon accumulation suggests that the main driver of biomass accumulation in old-growth forests of Borneo is not species-specific characteristics of tree species.     

Carbon storage in evergreen broad-leaf forests in mid-subtropical region of China at four succession stages

To better understand the effect of forest succession on carbon sequestration, we investigated carbon stock and allocation of evergreen broadleaf forest, a major zonal forest in subtropical China. We so...     

Predictive models for biomass and carbon stocks estimation in Grewia optiva on degraded lands in western Himalaya

Grewia optiva Drummond is one of important agroforestry tree species grown by the farmers in the lower and mid-hills of western Himalaya. Different models viz., monomolicular, logistic, gompetz, allometric, rechards, chapman and linear were fitted to the relationship between total biomass and diameter at breast height (DBH) as independent variable. The adjusted R² values were more than 0.924 for all the seven models implying that all models are apparently equally efficient. Out of the six non-linear models, allometric model (Y = a × DBH ^b ) fulfils the validation criterion to the best possible extent and is thus considered as best performing. Biomass in different tree components was fitted to allometric models using DBH as explanatory variable, the adjusted R² for fitted functions varied from 0.872 to 0.965 for different biomass components. The t values for all the components were found non-significant (p > 0.05), thereby indicating that model is valid. Using the developed model, the estimated total biomass varied from 6.62 Mg ha^?1 in 4 year to 46.64 Mg ha^?1 in 23 year old plantation. MAI in biomass varied from 1.66–2.05 Mg ha^?1 yr^?1. The total biomass carbon stocks varied from 1.99 Mg ha^?1 in 4 year to 15.27 Mg ha^?1 in 23 year old plantation. Rate of carbon sequestration varied from 0.63–0.81 Mg ha^?1 yr^?1. Carbon storage in the soil up to 30 cm soil depth varied from 25.4 to 33.6 Mg ha^?1.     

Profiles of carbon stocks in forest, reforestation and agricultural land, Northern Thailand

A study was conducted to assess carbon stocks in various forms and land-use types and reliably estimate the impact of land use on C stocks in the Nam Yao sub-watershed (19°05'10"N, 100°37'02"E), Thailand. The carbon stocks of aboveground, soil organic and fine root within primary forest, reforestation and agricultural land were estimated through field data collection. Results revealed that the amount of total carbon stock of forests (357.62 ± 28.51 Mg·ha-1, simplified expression of Mg (carbon)·ha-1) was significantly greater (P＜ 0.05) than the reforestation (195.25 ±14.38 Mg·ha-1) and the agricultural land (103.10±18.24 Mg·ha-1). Soil organic carbon in the forests (196.24 ±22.81 Mg·ha-1) was also significantly greater (P＜ 0.05) than the reforestation (146.83± 7.22 Mg·ha-1) and the agricultural land (95.09 ± 14.18 Mg·ha-1). The differences in carbon stocks across land-use types are the primary consequence of variations in the vegetation biomass and the soil organic matter. Fine root carbon was a small fraction of carbon stocks in all land-use types. Most of the soil organic carbon and fine root carbon content was found in the upper 40-cm layer and decreased with soil depth. The aboveground carbon(soil organic carbon: fine root carbon ratios (ABGC: SOC: FRC), was 5:8:1, 2:8:1, and 3:50:1 for the forest, reforestation and agricultural land, respectively. These results indicate that a relatively large proportion of the C loss is due to forest conversion to agricultural land. However, the C can be effectively recaptured through reforestation where high levels of C are stored in biomass as carbon sinks, facilitating carbon dioxide mitigation.     

Short-term changes in the soil carbon stocks of young oil palm-based agroforestry systems in the eastern Amazon

The current expansion of the oil palm (Elaeis guineensis Jacq.) in the Brazilian Amazon has mainly occurred within smallholder agricultural and degraded areas. Under the social and environmental scenarios associated with these areas, oil palm-based agroforestry systems represent a potentially sustainable method of expanding the crop. The capacity of such systems to store carbon (C) in the soil is an important ecosystem service that is currently not well understood. Here, we quantified the spatial variation of soil C stocks in young (2.5-year-old) oil palm-based agroforestry systems with contrasting species diversity (high vs. low); both systems were compared with a ~10-year-old forest regrowth site and a 9-year-old traditional agroforestry system. The oil palm-based agroforestry system consisted of series of double rows of oil palm and strips of various herbaceous, shrub, and tree species. The mean (±standard error) soil C stocks at 0–50 cm depth were significantly higher in the low (91.8 ± 3.1 Mg C ha^?1) and high (87.6 ± 3.3 Mg C ha^?1) species diversity oil palm-based agroforestry systems than in the forest regrowth (71.0 ± 2.4 Mg C ha^?1) and traditional agroforestry (68.4 ± 4.9 Mg C ha^?1) sites. In general, no clear spatial pattern of soil C stocks could be identified in the oil palm-based agroforestry systems. The significant difference in soil carbon between the oil palm area (under oil palm: 12.7 ± 2.3 Mg C ha^?1 and between oil palm: 10.6 ± 0.5 Mg C ha^?1) and the strip area (17.0 ± 1.4 Mg C ha^?1) at 0–5 cm depth very likely reflects the high input of organic fertilizer in the strip area of the high species diversity oil palm-based agroforestry system treatment. Overall, our results indicate a high level of early net accumulation of soil C in the oil palm-based agroforestry systems (6.6–8.3 Mg C ha^?1 year^?1) that likely reflects the combination of fire-free land preparation, organic fertilization, and the input of plant residues from pruning and weeding.     

Conversion of tropical moist forest into cacao agroforest: consequences for carbon pools and annual C sequestration

Tropical forests store a large part of the terrestrial carbon and play a key role in the global carbon (C) cycle. In parts of Southeast Asia, conversion of natural forest to cacao agroforestry systems is an important driver of deforestation, resulting in C losses from biomass and soil to the atmosphere. This case study from Sulawesi, Indonesia, compares natural forest with nearby shaded cacao agroforests for all major above and belowground biomass C pools (n = 6 plots) and net primary production (n = 3 plots). Total biomass (above- and belowground to 250 cm soil depth) in the forest (approx. 150 Mg C ha^?1) was more than eight times higher than in the agroforest (19 Mg C ha^?1). Total net primary production (NPP, above- and belowground) was larger in the forest than in the agroforest (approx. 29 vs. 20 Mg dry matter (DM) ha^?1 year^?1), while wood increment was twice as high in the forest (approx. 6 vs. 3 Mg DM ha^?1 year^?1). The SOC pools to 250 cm depth amounted to 134 and 78 Mg C ha^?1 in the forest and agroforest stands, respectively. Replacement of tropical moist forest by cacao agroforest reduces the biomass C pool by approximately 130 Mg C ha^?1; another 50 Mg C ha^?1 may be released from the soil. Further, the replacement of forest by cacao agroforest also results in a 70–80 % decrease of the annual C sequestration potential due to a significantly smaller stem increment.     

Earthworm population and microbial activity temporal dynamics in a Caspian Hyrcanian mixed forest

Few studies have analyzed how tree species within a mixed natural forest affect the dynamics of soil chemical properties and soil biological activity. This study examines seasonal changes in earthworm populations and microbial respiration under several forest species (Carpinus betulus, Ulmus minor, Pterocarya fraxinifolia, Alnus glutinosa, Populus caspica and Quercus castaneifolia) in a temperate mixed forest situated in northern Iran. Soil samplings were taken under six individual tree species (n = 5) in April, June, August and October (a total of 30 trees each month) to examine seasonal variability in soil chemical properties and soil biological activity. Earthworm density/biomass varied seasonally but not significantly between tree species. Maximum values were found in spring (10.04 m^?2/16.06 mg m^?2) and autumn (9.7 m^?2/16.98 mg m^?2) and minimum in the summer (0.43 m^?2/1.26 mg m^?2). Soil microbial respiration did not differ between tree species and showed similar temporal trends in all soils under different tree species. In contrast to earthworm activity, maximum microbial activity was measured in summer (0.44 mg CO₂–C g soil^?1 day^?1) and minimum in winter (0.24 mg CO₂–C g soil^?1 day^?1). This study shows that although tree species affected soil chemical properties (pH, organic C, total N content of mineral soils), earthworm density/biomass and microbial respiration are not affected by tree species but are controlled by tree activity and climate with strong seasonal dynamics in this temperate forest.     

Vegetation structural characteristics and topographic factors in the remnant moist Afromontane forest of Wondo Genet, south central Ethiopia

For forest ecosystem management to be effective, knowledge of the horizontal and vertical structural diversity of a forest is essential. The moist Afromontane highlands of Wondo Genet in south-central Ethiopia present an opportunity to restore and rehabilitate and enhance the ecosystem services to be obtained from this forest sustainably. We focused on the forest structural characteristics to better understand the current forest conditions to assist in the sustainable management of this resource. A total of 75 (20 m × 20 m) quadrats were sampled and diameter at breast height (DBH) ≥ 2 cm and stem height ≥ 2 m were measured. Species identity and abundance, elevation, slope, and aspect were recorded for each plot. Structural characteristics were computed for each plot. Relationship of topographic factors with vegetation characteristics was conducted using R-Software. A total of 72 woody species was recorded. Whereas, the overall diameter distribution shows an inverted J-shaped curve, the basal area followed a bell-shaped pattern. Five types of population structures are revealed. The mean tree density and basal area was 397.3 stems·ha^?1 and 31.4 m²·ha^?1, respectively. Only 2.8% of the tree species have densities of >25 stems·ha^?1 and the percentage distribution of trees show 56.2% in the DBH class 2–10 cm, indicating that the forest is dominated by medium-sized trees. Celtis africana (8.81 m²·ha^?1) and Pouteria adolfi-friedericii (5.13 m²·ha^?1) make the highest contribution to the basal area and species importance value index. The families/species with the highest importance value index are Ulmaceae, Fabacea and Sapotaceae. Species abundance (r ² =0.32, p <0.001) and species richness (r ² =0.50, p <0.001) are positively related with tree density. Tree density is negatively related with elevation (r ² =?0.36, p <0.001), slope (r ² =?0.15, p <0.001) and aspect (r ² =?0.07, p <0.05). While basal area is negatively related with elevation (r ² =?0.14, p <0.001), it has a positive relationship with tree density (r ² =0.28, p <0.001 and species richness (r ² =0.098). Species with poor population structure should be assisted by restoration tasks and further anthropogenic disturbance such as illegal logging and fuel wood extraction should be restricted.     

Stocks and dynamics of soil organic carbon and coarse woody debris in three managed and unmanaged temperate forests

The effect of forest conservation on the organic carbon (C) stock of temperate forest soils is hardly investigated. Coarse woody debris (CWD) represents an important C reservoir in unmanaged forests and potential source of C input to soils. Here, we compared aboveground CWD and soil C stocks at the stand level of three unmanaged and three adjacent managed forests in different geological and climatic regions of Bavaria, Germany. CWD accumulated over 40–100 years and yielded C stocks of 11 Mg C ha^?1 in the unmanaged spruce forest and 23 and 30 Mg C ha^?1 in the two unmanaged beech–oak forests. C stocks of the organic layer were smaller in the beech–oak forests (8 and 19 Mg C ha^?1) and greater in the spruce forest (36 Mg C ha^?1) than the C stock of CWD. Elevated aboveground CWD stocks did not coincide with greater C stocks in the organic layers and the mineral soils of the unmanaged forests. However, radiocarbon signatures of the O _e and O _a horizons differed among unmanaged and managed beech–oak forests. We attributed these differences to partly faster turnover of organic C, stimulated by greater CWD input in the unmanaged forest. Alternatively, the slower turnover of organic C in the managed forests resulted from lower litter quality following thinning or different tree species composition. Radiocarbon signatures of water-extractable dissolved organic carbon (DOC) from the top mineral soils point to CWD as potent DOC source. Our results suggest that 40–100 years of forest protection is too short to generate significant changes in C stocks and radiocarbon signatures of forest soils at the stand level.     

Temporal dynamics and spatial variations of forest vegetation carbon stock in Liaoning Province,China

There are many uncertainties in the estimation of forest car- bon sequestration in China, especially in Liaoning Province where vari- ous forest inventory data have not been fully utilized. By using forest inventory data, we estimated forest vegetation carbon stock of Liaoning Province between 1993 and 2005. Results showed that forest biomass carbon stock increased from 68.91 Tg C in 1993 to 97.51 Tg C in 2005, whereas mean carbon density increased from 18.48 Mg·ha -1 C to 22.33 Mg·ha -1 C. The carbon stora...     

Long-term response to area of competition control in <Emphasis Type="Italic">Eucalyptus globulus</Emphasis> plantations

Numerous studies have quantified the responses to vegetation management in Eucalyptus plantations but most publications have reported early responses in tree growth and a gap in knowledge exist about the magnitude and duration of growth responses throughout the whole rotation. We evaluated the long-term response (9 years-old) of E. globulus across a gradient of sites to different intensity levels of free area of competing vegetation around individual tree seedlings. Competing vegetation intensity levels considered free areas ranging between 0 (control) to 2.54 m² plus a treatment with total weed control. Competing vegetation biomass production during the first growing season was 2.9, 6.5, 2.2 and 12.9 Mg ha^?1, for sites ranging from low to high annual rainfall. Across sites, maximum response in stand volume ranged between 58 and 262 m³ ha^?1 at age 9 years and was proportional to the amount of competing biomass controlled during the first growing season. Total competing vegetation control showed the largest response in stand volume at sites with 2.9 and 12.9 Mg ha^?1 of competing vegetation. However, the 2.54 m² vegetation control treatment showed the maximum response for sites with 2.2 and 6.5 Mg ha^?1 of competing vegetation. The duration of response for vegetation control treatments ranged between 5 and 9 years. However, at the site with the largest accumulation of competing vegetation biomass the response to vegetation control showed a sustained and divergent response. Our results suggest that vegetation control improved site resources acquisition increasing long-term stand productivity by reducing environmental limitations to tree growth differentially at each site.     

Do thinnings influence biomass and soil carbon stocks in Mediterranean maritime pinewoods?

The effects of silvicultural treatments on carbon sequestration are poorly understood, particularly in areas like the Mediterranean where soil fertility is low and climatic conditions can be harsh. In order to improve our understanding of these effects, a long-term thinning experiment in a stand of Mediterranean maritime pine (Pinus pinaster Ait.) was studied to identify the effects of thinning on soil carbon (forest floor and mineral soil), above and belowground biomass and fine and coarse woody debris. The study site was a 59-year-old pinewood, where three thinnings of differing intensities were applied: unthinned (control), moderate thinning and heavy thinning. The three thinning interventions (for the managed plots) involved whole-tree harvesting. The results revealed no differences between the different thinning treatments as regards the total soil carbon pool (forest floor + mineral soil). However, differences were detected in the case of living aboveground biomass and total dead wood debris between unthinned and thinned plots; the former containing larger amounts of carbon. The total carbon present in the unthinned plots was 317 Mg ha^?1; in the moderately thinned plots, it was 256 Mg ha^?1 and in the case of heavily thinned plots, 234 Mg ha^?1. Quantification of these carbon compartments can be used as an indicator of total carbon stocks under different forest management regimes and thus identify the most appropriate to mitigate the effects of global change. Our results indicated that thinning do not alter the total soil carbon content at medium term, suggesting the sustainability of these silvicultural treatments.     

Sapling biomass allometry and carbon content in five afforestation species on marginal farmland in semi-arid Benin

Allometric equations are routinely used in the estimation of biomass stocks for carbon accounting within forest ecosystems. However, generic equations may not reflect the growth trajectories of afforestation species that are introduced to degraded farmland characterized by water and nutrient limitations. Using sequential measurements of the height, basal diameter, and above- and belowground biomass of juvenile trees, we developed allometric equations for five woody species (Moringa oleifera Lam., Leucaena leucocephala Lam., Jatropha curcas L., Anacardium occidentale L. and Parkia biglobosa Jacq.) subjected to a gradient of water and nutrient availability in an afforestation trial on degraded cropland in the semi-arid zone of Benin, West Africa. For three of the species studied, the allometric relationships between basal diameter and biomass components (i.e. leaves, stems and roots) were described best by a simple power-law model (R² > 0.93). The incorporation of species-specific height–diameter relationships and total height as additional predictors in the power-law function also produced reasonable estimates of biomass. Fifteen months after planting, roots accounted for 10–58% of the total biomass while the root-to-shoot ratio ranged between 0.16 and 0.73. The total biomass of the saplings ranged between 1.4 and 10.3 Mg ha^?1, yielding 0.6–4.3 Mg C ha^?1, far exceeding the biomass in the traditional fallow system. The rate of stem carbon sequestration measured ca. 0.62 Mg C ha^?1 year^?1. Overall, the allometric equations developed in this study are generally useful for assessing the standing shoot and root biomass of the five afforestation species during the juvenile growth stage and can help in reporting and verifying carbon stocks in young forests.     

Carbon dynamics and stability between native Masson pine and exotic slash pine plantations in subtropical China

Afforestation and ecological restoration have often been carried out with fast-growing exotic tree species because of their high apparent growth and yield. Moreover, fast-growing forest plantations have become an important component of mitigation measures to offset greenhouse gas emissions. However, information on the long-term performance of exotic and fast-growing species is often lacking especially with respect to their vulnerability to disturbance compared to native species. We compared carbon (C) storage and C accumulation rates in vegetation (above- and belowground) and soil in 21-year-old exotic slash pine (Pinus elliottii Engelm.) and native Masson pine (Pinus massoniana Lamb.) plantations, as well as their responses to a severe ice storm in 2008. Our results showed that mean C storage was 116.77 ± 7.49 t C ha^?1 in slash pine plantation and 117.89 ± 8.27 t C ha^?1 in Masson pine plantation. The aboveground C increased at a rate of 2.18 t C ha^?1 year^?1 in Masson pine and 2.23 t C ha^?1 year^?1 in slash pine plantation, and there was no significant difference in C storage accumulation between the two plantation types. However, we observed significant differences in ice storm damage with nearly 7.5 % of aboveground biomass loss in slash pine plantation compared with only 0.3 % loss in Masson pine plantation. Our findings indicated that the native pine species was more resistant to ice storm because of their adaptive biological traits (tree shape, crown structure, and leaf surface area). Overall, the native pine species might be a safer choice for both afforestation and ecological restoration in our study region.     

Biomass production,carbon sequestration and nutrient characteristics of 22-year-old support trees in black pepper (<Emphasis Type="Italic">Piper nigrum</Emphasis>. L) production systems in Kerala,India

Diverse kinds of fast growing multipurpose trees are traditionally grown as support trees (standards) for trailing black pepper vines in the humid tropics of India. Apart from differential black pepper yields, such trees exhibit considerable variability to accumulate biomass, carbon and nutrients. An attempt was made to assess the biomass production, carbon sequestration potential (tree + soil) and nutrient stocks of six multipurpose tree species (age: 22 years) used for trailing black pepper vines (Acacia auriculiformis, Artocarpus heterophyllus, Grevillea robusta, Macaranga peltata, Ailanthus triphysa and Casuarina equisetifolia). Results indicate that G. robusta showed the highest total biomass production (365.72 Mg ha^?1), with A. triphysa having the least value (155.13 Mg ha^?1). Biomass allocation among tissue types followed the order stemwood > roots > branchwood > twigs > leaves. Total C stocks were also highest for G. robusta (169 Mg C ha^?1), followed by A. auriculiformis (155 Mg C ha^?1). Mean annual carbon increment also followed a similar trend. Among the various tissue fractions, stemwood accounted for the highest N, P and K stocks, implying the potential for nutrient export from the site through wood harvest. All the support trees showed significantly higher soil carbon content compared to the treeless control. Soil N, P and K contents were higher under A. auriculiformis than other species. Nitrogen fixation potential, successional stage of the species, stand age and tree management practices such as lopping may modify the biomass allocation patterns and system productivity.     

Effect of tree spacing on growth and wood density of 38-year-old Cariniana legalis trees in Brazil

Average wood density of 38-year-old Cariniana legalis (Mart.) Kuntze, a Brazilian native forest species, was found to increase with faster growth and lower stocking, while decreasing from pith to bark. A complete randomised block design was planted with five blocks. Ten trees were harvested in each of three spacing treatments. We hypothesised that the stand stemwood production would not significantly differ depending on tree spacing. However, tree growth would be higher in the wider spacing and wood density would be higher in the narrower spacing. The diameter growth of trees was higher at 3 m × 2.5 m than at 3 m × 2 m and 3 m × 1.5 m. Nevertheless, this higher individual tree growth at 3 m × 2.5 m did not compensate for the greater tree stock density at 3 m × 1.5 m with stand stemwood production at 38 years of 530 m3 ha^?1 and 649 m³ ha^?1, respectively. These results suggest that C. legalis, which can produce up to 17 m³ ha^?1 y^?1 of medium-to high-density timber – about 800 kg m^?3 – is a promising native species for forest plantations in Brazil.