Untersuchungen zum Einfluß von Konkurrenz auf die Bestandesentwicklung von Sommerraps bei unterschiedlicher Gunst der Umweltbedingungen

Investigations on the effect of competition with regard to the stand development of spring oilseed rape as dependent on environmental conditions
Field experiments were conducted to investigate the frequency distribution in spring oilseed rape as dependent on environmental growing conditions and plant density. As measures for frequency distribution skewness and kurtosis were estimated. The plant character determined was single plant dry matter. To understand the changes in the stand structure, samples were taken at the following developmental stages: rosette, begin of flowering, end of flowering and ripeness (harvest). Plant densities ranged from 60 plants/m², 90 plants/m² till 120 plants/m²
The results show that under unfavourable growing conditions, generally speaking, the values of (positive) skewness and kurtosis increase with the development of the plants; under favourable growing conditions, however, the values of skewness and kurtosis decrease from rosette till harvest.
Under unfavourable growing conditions as well as favourable ones, the values of skewness and kurtosis increase with strengthening of plant competition as consequence of increasing plant density. However, the trend during development and the height of the values as well as the quantitative relationships are strongly controlled by the general growing conditions. Frequency distribution, therefore, is not only a result of plant competition but also of differences in resources due to the general conditions of the location.     

Untersuchungen zum Einfluß von Konkurrenzbedingungen auf die Entwicklung der Bestandesstruktur von Sommerraps

Investigations on the effect of competition on stand structure of spring oilseed rape
On the basis of a field experiment with spring oilseed rape, frequency distributions of plant dry matter and plant height were investigated. Samples were taken at four developmental stages: rosette, begin of flowering, end of flowering, ripeness (harvest). To understand the effect of competition in a plant population on frequency distribution, distance of rows (16 cm, 32 cm) and number of plants (60, 90 and 120 plants/m²) were varied. Differences in frequency distribution were estimated by skewness and kurtosis.
The results show that with age of the plant population the skewness of the plant character dry matter decreases from a high positive value till ripeness to a low positive value; kurtosis also decreases during development and tends to a value of 0 at ripeness. The effect of competition as produced by number of plants for area is mirrored by a positive skewness increasing with increasing number of plants per area. The value for kurtosis also increases with the stronger competition as consequence of plant density. Row distance did not show specific effects of frequency distribution. Plant height did, generally speaking, not react to competition.     

Comparison of Harvest Index and Grain/Straw-Ratio with Applications to Winter Oilseed Rape

To compare the statistical properties of harvest index and grain/straw-ratio their variabilities (expressed by the relative measure coefficient of variation), their skewnesses and kurtoses have been compared theoretically.
With regard to variability, the harvest index exhibits a clear superiority (= smaller variability) compared to the grain/straw-ratio. For skewness and kurtosis no such generally valid relationships exist. But, for the range of the empirically most relevant parameter values one obtains approximately equal numerical values for skewness as well as for kurtosis of harvest index and grain/straw-ratio. Finally, the theoretical results and conclusions have been checked and applied to the experimental data sets from a field trial with ten European cultivars/lines of winter oilseed rape.     

On Harvest Indices of Winter Oilseed Rape (Brassica napus L.)

Ten quite different European winter rapeseed cultivars were grown in a 3-year field trial at one location. The traits grain yield and biomass were measured based on single plant measurements (first year) or on plot basis (second and third year). Individual harvest indices were calculated.
Besides a general discussion of the experimental results for this collection of winter rapeseed cultivars the main aim of the present investigations was an examination whether these experimental results are in agreement with the theoretical conclusions on harvest indices which have been published in this journal in three preceding publications.
Hence the investigations of this paper are divided into the three chapters: 1) Calculation of mean harvest indices, 2) Variability of harvest indices and 3) Correlations among harvest index, grain yield and biomass. In all three chapters the agreement between the empirically obtained and the theoretically expected values is quite good.     

Comments on the Calculation of Mean Harvest Indices

The harvest index Z is defined, for example for cereals, as the ratio between grain yield X and total yield Y: Z = X/Y. For individual measurements x_i of X and y_i of Y the individual harvest index is calculated by z_i= x_i/y_i. These individual measurements i are based on N different experimental units (single plants, random samples from plots or total plots).
Mean harvest indices are commonly calculated either by     or by     where the means of X, Y and Z are denoted by , and respectively.
Both approaches     are, in general, different. But, in this paper it is shown that for most situations which may occur in the field of practical applications in agronomy and plant breeding both procedures will lead to nearly identical results. Only in some special situations the two results differ significantly. These conditions are investigated and explicitly given.
Some explicit formulae (approximations) for the calculation of mean harvest indices are derived and discussed.
Finally, all the theoretical investigations and results are demonstrated and applied to a numerical example of winter-rapeseed data.     

Lichtaufnahme und Stoffproduktion eines konventionellen und eines epigonalen Genotyps der Weißen Lupine (Lupinus albus)

Light interception and dry matter production of a conventional and an epigonal genotype of the white lupine ( Lupinus albus )
Yield formation of two white lupin ( Lupinus albus ) genotypes differing substantially in growth habit was investigated. Grain production was quantified as the product of harvest index and dry matter production, the latter being a function of light interception and utilization. Experimental data were obtained from two-year field experiments with a combined variation of plant density and distribution. The conventional, freely branching cultivar Kalina showed higher leaf area indices and growth rates than the "epigonal" (little branching) genotype. The higher growth rates of Kalina were partly due to increased light interception, but were mainly a result of a higher light use efficiency. This can be explained with a more even light distribution over a greater leaf area. The more rapid increase of the harvest index in the epigonal genotype compared to Kalina only partially compensated for the differences in dry matter production.     

Stickstoffaufnahme und Stickstoffrückstände von Hauptfrucht-und Ausfallrapsbeständen

Nitrogen Uptake and Nitrogen Residuals of Winter Oil-Seed Rape and Fallout Rape
The objective of the investigation was a study of the relationship between seed dry-matter production and vegetative dry-matter production in oil-seed rape crops and their dependence on the production conditions. In addition to the relationship between the N-uptake during the vegetation period and the N-residue after harvest was of major interest. Furthermore the potential for N-uptake in fallout rape was measured. Over two vegetation periods factorial field experiments with winter oil-seed rape, cv. Lirabon, different drilling techniques and different nitrogen fertilization levels were tested. Measured traits were: the dry-matter accumulation including root mass and fall-off leaf-material mass, the N-uptake of both the oil-seed crops and the fallout rape stands, and, simultaneously, the soil NO₃-N content. Finally the harvest indices and the N-harvest indices were calculated.
Combined with a N-uptake of up to 330 kg N/ha, oil-seed rape crops produced up to 200 dt dry matter/ ha. At seed yield levels of 33dt/ha (d.m.), harvest indices varied from 0.14—0.23 and N-harvest indices varied from 0.30–0.50. As a result of the residue of vegetative plant material at harvest, leaf losses before harvest and the soil NO₃-N-contents at harvest up to 275 kg N/ha remained in the field. After the harvest of oil-seed rape, the soil NO₃-N-contents were quickly reduced by emerging and growing fallout rape stands. However, following soil-preparation measures in the autumn, a continuous rise in the soil NO₃-N-content was observed.     

How does the harvest index affect water‐use efficiency and nutrient‐utilization efficiency of sunflowers (Helianthus annuus L.)?

Besides oilseed rape and soybean, sunflower is one of the most important annual oilcrops. Apart from seed yield, which is the most important trait in many crops, oil yield is of utmost interest for sunflower breeders and farmers. Compared to the seed yield of sunflowers, their total biomass is often very high, and it is of great agronomic interest to increase the ratio of both parameters, which is defined as harvest index. With an increase in harvest index, improvements of resource‐use efficiencies can be expected. To generate an adequate seed yield, water and nutrients are key factors and their efficient use becomes increasingly important under conditions of aridity and higher production goals. A pot experiment was conducted in summer 2017 to determine differences in harvest index of various sunflower genotypes which could have an influence on the use efficiencies of water and nutrients (N, P, K). In total, 25 genotypes of different origin and with various morphological traits were investigated under optimal growth conditions. The harvest index varied from 29% to 47% and showed significant differences among some genotypes. The harvest index was less affected by the total biomass of the plants but more by the seed yield. The seed yield itself was mainly determined by the single seed weight rather than by the number of seeds. The experiment also confirmed significant positive correlations between harvest indices and all use efficiencies. Genotypes with high harvest indices were characterized by high oil concentrations in the seeds and particularly by high oil yields. In conclusion, cultivation of sunflower genotypes with high harvest indices most likely results in high water‐use efficiency and high nutrient‐utilization efficiencies as well as in high oil yields.     

‘美洲黑杨’不同家系苗期生长特征比较

为获得适合长江中下游江汉平原发展的杨树新品种,以优良美洲黑杨无性系为亲本,配制了4个杂交组合,对获得的子代群体无性系化后进行生长性状调查,比较不同家系生长特征和生长节律变化。结果表明：ZS8×2-7的苗高和地径均服从正态分布(P=0.200>0.050),2-2×2-46的地径服从正态分布(P=0.057>0.050),其余家系的生长性状偏度值在-0.482~0.304之间,峰度值大多在-0.363~0.155之间,接近正态分布,4个家系苗高分布呈左偏态,大于均值的株数较多,地径分布呈右偏态,小于均值的株数相对较多;2个生长性状在家系间均存在显著差异(P<0.05),但优势并不明显,苗高均值最大的2-2×2-38比均值最小的ZS8×2-7仅高7.14%,地径均值最大的2-2×2-9比均值最小的ZS8×2-7仅粗7.52%;4个家系从6月到10月的生长节律相近,其中在7月初到7月中旬之间可能出现了夏季休眠。基于各家系生长性状的分布情况,均可开展一定强度的早期选择和淘汰以选择优良无性系。     

Nitrogen Uptake and Utilization in Winter Wheat under Different Fertilization Regimes, with Particular Reference to Main Stems and Tillers

Two-year field trials with winter wheat under nine nitrogen (N) fertilizer treatments were conducted at TU Munich's research station in Southern Bavaria. The study investigated N uptake and N utilization of wheat stands for varying N fertilizer strategies, with emphasis on N application during stem elongation. N use was analysed in individual plants, separated into main stems and tillers to estimate the effect on the tiller category. While grain N uptake showed only slight differences, substantial differences were recorded for N uptake of straw, leading to significant variation in N harvest index as affected by N fertilizer treatment. Individual plant analysis indicated that N harvest indices of tillers were lower than those of main stems due to high N uptake into the straw of tillers. This was not modified by any of the N fertilization strategies. It is presumed that for tillers the translocation of nitrogen from straw into kernels is less effective than for main stems since ears of tillers produce a lower sink size. N fertilization with emphasis on stem elongation led to a below-average number of tillers and above-average N harvest indices for main stems as well as tillers. This was confirmed by a negative correlation between plant density and N harvest index. High N content in straw was not found to be necessary for achieving high N content in kernels. In addition to economical benefits, a N fertilization strategy with emphasis on application during stem elongation has environmental benefits in intensive wheat production systems.     

The effect of skewness on selection in a plant breeding program

Summary The distribution of many plant characters is skewed. The standard formula for the expected gain from selection is based on the assumption that the character value is the sum of two independent normal variables, one genetic and the other environmental. If the genetic variable is normal, but the environmental variable has a positively skewed distribution, then upwards selection will give a lower gain than that expected if both distributions are normal. If the distribution of the environmental variable is negatively skewed, then upwards selection will give a larger than expected gain. Of course, with downwards selection, the results will be reversed. This leads to asymmetry of response if both upwards and downwards selection are used. In cases where unexpected responses to selection are obtained and normality of the character distribution has been assumed, skewness of the data should be checked as possible cause.     

海南阿陀岭4种加勒比松林分直径分布规律研究

为了给海南省低质低效次生林分的改造经营提供科学、合理的参考依据,利用正态分布、对数正态分布、Weibull分布、Γ分布4种分布函数分别对阿陀岭的4种加勒比松林分的直径分布规律进行了分析研究。结果表明：（1）4种林分的直径分布曲线均为左偏态,林木株数均偏向中小径阶;（2）加勒比松与台湾相思混交林和加勒比松与海南柃混交林用Weibull分布函数拟合的效果更好,加勒比松与浙江润楠混交林用Γ分布函数拟合的效果比较理想,加勒比松纯林用Weibull分布和Γ分布函数拟合效果接近,无显著差异。说明Weibull分布对4种林分的适应性和灵活性最强,正态分布最差,Γ分布和对数正态分布居中。     

Multivariate analysis of water-related agroclimatic factors limiting spring wheat yields on the Canadian prairies

Water use by spring wheat and soil water contents at meteorological stations on the Canadian prairies were simulated with the Versatile Soil Moisture Budget model for different crop growth stages. Six water-related agroclimatic indices at five growth stages (seeding–emergence, emergence–jointing, jointing–heading, heading–soft dough and soft dough–harvest) and previous non-growing season were correlated to spring wheat yields in the three prairies provinces and in the entire prairie region for the years 1976–2006. Principal component analysis was applied to explore major modes of joint variability in the regional water-related agroclimatic indices. Canonical correlation analysis was employed to further identify joint variability patterns of the water-related indices associated with regional spring wheat yields. Results showed some common features of the effects of the water-related factors at different growth stages: lower-than-normal moisture stress at the jointing–heading stage favoured spring wheat yields in all three provinces. Regional differences were also seen, for example, a slight moisture stress at the heading–soft dough stage could be beneficial to spring wheat yields in Manitoba because of its relatively wetter climate compared to the other two provinces. The results can be used for a better understanding of the effects of water-related agroclimatic conditions at different growth stages on final spring wheat yields on the Canadian prairies, leading to the improvement of crop management. The results can also be used in regional yield forecasting and in the projection of climate change impacts on crop production. This study provided an example of how to quantify crop–climate relationships by the use of statistical multivariate analysis tools.     

Note on the Effect of Non-regular Spatial Patterns of Plants on Yield per Area Based on a Logarithmic Relationship between Single Plant Yield and Individual Area

Based on several simplifying assumptions, a previously developed stochastic approach allows an estimation of the effects of non-regular spatial patterns of the distribution of individual plants on yield per area (F). In this approach, two random variables were attached to each plant: single plant yield (E) and individual space per plant (A). The latter was estimated by the area of Thiessen-polygons. Yield per area was calculated theoretically by the expectation of the ratio E/A. Based on a logarithmic relationship between E and A the expectation of E/A can be expressed by an approximation which depends on the mean and on the variance of the individual plant spaces. An improvement of this approximation can be easily obtained by including skewness and kurtosis of the distribution of individual plant areas. Finally, all theoretical concepts and results were applied to an experimental data set of winter oilseed rape ( Brassica napus L.).     

辽河流域玉米籽粒脱水特点及适宜收获期分析

辽河流域处于中国东北春玉米区南部,积温资源相对丰富,在该区域推广玉米机械粒收技术具有较好的热量资源基础,但区域内玉米收获时籽粒含水率偏高,机械粒收的破碎率、损失率偏高等质量问题突出。分析区域内主推品种的籽粒脱水特征、基于热量资源条件确定机械粒收的适宜时间,是解决上述问题的合理途径。2017年选择该区域主推的29个不同熟期玉米品种,在开鲁县和铁岭县开展了籽粒脱水动态观测试验。结合流域内常年春玉米播种日期、不同品种生长发育及籽粒脱水积温需求、历史气象数据等分析结果,建立不同品种在辽河流域适宜机械粒收时期的预测方法。结果显示, Logistic Power模型可以很好地模拟春玉米籽粒含水率变化过程。不同品种籽粒实收含水率与模拟含水率间存在极显著的线性关系,决定系数R2为0.916 (n=45),均方根误差RMSE为1.217。研究建立的不同品种籽粒含水率模型具有极佳的区域适用性,以2017年国审的4个宜机收品种及流域内2个主栽品种研究,明确了不同品种适宜机械粒收时期的分布规律。国审品种中,德育919和京农科728自播种至籽粒含水率降至25%活动积温需求低于3200°C d,在辽河流域大部地区可于9月中下旬达到高质量机械粒收的籽粒含水率要求。泽玉8911和吉单66积温需求低于3400°C d,可于10月上中旬在流域内实现机械粒收,较上述德育919和京农科728晚10～20d。而当地主栽的辽单575和京科968脱水至适宜籽粒含水率的积温需求较泽玉8911和吉单66多200°Cd,无法在当地常规收获期实现高质量的机械粒收。本研究检验了基于LogisticPower模型的籽粒含水率预测模型在区域分析应用中的精度。通过比较国审宜机收品种与当地主栽品种的籽粒含水率变化、成熟和脱水的积温需求以及适宜机械粒收日期的空间分布规律,更新现有品种有助于在辽河流域实现常规收获期内的高质量机械粒收。     

谷子各器官干物质量分配与产量的相关性分析

为了明确谷子各器官干物质量分配情况及其与产量的关系,本研究以6个比较有代表性的品种为研究对象,对其单株干物质量、收获指数、各器官干物质分配情况和产量的差异进行了比较研究。结果显示:不同品种谷子各器官干物质量分配规律基本一致,即穗比例>茎比例>叶比例>叶鞘比例抑根比例。杂交种‘长杂谷466’(CZ466)收获指数、单穗重和产量均是最大,株高较低;常规种‘长生13’(CS13)株高最低,产量及收获指数较高。所有品种谷子穗干物质量与单株总干物质量均显著正相关;叶鞘干物质量与穗干物质量均呈显著正相关。综上表明CZ466和CS13是2个产量高、株高矮、收获指数高的优良品种;推断谷子叶鞘和单株总干物质量与产量有较密切的相关性,其他器官干物质量与产量的相关性因品种而异。     

Relationship between grain yield and grain nitrogen concentration in winter triticale

Summary Ten hexaploid winter triticale lines were grown for two cropping periods at three locations in western Switzerland. Averaged across the six environments, the differences between lines were statistically significant (P=0.05) for grain yield, above-ground biomass, N uptake, grain N yield, nitrogen harvest index, grain N concentration and straw N concentration. There were significant line x environment interactions for all traits. Grain yield and grain N concentration were inversely related (r=–0.74^**). Diagrams in which grain yields were plotted against grain N concentration were used to identify lines with a consistently unusual combination of grain yield and grain N concentration. Despite comparable grain yields, Line 3 had a high grain N concentration, while that of Line 7 was low. Line 3 was superior to Line 7 in both N uptake and N harvest index. Averaged across environments and lines, the N harvest index was 0.73 which corresponds to N harvest indices reported for bread wheat in the same region. We considered the feasibility of developing triticale lines which would outperform the best recent ones in N uptake and partitioning. However, we doubted that this would bring about a marked increase in grain N concentration, because, in the long run, the expected genetic progress in grain yield will lead to a dilution of grain protein by grain carbohydrate increments.Abbreviations GNC grain N concentration - GNY grain N yield - GY grain yield - HI above-ground dry matter harvest index - NHI nitrogen harvest index - SNC straw N concentration - TB total above-ground biomass - TPN total plant N     

收获期对不同生育期高油大豆油分及产量的影响

利用不同生育期高油大豆品种的收获期试验,来研究收获期对不同生育期高油大豆品种油分及产量的影响。采用分期收获的方法对6个不同生育期高油大豆品种进行处理,利用近红外谷物品质分析仪进行油分分析,DPS数据处理系统进行数据处理。试验表明,收获期对大豆油分含量和产量有显著的影响。从初熟期开始,随着收获期的延迟,各品种油分含量呈先上升后下降有规律的变化趋势,6个品种的油分含量变化幅度为0.37-0.87个百分点,每个品种都有一个最佳的收获时期以获得最高的油分含量,但油分含量变化情况和品种的生育期相关性不大,是由品种本身的遗传特性决定的。播期对产量也有较明显的影响作用,四个生育期相对较长的品种黑农44、垦农19、合丰48号和合丰47号产量变化较大,达到了极显著水平,而两个早熟品种的产量变化较小,不显著,产量变化总体趋势是晚熟品种＞中熟品种＞早熟品种。不同生育期的高油品种适期收获能获得较高的油分和产量。     

The Effects of Timing of an Interim Harvest on the Yield and Composition of Lablab purpureus

Lablab purpureus was grown in two field experiments in consecutive years to evaluate the effects of the timing of an interim harvest on the yield and nutritive value of the harvested material and the subsequent regrowth, which was determined from the second and final harvest. Delaying the first harvest to 50–70 days post-sowing increased the total (first plus second harvests) dry matter (DM) yield, with a greater first harvest and reduced second harvest DM yield. The delay also reduced the crude protein concentration of the first harvest and tended to increase its modified acid-detergent (MAD) fibre concentrations. The delay increased the crude protein concentration and decreased the MAD fibre concentration of the second harvest. The total crude protein yield of both harvests increased with late interim harvesting. The first harvest plant calcium concentration increased and phosphorus concentration decreased with a delay in the interim harvest. It is concluded that, in the difficult growing conditions of the Sahelian zone of sub-Saharan Africa, delaying the interim harvest of Lablab purpureus until 50–70 days post-sowing will have beneficial effects on total dry matter and crude protein yields.     

Nitrogen uptake and nitrogen harvest index in durum wheat cultivars varying in their grain protein concentration

Summary In a field experiment, plant (excluding roots) and grain nitrogen at harvest were estimated in 15 durum wheat cultivars varying in their grain protein concentration. They showed significant variation in grain yield, grain protein concentration, biological yield, total plant nitrogen at harvest and residual nitrogen in straw. Harvest index and nitrogen harvest index were calculated from this primary data. Nitrogen harvest index varied from 57 to 83%. Plant nitrogen showed significant positive correlation with biological yield, grain yield and grain protein yield, but the correlations with grain protein concentration, harvest index and nitrogen harvest index were not significant. Nitrogen harvest index was positively correlated with harvest index indicating that the distribution of N between straw and grain to a large extent, but not entirely, depends upon the partitioning of dry matter between the two. Grain protein concentration was neither correlated significantly to plant nitrogen nor to nitrogen harvest index.