Variation in the relative abundance of the large pine weevil among Sitka spruce plantation sites

Between 1993 and 1994 over 160 forest sites were sampled toobtain indices of pine weevil (Hylobius abietisL.) densities.Overall 48 variables were tested for their ability to explainvariation in weevil numbers between compartments. The modelsgenerated from the sample data identified 14 variables as significantin influencing weevil densities. In this research note the variablesare identified, and their relationship to forest managementdiscussed     

Interception of precipitation by pole-stage Sitka spruce and lodgepole pine and mature Sitka spruce at Kielder Forest, Northumberland

Interception loss was measured indirectly for 3 years in adjacentunthinned 25-year-old stands of Sitka spruce and lodgepole pineand for 2 years in a 63-year-old selectively thinned stand ofSitka spruce. For each stand measurements were made of grossprecipitation, throughfall and stemflow. Interception loss wasderived by subtracting the sum of throughfall and stemflow (netprecipitation) from gross precipitation. For the years 1977–78, 1978–79 and 1979–80,interception losses in the pole-stage Sitka spruce were 32,28 and 27 per cent of gross precipitation respectively whilethose in the lodgepole pine were 33, 29 and 26 per cent. For1979–80 and 1980–81 interception losses in the matureSitka spruce were 44 and 53 per cent of gross precipitation.The average interception loss was 29 per cent for both pole-stagecrops and 49 per cent for mature Sitka spruce. The proportions of net precipitation reaching the ground asstemflow and throughfall were 0.18 and 0.82 respectively forthe pole-stage Sitka spruce, 0.14 and 0.86 for the pole-stagelodepole pine and 0.02 and 0.98 for the mature Sitka spruce.     

Internal cycling of nitrogen, potassium and magnesium in young Sitka spruce

Potassium (K) and magnesium (Mg) are essential macro-nutrients, but little is known about how they are cycled within plants. Stable isotope studies have shown that the internal cycling of nitrogen (N) is independent of current nutrient supply in temperate tree species. This is ecologically significant because it allows trees to produce rapid shoot growth in spring independent of current soil N uptake. We used stable isotopes to quantify N, K and Mg in new shoots of Sitka spruce (Picea sitchensis (Bong.) Carr.) seedlings and to compare the relative contributions from current uptake and internal cycling. Two-year-old Sitka spruce seedlings were labeled with (15)N, (41)K and (26)Mg in an abundant or a limited supply for one growing season. The trees were repotted in the subsequent dormant season to prevent further root uptake of enriched isotopes and provided with an abundant or a limited supply of unlabeled nutrients until they were harvested in early summer of the following year. The supply was switched for half the trees in the second year to create four nutrient regimes. Enrichment of (15)N, (41)K and (26)Mg in current-year growth was attributed to internally cycled N, K and Mg uptake from the previous year. The internal cycling of N, K and Mg in new growth was significantly affected by the first-year nutrient treatments. The second-year nutrient supply affected the growth rates of the trees, but had no effect on the amounts of N, K or Mg contributed from internal cycling. Thus, internal cycling of K and Mg in Sitka spruce are, like that of N, independent of current nutrient supply.     

Wettability of Sitka spruce timber stored in a Scottish sawmill yard

Kiln-dried and air-dried Sitka spruce battens were exposed to outdoor weathering within stickered, close packed and wrapped packs and at the upper surface of these packs. A four-week exposure period was used in James Jones and Sons Ltd. sawmill yard near Aboyne, NE Scotland. The wettability of batten samples removed from the packs of timber was determined by a drop contact angle method using a video tape technique. Timber exposed to air drying before weathering in the experimental packs generally had a greater wettability than kiln-dried timber at the end of the exposure period. However, kiln-dried timber stored in stickered packs developed a greater wettability than similarly stored air-dried timber, suggesting that the kiln drying process increased the susceptibility of timber to developing increased wettability. Close-packed timber protected from sunlight and rain water, by covering the upper surface of the pack with microporous plastic sheeting, maintained a lower wettability than unprotected close-packed or stickered timber. Fungal spoilage was unlikely to develop on timber stored in pack types which developed high timber wettability, the storage conditions leading to an increase in wettability were adverse to fungal development. Kiln-dried timber stored in stickered packs in the sawmill yard may become particularly vulnerable to fungal attack if subsequently exposed to damp conditions in service. Received 15 October 1998     

Bending and shear properties of compressed Sitka spruce

We examined the bending and shear properties of compressed wood using small and clear specimens of Sitka spruce (Picea sitchensis Carr.). For measuring the bending properties, three-point bending tests were conducted under the span/depth ratio of 14, which is standardized in the American Society for Testing and Materials [ASTM D143-94 (2005a)] and Japanese Industrial Standards [JIS Z2101-94 (1994)]. In the bending test, the load, deflection at the midspan, and strain at the bottom of the midspan were simultaneously measured, and Young’s modulus and bending strength were obtained by elementary beam theory. For obtaining the shear modulus and shear strength, asymmetric four-point bending tests were conducted using the specimens with rectangular and side-grooved cross sections, respectively, and the influence of the compression ratio on the shear properties was examined. The results are summarized as follows: (1) Young’s modulus increased with increasing compression ratio when it was determined by the load–strain relation. Nevertheless, this tendency was rather obscured when Young’s modulus was determined by the load–deflection relation. Hence, it is preferable that Young’s modulus is measured from the load–strain relation. (2) The shear modulus in the longitudinal–tangential plane was maximum at the compression ratio of 50%, whereas that in the longitudinal–radial plane was minimum at the compression ratio of 50%. (3) The influence of the compression on the bending and shear strength ratio was not significant.     

Differences in mycorrhizal status among clones of Sitka spruce

Cuttings of Sitka spruce clones with the same maternal parentage were rooted under mist and examined after they had been grown for a year in a forest nursery. All short roots were found to be mycorrhizal, but one clone differed from the others in the proportion of the roots colonized by different mycorrhizal fungi. Substantial clonal differences in root morphology were partially explained by these mycorrhizal differences. The significance of the host-fungus genotype interaction is discussed.     

Autumn frost damage: clonal variation in Sitka spruce

Current-year needles of Sitka spruce (Picea sitchensis) can be injured by frost after hardening of stem and buds in the autumn, and in Scotland trees of southern provenances are most susceptible to damage. In October 1993 a frost of −5°C caused needle damage to clonal plants of Alaskan, Queen Charlotte Islands, and Washington provenances of Sitka spruce that had been selected for variation in the onset of root dormancy. Visual damage assessment revealed significant differences among clones in the severity of damage, and this damage was positively correlated with the lateness of root dormancy. Measurement of electrolyte leakage from shoot samples confirmed the observed differences between clones. The large clonal differences in frost hardiness found in this study demonstrate a potential for selecting frost tolerant trees from within southern provenances of Sitka spruce.     

Vibrational properties of Sitka spruce heat-treated in nitrogen gas

Sitka spruce (Picea sitchensis Carr.) wood was heated for 0.5–16.Oh at temperatures of 120°–200°C in nitrogen gas or air. The values for Young's modulus, shear modulus, and loss tangent were measured by free-free flexural vibration tests. X-ray diffractometry was carried out to estimate the crystallinity index and crystallite width. The results obtained are as follows: (1) Density decreased at higher temperatures and longer heating times. The specific Young's modulus, specific shear modulus, crystallinity index, and crystallite width increased during the initial stage and were constant after this stage at 120°C and 160°C, whereas they increased during the initial stage and decreased later when the temperature was high. Loss tangent in the longitudinal direction increased under all conditions, whereas that in the radial direction increased at 120°C and decreased at 160°C and 200°C. (2) From the relation between Young's modulus and moisture content, it can be safely said that Young's modulus is increased by the crystallization and the decrement in equilibrium moisture content, and that crystallization (rather than degradation) is predominant at the initial stage of the heat treatment, whereas the latter is predominant as the heating time increases. (3) It is implied that the specific Young's modulus, specific shear modulus, crystallinity index, and crystallite width decreased more in air than in nitrogen gas because of oxidation in air.This study was presented in part at the 43th Annual Meeting of Japan Wood Research Society at Morioka, August 1993, the 44th Annual Meeting of Japan Wood Research Society at Nara, April 1994, and the 45th Annual Meeting of Japan Wood Research Society at Tokyo, April 1995     

Trade-offs between seedling growth,thinning and stand stability in Sitka spruce stands: a modelling analysis

The forest industry is increasingly adopting alternative silvicultural systems, involving regeneration beneath an existing forest canopy, rather than clear-felling and replanting. To apply these silvicultural systems in windy regions such as Britain and Ireland, it is essential that the interactions between thinning intensity, stand stability and seedling growth are properly understood. Here, we present a modelling analysis of the three key relationships between: (i) stand density and the proportion of incident radiation transmitted through a forest canopy as a stand is thinned; (ii) transmitted radiation and seedling growth, and (iii) stand density and stand stability. These relationships were examined using separate models of radiative transfer (MAESTRO), seedling growth, and stand stability/wind risk (ForestGALES). Output from the three models was synthesised to calculate whether a given stand thinned to a pre-defined stability limit would allow sufficient light to penetrate the canopy for seedling growth. A minimum transmittance of 20% was identified as a requirement for seedling growth, which corresponds to removing 45% of stand basal area. A thinning of this intensity left some stands susceptible to unacceptable wind damage, especially in old or previously thinned stands on soils where rooting is impeded. The results emphasised the fact that rooting conditions, thinning history and age of intervention are major constraints on the silvicultural options. In general, older stands are not suitable for conversion to continuous cover forestry (CCF) systems, and the transformation process should begin at pole stage, when heavy thinning does not leave the stand unstable. The analysis approach used here illustrates the potential for combining models to address complex forest management issues.     

Regeneration dynamics of Sitka spruce in artificially created forest gaps

This study examined the variation in the development of naturally regenerated and planted seedlings of Sitka spruce (Picea sitchensis (Bong.) Carr.) within gaps cut in a 32-year-old stand of the same species. The circular gaps were 20 m in diameter and designed to allow sunlight into only half of the gap floor at midsummer given the latitude of 56°45′N. Eight plots (8 m × 3 m) were laid out along a north–south transect through each gap (four within the gap and two each under the closed canopy north and south of the gap). Each plot was sub-divided and seedlings were planted into one part and the other part was left to naturally regenerate. In subsequent seasons, plots were further subdivided into ‘weed free’ and ‘vegetation left untouched’. Results showed that while the two central plots within the gaps had the highest value of canopy openness, the highest accumulated temperature and lowest soil moisture were recorded in plots that received direct sunlight. However, level of germination was significantly higher in the shaded area of the gap than in the part that received direct sunshine suggesting that higher moisture levels in shaded areas are important to successful germination. Minimal germination was recorded in the plots beneath the canopy. Seedling survival was significantly influenced by the influx of competing vegetation, but only in the part of the gaps that received direct sunlight. The success of Sitka spruce regeneration within gaps appears to depend on sufficient moisture and light to support regeneration and early growth, but not too much light to encourage the development of competing vegetation. The permanently shaded areas of the gaps appeared to offer ground conditions with sufficient moisture and light to ensure successful germination and early growth of seedlings, but without excessive competition from other vegetation.     

Effects of seed size on seedling attributes in Sitka spruce

No statistically-significant effects of size (small and large) of wind-pollinated seeds from 18 seed-orchard clones could be found in the attributes of 8-month-old seedlings of Sitka spruce (Picea sitchensis (Bong.) Carr.). Clonal variations in seedling attributes was highly significant (P < 0.01), but these variations were within regional specifications for operationally-produced nursery stock.     

Biomass and net productivity for spruce plantation

SiteandmethodThesiteislocatedintheSuiIingDistrictofHeiIong-jiangProvincefromN47"26'toN48"o6',E127"37'toE128"28'.TheaveragesIopeis15".Themeanelevationis349m.ThemeanannuaItem-peratureis-o.4t.TheaccumuIatedtemperatureof31ooCis1986C-Theamountofprecipitationisfrom6ooto8oomm.Thefrost-freeseasonisabout13od.ThemainsoilisdarkbrownsoiI.The17plots(o.o4hm')fordifferentages,differ-entsitesweremeasured.OnesampletreewasseIectedineverypIot,ThefalIenstemanalysistreesweredividedwithonemeterortwometer…     

Floristic composition and spatial distribution of germinable seeds in a spruce plantation

Introduction

The logging of several spruce plantations is planned for the next years in the Eifel National Park (Western Germany). To understand the ecological mechanisms of spontaneous forest regeneration that follows these impacts, it is crucial to investigate the species composition and functional diversity of the buried germinable seeds in those stands.

Methods

We analysed the soil seed bank by germination method and characterised germination dynamics and distribution of seedlings throughout the samples according to a sigmoid model and cumulative curves, respectively. To compare the proportions of functional and syntaxonomical groups between seed bank and aboveground vegetation, we applied the Fisher??s exact test. Additionally, we described the spatial distribution of the species in the seed bank according to semivariograms.

Results

The studied seed bank reached a density of about 5,300 seed?m^?2. Most of the species detected in the seed bank are related to open vegetation, forest fringes and early successional stages of forest regeneration. In spite of a low floristic similarity between seed bank and aboveground vegetation, the functional and syntaxonomical composition is very similar. Further studies considering the distribution of seeds in the soil and the changes of the seed bank composition during the regeneration process are required.     

Mycorrhizal and associated fungi of Sitka spruce in Irish forest mixed stands

Identity of mycorrhizas and isolation of symbionts and associated fungi from Sitka spruce growing in pure and mixed stands with either Japanese larch or Lodgepole pine are described and compared. More mycorrhizal types and sporocarps of the Agaricales were collected from mixed stands. Mycorrhizas of Lactarius rufus, Paxillus involutus and Suillus spp. were more prevalent on roots from mixed stands. The most common unidentified mycorrhizal type (type B) had features similar to synthesized mycorrhizas of two Basidiomycete isolates. Suillus grevillei and three un-identified types were associated specifically with Japanese larch. The main associating fungi were Oidiodendron sp. and Mycelium radicis atrovirens. The association of a “nurse” tree with Sitka spruce provides a more diverse mycorrhizal flora, the majority of which are shared between the tree species.     

Isolation of symbiont and associated fungi from ectomycorrhizas of Sitka spruce

Isolation of symbiont and associated fungi from ectomycorrhizas of Sitka spruce from three Irish forest plantations is described. Frequent emergence of dark sterile mycelia and Oidiodendron spp., predominantly O. maius, restricted the recovery of potential symbionts. Inhibition of the former by benomyl resulted in a sixfold increase in basidiomycetes isolated. Pure culture syntheses confirmed the ectomycorrhizal ability of potential symbiont isolates and demonstrated the parasitic nature of dark sterile mycelial types. Antagonism of Mycelium radicis atrovirens, Phytophthora cinnamomi and Heterobasidion annosum by O. maius observed in paired cultures supports the possible involvement of this species in biological control of conifer root pathogens.