关于异龄林大小等级划分标准的探讨

在森林资源调查中,对异龄林用平均年龄和龄组进行归类,会缩小不同林分之间的结构差异,并造成林分年龄的偏低估计。利用海南省热带天然林样地资料。在对异龄林结构进行分析的基础上,提出用占总株数20％的大径级林木来计算异龄林的平均直径、平均树高等林分特征因子,并根据平均直径大小按“径组”（或粗度级）将异龄林分为5级,基本与同龄林的5个龄组相对应。便于对森林资源的统计和汇总分析。提出的径组划分标准可为异龄林的调查和统计提供参考,也为开展这方面的进一步研究提供借鉴。     

Modelling near-natural silvicultural regimes for beech – an economic sensitivity analysis

Near-natural silvicultural regimes for beech (Fagus sylvatica L.), in the form of uneven-aged selective felling, receive increasing interest in Denmark. This is partly due to the ability of this system to preserve the forest climate and maintain important ecological functions such as bio-geochemical cycling and biodiversity conservation. The optimal age to convert from even-aged management to uneven-aged selective felling seems to be as early as possible, about 55 years, when sufficient natural seeding can be expected. A fixed conversion period of 100 years is assumed. This regime, here analysed by the use of a so-called chessboard model, appears to be economically superior to clear felling if the regime is initiated in medium-aged stands down to the age of 55 years, assuming a high site quality and a real discount rate of 3% per annum. Uneven-aged selective felling seems to be economically superior to even-aged natural regeneration, assuming that: (i) the stumpage prices of regeneration harvests are increased by 10%, or (ii) the diameters of regeneration harvests exceed those of even-aged management by about 17% simultaneous with an identical increase of the maximum stumpage price. Clear felling seems to be the more profitable regime only if: (i) the stand is close to the economic optimal rotation age for clear felling, and (ii) when the stumpage prices of regeneration harvests achieved by use of the uneven-aged selective management regime are reduced by 10% or more due to quality defects caused by prolonged rotation ages. The above results are sensitive to variation of stumpage prices, but less so to variation of regeneration costs associated with near-natural management systems. However, the near-natural silvicultural regimes may be unable to fulfill the liquidity objectives following from the application of traditional management systems.     

Evaluation of forest management systems under risk of wildfire

We evaluate the economic efficiency of even- and uneven-aged management systems under risk of wildfire. The management problems are formulated for a mixed-conifer stand and approximations of the optimal solutions are obtained using simulation optimization. The Northern Idaho variant of the Forest Vegetation Simulator and its Fire and Fuels Extension is used to predict stand growth and fire effects. Interest rate and fire risk are found to be critical determinants of the superior stand management system and timber supply. Uneven-aged management is superior with higher interest rates with or without fire risk. Alterations in the interest rate affect optimal stocking levels of uneven-aged stands, but have only minor effects on the long-run timber supply. Higher interest rates reduce rotation length and regeneration investments of even-aged stands, which lead to markedly reduced timber supply. Increasing fire risk increases the relative efficiency of even-aged management because a single age cohort is less susceptible to fire damage over the course of the rotation than multiple cohorts in uneven-aged stands. Higher fire risk reduces optimal diameter limit under uneven-aged management and decreases optimal rotation length and planting density under even-aged management.     

Optimal management of uneven-aged Norway spruce stands

An optimization model is specified to analyze forest management without any restrictions on the forest management system. The data on forest growth comes from unique field experiments and is used to estimate a nonlinear transition matrix or size-structured model for Norway spruce. The objective function includes detailed harvesting cost specifications and the optimization problem is solved in its most general dynamic form. In optimal uneven-aged management, stand density is shown to be dominated by limitations in natural regeneration. If the goal is volume maximization, even-aged management with artificial regeneration (and thinnings from above) is superior to uneven-aged management. After including regeneration and harvesting costs, the interest rate, and the price differential between saw timber and pulpwood, uneven-aged management becomes superior to even-aged management. However, in the short term the superiority is conditional on the initial stand state.     

美国的森林植被模型

从1973年以来, 森林植被模型在美国就被作为森林生长模拟模型来使用, 广泛用于评价林分目前的生长状况和预测不同经营管理措施下林分未来的动态变化。其能够模拟各种不同的森林类型和林分结构, 包括同龄林、异龄林、纯林和混交林, 以及单层混交林和复层混交林等。同时, 森林植被模型能够模拟大多数主要森林树种、森林类型和林分条件在不同森林培育措施(包括疏伐、施肥等)下的变化状况。森林植被模拟体系还能够评价和预测病虫害和火灾等干扰因子对林分状况的影响。目前这个模型已经成为一个高度综合的分析工具体系。     

Beyond Even- vs. Uneven-Aged Management

The effect of forest management has been to override the dynamics produced by natural disturbances with a dynamic of human design. Under even-aged management, the forest is held in an early successional state, providing some habitat for species of open field and edge but little for species of the forest interior. Alternatively, uneven-aged management may maintain forest interior conditions but does not provide the large-scale heterogeneity created by pre-settlement disturbance processes. The traditional choice between even- and uneven-aged management approaches condemns the landscape to one of two mutually exclusive dynamics, causing the loss of natural landscape heterogeieity and the biodiversity it supports. The conservation of biodiversity will require that silviculture move beyond the artificial dichotomy of even- versus uneven-aged management to accommodate the full range of patterns, intensities, and frequencies of the natural disturbance regime. Traditional emphases on "regenerating the stand" and "optimizing growing stock" must be replaced by an emphasis on establishing the appropriate-sized cohort in the context of landscape-scale objectives.     

林木生长和收获预估模型的研究动态

从近年来林木生长和收获模型的研究中,可以看出有如下的发展动向:（1）由林分水平模型向径级水平模型、单木水平模型转变;（2）从林木的自然生长模型发展来考虑人为育林措施的经营模型;（3）由简单的人工同龄纯林模型深化到复杂的天然异龄混交林;（4）关注点从测树学调查因子的经验性模型转向生理生态的机理性模型。还就今后我国的林木生长模型研究提出了几点建议。     

Partial cutting in mixedwood stands: Effects of treatment configuration and intensity on stand structure,regeneration, and tree mortality

In temperate and boreal mixedwood forests of eastern North America, partial disturbances such as insect outbreaks and gap dynamics result in the development of irregular forest structures. From a forest ecosystem management perspective, management of these forests should therefore include silvicultural regimes that incorporate medium- to high-retention harvesting. We present 12-year results of a field experiment undertaken to evaluate the effects of variable retention harvesting on stand structure, recruitment, and mortality. Treatments were gap harvesting (GAP), diameter-limit harvesting (DL), careful logging (CL), and careful logging followed by scarification (CL + SCAR), and an unharvested control. Although post-harvest basal area in the GAP treatment was significantly lower than that of controls, it maintained a diameter distribution profile and densities of balsam fir regeneration similar to those of pre-harvest conditions. Lower retention treatments (DL, CL, and CL + SCAR) tended to favor regeneration of pioneer, shade-intolerant species. Except for black spruce (for which mortality was highest in DL), stem mortality was similar among harvesting treatments. From an ecosystem management perspective, this study suggests that gap harvesting can maintain, in the short term, forest stand composition and structure similar to unharvested forests, and could be used where management objectives include the maintenance of late successional forest conditions.     

Dynamics and drivers of aboveground biomass accumulation during recovery from selective harvesting in an uneven-aged forest

European Journal of Forest Research - Selective harvesting is a common silvicultural practice to ensure sustainable use of uneven-aged forests, but our understanding of how multiple forest...     

How to quantify forest management intensity in Central European forests

Existing approaches for the assessment of forest management intensity lack a widely accepted, purely quantitative measure for ranking a set of forest stands along a gradient of management intensity. We have developed a silvicultural management intensity indicator (SMI) which combines three main characteristics of a given stand: tree species, stand age and aboveground, living and dead wooden biomass. Data on these three factors are used as input to represent the risk of stand loss, which is a function of tree species and stand age, and stand density, which is a function of the silvicultural regime, stand age and tree species. Consequently, the indicator consists of a risk component (SMI_r) and a density component (SMI_d). We used SMI to rank traditional management of the main Central European tree species: Norway spruce (Picea abies [Karst.] L.), European beech (Fagus sylvatica L.), Scots pine (Pinus sylvestris L.), and oak (Quercus robur L. and Quercus petraea L.). By analysing SMI over their whole rotation period, we found the following ranking of management intensity: oak<beech<pine?spruce. Additionally, we quantified the SMI of actual research plots of the German Biodiversity exploratories, which represent unmanaged and managed forest stands including conifer forests cultivated outside their natural range. SMI not only successfully separate managed from unmanaged forests, but also reflected the variability of forest management and stand properties across the entire sample and within the different management groups. We suggest using SMI to quantify silvicultual management intensity of stands differing in species composition, age, silvicultural system (even-aged vs. uneven-aged), thinning grade and stages of stand conversion from one stand type into another. Using SMI may facilitate the assessment of the impact of forest management intensity on biodiversity in temperate forests.     

Tree size distribution and abundance explain structural complexity differentially within stands of even-aged and uneven-aged structure types

Characterizations of physical structural complexity are an important surrogate for the potential of forested stands to provide desired ecosystem services such as biodiversity. Distinguishing between stands with different structural conditions is not only a necessary feature of useful structural metrics and indices, but how such measures vary among stands can reveal clues to the ecological processes driving structure. We used stand inventory metrics and indices of structural complexity to differentiate between even-aged and uneven-aged structure types using 10 stem-mapped coniferous stands of each type distributed across Switzerland. Within each structure type, we further explored relationships among stand inventory metrics and structural indices over a roughly 10-year period of management intervention. The even-aged and uneven-aged structure types were clearly differentiated using both stand inventory metrics and spatially explicit structural complexity indices. Overall, structural complexity within even-aged stands was strongly related to, and best predicted by, metrics including the distribution of basal area among canopy layers, while complexity in the uneven-aged stands was most strongly related to, and best predicted by, metrics including measures of abundance. Although predictive models could be developed for canopy position mixture, diameter differentiation, and small-scale structural complexity (but not spatial aggregation) using only stand inventory metrics, the prediction success after only a single management intervention was lower than expected. These results indicate that research to explore small-scale structural complexity requires detailed spatially explicit inventory data and that management to enhance structural complexity may require the manipulation of different attributes in stands of even-aged (diameter distribution) and uneven-aged (total abundance) structure types.     

Analysis of the changes in forest ecosystem functions,structure and composition in the Black Sea region of Turkey

We used geographical information system to analyze changes in forest ecosystem functions, structure and composition in a typical department of forest management area consisting of four forest management planning units in Turkey. To assess these effects over a 25 year period we compiled data from three forest management plans that were made in 1986, 2001 and 2011. Temporal changes in forest ecosystem functions were estimated based on the three pillars of forest sustainability: economics, ecology and socio-culture. We assessed a few indicators such as land-use and forest cover, forest types, tree species, development stage, stand age classes, crown closure, growing stock and its increment, and timber biomass. The results of the case study suggested a shift in forest values away from economic values toward ecological and socio-cultural values over last two planning periods. Forest ecosystem structure improved, due mainly to increasing forest area, decreasing non-forest areas (especially in settlement and agricultural areas), forestation on forest openings, rehabilitation of degraded forests, conversion of even-aged forests to uneven-aged forests and conversion of coppice forests to high forests with greater growing stock increments. There were also favorable changes in forest management planning approaches.     

Modeling silviculture after natural disturbance to sustain biodiversity in the longleaf pine (Pinus palustris) ecosystem: balancing complexity and implementation

Modeling silviculture after natural disturbance to maintain biodiversity is a popular concept, yet its application remains elusive. We discuss difficulties inherent to this idea, and suggest approaches to facilitate implementation, using longleaf pine (Pinus palustris) as an example. Natural disturbance regimes are spatially and temporally variable. Variability leads to a range of structural outcomes, or results in different pathways leading to similar structures. In longleaf pine, lightning, hurricanes, surface fires, and windthrow all lead to similar structures, but at different rates. Consequently, a manager can select among various natural disturbance patterns when searching for an appropriate silvicultural model. This facilitates management by providing flexibility to meet a range of objectives. The outcomes of natural disturbances are inherently different from those of silviculture, for example, harvesting always removes boles. It is instructive to think of silvicultural disturbances along a gradient in structural outcomes, reflecting degree of disparity with natural disturbance. In longleaf pine this might involve managing for two-cohort structure, instead of multi-cohort structure characteristic of old growth stands. While two-cohort structure is a simplification over the old growth condition, it is an improvement over single-cohort management. Reducing structural disparity between managed and unmanaged forests is key to sustaining biodiversity because of linkages that exist between structural elements, forest biota, and ecosystem processes. Finally, interactions of frequency, severity, intensity, seasonality, and spatial pattern define a disturbance regime. These components may not have equal weight in affecting biodiversity. Some are easier to emulate with silviculture than are others. For instance, ecologists consider growing-season fire more reflective of the natural fire regime in longleaf pine and critical for maintenance of biodiversity. However, dormant season fire is easier to use and recent work with native plants suggests that seasonality of fire may be less critical to maintenance of species richness, as one component of biodiversity, than is generally believed. Science can advance the goal of modeling silviculture after natural disturbances by better illustrating cause and effect relationships among components of disturbance regimes and the structure and function of ecosystems. Wide application requires approaches that are adaptable to different operational situations and landowner objectives. A key point for managers to remember is that strict adherence to a silvicultural regime that closely parallels a natural disturbance regime may not always be necessary to maintain biodiversity. We outline examples of silvicultural systems for longleaf pine that demonstrates these ideas.     

Carbon dynamics of North American boreal forest after stand replacing wildfire and clearcut logging

Boreal forest carbon (C) storage and sequestration is a critical element for global C management and is largely disturbance driven. The disturbance regime can be natural or anthropogenic with varying intensity and frequency that differ temporally and spatially the boreal forest. The objective of this review was to synthesize the literature on C dynamics of North American boreal forests after most common disturbances, stand replacing wildfire and clearcut logging. Forest ecosystem C is stored in four major pools: live biomass, dead biomass, organic soil horizons, and mineral soil. Carbon cycling among these pools is inter-related and largely determined by disturbance type and time since disturbance. Following a stand replacing disturbance, (1) live biomass increases rapidly leading to the maximal biomass stage, then stabilizes or slightly declines at old-growth or gap dynamics stage at which late-successional tree species dominate the stand; (2) dead woody material carbon generally follows a U-shaped pattern during succession; (3) forest floor carbon increases throughout stand development; and (4) mineral soil carbon appears to be more or less stable throughout stand development. Wildfire and harvesting differ in many ways, fire being more of a chemical and harvesting a mechanical disturbance. Fire consumes forest floor and small live vegetation and foliage, whereas logging removes large stems. Overall, the effects of the two disturbances on C dynamics in boreal forest are poorly understood. There is also a scarcity of literature dealing with C dynamics of plant coarse and fine roots, understory vegetation, small-sized and buried dead material, forest floor, and mineral soil.     

A density management diagram for Norway spruce in the temperate European montane region

Norway spruce is one of the most important conifer tree species in Europe, paramount for timber provision, habitat, recreation, and protection of mountain roads and settlements from natural hazards. Although natural Norway spruce forests exhibit diverse structures, even-aged stands can arise after disturbance or as the result of common silvicultural practice, including off-site afforestation. Many even-aged Norway spruce forests face issues such as senescence, insufficient regeneration, mechanical stability, sensitivity to biotic disturbances, and restoration. We propose the use of Density Management Diagrams (DMD), stand-scale graphical models designed to project growth and yield of even-aged forests, as a heuristic tool for assessing the structure and development of even-aged Norway spruce stands. DMDs are predicated on basic tree allometry and the assumption that self-thinning occurs predictably in forest stands. We designed a DMD for Norway spruce in temperate Europe based on wide-ranging forest inventory data. Quantitative relationships between tree- and stand-level variables that describe resistance to selected natural disturbances were superimposed on the DMD. These susceptibility zones were used to demonstrate assessment and possible management actions related to, for example, windfirmness and effectiveness of the protective function against rockfall or avalanches. The Norway spruce DMD provides forest managers and silviculturists a simple, easy-to-use, tool for evaluating stand dynamics and scheduling needed density management actions.     

Growth and productivity of black spruce in even- and uneven-aged stands at the limit of the closed boreal forest

The increasing commercial interest and advancing exploitation of new remote territories of the boreal forest require deeper knowledge of the productivity of these ecosystems. Canadian boreal forests are commonly assumed to be evenly aged, but recent studies show that frequent small-scale disturbances can lead to uneven-aged class distributions. However, how age distribution affects tree growth and stand productivity at high latitudes remains an unanswered question. Dynamics of tree growth in even- and uneven-aged stands at the limit of the closed black spruce (Picea mariana) forest in Quebec (Canada) were assessed on 18 plots with ages ranging from 77 to 340 years. Height, diameter and age of all trees were measured. Stem analysis was performed on the 10 dominant trees of each plot by measuring tree-ring widths on discs collected each meter from the stem, and the growth dynamics in height, diameter and volume were estimated according to tree age. Although growth followed a sigmoid pattern with similar shapes and asymptotes in even- and uneven-aged stands, trees in the latter showed curves more flattened and with increases delayed in time. Growth rates in even-aged plots were at least twice those of uneven-aged plots. The vigorous growth rates occurred earlier in trees of even-aged plots with a culmination of the mean annual increment in height, diameter and volume estimated at 40–80 years, 90–110 years earlier than in uneven-aged plots. Stand volume ranged between 30 and 238 m³ ha⁻¹ with 75% of stands showing values lower than 120 m³ ha⁻¹ and higher volumes occurring at greater dominant heights and stand densities. Results demonstrated the different growth dynamics of black spruce in single- and multi-cohort stands and suggested the need for information on the stand structure when estimating the effective or potential growth performance for forest management of this species.     

Emulating natural disturbances: the role of silviculture in creating even-aged and complex structures in the black spruce boreal forest of eastern North America

Ecosystem-based forest management is based on the principle of emulating regional natural disturbance regimes with forest management. An interesting area for a case study of the potential of ecosystem-based forest management is the boreal forest of north-western Québec and north-eastern Ontario, where the disturbance regime creates a mosaic of stands with both complex and simple structures. Old-growth stands of this region have multi-storied, open structures, thick soil organic layers, and are unproductive, while young post-fire stands established following severe fires that consumed most of the organic soil show dense and even-sized/aged structures and are more productive. Current forest management emulates the effects of low severity fires, which only partially consume the organic layers, and could lead to unproductive even-aged stands. The natural disturbance and forest management regimes differ in such a way that both young productive and old-growth forests could ultimately be under-represented on the landscape under a fully regulated forest management regime. Two major challenges for ecosystem-based forest management of this region are thus to: (1) maintain complex structures associated with old-growth forests, and (2) promote the establishment of productive post-harvest stands, while at the same time maintaining harvested volume. We discuss different silvicultural approaches that offer solutions to these challenges, namely the use of (1) partial harvesting to create or maintain complex structures typical of old-growth stands, and (2) site preparation techniques to emulate severe soil burns and create productive post-harvest stands. A similar approach could be applied to any region where the natural disturbance regime creates a landscape where both even-aged stands established after stand-replacing disturbances and irregular old-growth stands created by smaller scale disturbances are significant.     

Species Interactions in the Dynamics of Even- and Uneven-Aged Boreal Forests

Many boreal tree stands are neither clearly even-aged nor clearly uneven-aged. The stands may undergo a series of stages, during which an even-aged stand is transformed into two-storied mixed stand, and finally to multistoried or uneven-aged stand structure. The species composition often changes during the succession of stand stages. This study developed models for stand dynamics that can be used in different stand structures and species compositions. The model set consists of species-specific individual-tree diameter increment and survival models, and models for ingrowth. Separate models were developed for Scots pine, Norway spruce, and hardwood species. The models were used in a growth simulator, to give illustrative examples on species influences and stand dynamics. Methods to simulate residual variation around diameter increment and ingrowth models are also presented. The results suggest that mixed stands are more productive than one-species stands. Spruce in particular benefits from an admixture of other species. Mixed species improve diameter increment, decrease mortality, and increase ingrowth. Pine is a more beneficial admixture than birch. Simulations showed that uneven-aged management of spruce forests is sustainable and productive, and even-aged conifer stands growing on medium sites can be converted into uneven-aged mixed stands by a series of strong high thinnings.     

Market impacts of hypothetical fuel treatment thinning programs on federal lands in the western United States

This paper addresses the economics of forest fuel thinning programs on federal lands in the U.S. West, and presents a model of regional timber and product market impacts. The issue of economics is vital to the debate about fire management, and this paper presents market implications of two alternative silvicultural strategies, even-aged and uneven-aged thinning. Projections are based on a regional market model called FTM—West (Fuel Treatment Market model—West), which uses the method of price-endogenous linear programming to project annual market equilibria for softwood timber and wood products in the western United States from 1997 to 2020. The model takes into account variability in tree and log size, as well as economic effects of variable size class on harvest costs, log value, product recovery and mill capacity. Results show large potential market impacts from expanded thinning on federal lands, but impacts vary by silvicultural regime due to differences in size–class distributions of trees available under different thinning regimes. A hypothetical even-aged thinning program (“thin-from-below” strategy) results in net negative market welfare over the projection period (2005–2020), while a hypothetical uneven-aged thinning program (thinning based on stand density index) results in positive net market welfare. Net welfare results are the same over a range of different subsidy and administrative fee assumptions. An implication is that even-aged thinning regimes on federal lands in the U.S. West are less economical and therefore will be less effective.     

Simulated cavity tree dynamics under alternative timber harvest regimes

We modeled cavity tree abundance on a landscape as a function of forest stand age classes and as a function of aggregate stand size classes. We explored the impact of five timber harvest regimes on cavity tree abundance on a 3261 ha landscape in southeast Missouri, USA, by linking the stand level cavity tree distribution model to the landscape age structure simulated by the LANDIS model. Over 100 years, mean cavity tree density increased constantly under all timber harvest regimes except for even-aged intensive management. This was due in large part to the continued maturation of the numerous stands that were >70 years old at the start of the simulations. However, compared to the no harvest (control) regime, the uneven-aged, the mixed, the even-aged long rotation, and the even-aged intensive harvest regimes reduced the cavity tree density by 9–11, 11–13, 15–18, and 28–34%, respectively, as more old stands were cut. Forest managers and planners can use this information to evaluate the practical consequences of alternative timber harvest regimes and consider the need for activities such as cavity tree retention.