THE EFFECT OF METHOD OF SOWING ON THE ESTABLISHMENT OF GRASS SEED CROPS

Methods used by growers to establish seed crops of perennial ryegrass, cocksfoot, timothy and meadow fescue have been examined. While satisfactory establishments were obtained by sowing perennial ryegrass broadcast, cocksfoot, timothy and meadow fescue should be sown in rows, especially if they are to be sown under a cover crop.     

A COMPARISON OF BROADCAST AND WIDE-ROW SPACED GRASSES WHEN MANAGED FOR FOGGAGE PRODUCTION

A small-plot trial comparing perennial ryegrass, cocksfoot, timothy and meadow fescue sown broadcast and also in 21 inch rows was conducted at Auchincruive from 1954 to 1957. The cutting treatments were arranged so that measurement could be made of both summer and winter production.
Of the four species, cocksfoot was the most suitable for foggage production. Perennial ryegrass suffered severely from winter killing. Timothy and meadow fescue, although persistent, did not yield as much as cocksfoot.
Total dry-matter production over three years was higher from the cocksfoot rows than from the broadcast stand. For ryegrass and meadow fescue, broadcasting gave higher yields than row sowing while timothy showed no significant difference between sowing methods.
During the winter, all species showed a loss of dry matter. Losses for ryegrass, cocksfoot, timothy and meadow fescue were respectively 34%, 17%, 18% and 24%. Broadcast stands averaged 11% greater loss than the rows.     

Yield profile of tall fescue (Festuca arundinacea) in comparison with meadow fescue (F. pratensis) in Finland

Yield profile characteristics of tall fescue (Festuca arundinacea Schreb.), cv. Retu, were compared with those of meadow fescue (Festuca pratensis Huds.). The study was conducted in Finland and was based on official variety trial data recorded between 1980 and 1998 at 17 trial sites between latitudes 60° and 66°N. The crops were managed according to silage‐cutting regimes. The pattern of yield formation of the tall fescue cv. Retu differed significantly from that of meadow fescue cultivars, both within a growing season as well as in sward age. Tall fescue cv. Retu established slowly, and the dry‐matter (DM) yield from the first cut, made in the first year of harvest, was significantly lower than that for meadow fescue. In the second and third years, the DM yield from the first cut did not differ between tall fescue and meadow fescue. Tall fescue produced significantly higher DM yield as regrowth (second and third cuts) than meadow fescue. The sward age significantly affected total DM production. In first‐year swards, there were no significant differences in total DM yield between tall fescue and meadow fescue but, in the second‐ and third‐year swards, tall fescue produced significantly higher DM yields than meadow fescue. The difference in yield profile between tall fescue and meadow fescue was similar in all the environments included in the study. DM yield for the first cut (kg DM ha^–1) for tall fescue cv. Retu, in comparison with meadow fescue cultivars, was 2495 vs. 3099 (P < 0·001), 3735 vs. 3741 (NS, P=0·94) and 3553 vs. 3468 (NS, P=0·30) in the first‐, second‐ and third‐year swards respectively. The respective DM regrowth yields (second plus third cut) were: 6059 vs. 5416 (P < 0·001); 5445 vs. 4221 (P < 0·001); and 5580 vs. 4113 (P < 0·001) in first‐, second‐ and third‐year swards. Total DM yields per season for tall fescue vs. meadow fescue were (kg DM ha^–1) 8554 vs. 8515 (NS, P=0·69), 9180 vs. 7962 (P < 0·001) and 9133 vs. 7581 (P < 0·001) in first‐, second‐ and third‐year swards respectively. Over the three‐year sward rotation period, which is common in Finland, tall fescue produced on average 12% higher DM yield than meadow fescue. Both tall fescue and meadow fescue suffered little winter damage in Finnish conditions; the differences between cultivars of the two species were small. The results indicated that tall fescue cv. Retu is a productive and persistent forage species suited to Finnish growing conditions.     

EFFECTS OF WHITE CLOVER ON THE BOTANICAL COMPOSITION OF SWARDS OF RYEGRASS, TIMOTHY AND MEADOW FESCUE AND MIXTURES OF THESE THREE

Results are presented to show the effect of white clover (Trifolium repens) and of applied N on the hotanical composition of swards sown with perennial ryegrass (Lolium perernne), timothy (Phleum pratense), meadow fescue (Festuca pratensis) and mixtures of two and three of these species. White clover had a more favourahle effect on the growth of perennial ryegrass than on either timothy or meadow fescue both in terms of yield and in terms of tiller numbers. Tiller numbers of rough-stalked meadow grass (Poa trivialis) were increased and tiller numbers of Agrostis species were reduced by application of N and by inclusion of clover. Possible reasons for these effects are considered.     

THE EFFECT OF DEFOLIATION ON THE SEED-PRODUCING CAPACITY OF BRED VARIETIES OF GRASSES

Leafy grass varieties S24 perennial ryegrass, S26 cocksfoot, S215 meadow fescue and S51 timothy were cut in October and at various growth stages in spring; S53 meadow fescue was grazed at similar periods. Sub-plots were given different applications of N. The control plots had similar manurial treatments but were not defoliated. The seed yield of S24 following October cutting was significantly lower than the control in the first harvest year and significantly higher in the second; the mean yields over 2 years, as a result of cutting at this stage and before ear formation in March, were very similar to those of the control treatment. The mean yields from 3 harvest years for S26, S215 and S51 were generally improved by cutting in October and before ear formation, but cutting after ear formation reduced the number of heads and the seed yields. Extra N applied in conjunction with the defoliation had no significant effect on the number of seed heads or on seed yield. Double grazing of S53 in Sept. and Dec. reduced significantly its seed production in both harvest years. Application of N in spring gave a greater response than a similar application in autumn.     

THE EFFECT OF DRILL WIDTH ON THE SEED PRODUCTIVITY OF LEAFY VARIFTIES OF THREE GRASS SPECIES

Three leafy grass varieties, S215 meadow fescue, S37 cocksfoot and S51 timothy. were subjected to four dril spacings of 6, 15, 24 and 36 in. under two levels of nitrogen application. Over three harvest years, meadow fescue gave better seed yields with 6- and 15-in, drill spacings. Cocksfoot gave very similar yields from 1.5, 24- and 36-in, drills but a signincantly reduced yield from 6-in, spaced drills. Timothy produced significantly higher yields at the 24 in spacing.
The three varieties showed a significant response to nitrogen application.
Aphids seriously affected the vegetative growth of meadow fescue and timothy at the wider spacings, but the 6-in. drills m boths species remained relatively free of aphid attack, as also did the cocksfoot block.     

Effects of systemic fungal endophytes on the performance of meadow fescue and tall fescue in mixtures with red clover

A symbiosis between grasses and systemic fungal endophytes exists in both natural and agricultural grassland communities. Our objective was to examine the effects of systemic endophytes on the competitive ability of two agronomically important grass species: meadow fescue [Festuca pratensis (Huds.) syn. Schedonorus pratensis (Huds.) P. Beauv] and tall fescue [Festuca arundinacea (Schreb.) syn. Schedonorus phoenix (Scop.)]. Plants of meadow and tall fescue were grown for 48 days in replacement series of interspecific mixture with a legume (red clover, Trifolium pratense L.) in different nutrient environments in a greenhouse. Neither of the grass species gained endophyte‐promoted competitive advantage over red clover in grass–clover mixtures. Endophyte infection increased the growth of meadow fescue monocultures by 89% compared to endophyte‐free monocultures in high‐nutrient soils, but plant competition or the cost of endophyte infection to the meadow fescue decreased the yield in resource‐limited conditions. On average, endophyte‐infected and endophyte‐free meadow fescues produced 0·15 and 0·17 g, and 0·14 and 0·14 g dry biomass per plant in mixtures with red clover in high‐ and low‐nutrient soils respectively. In contrast to meadow fescue, endophyte‐promoted growth of tall fescue monocultures was not detected. Endophyte‐infected and endophyte‐free tall fescue monocultures produced 0·76 and 0·95 g biomass per pot, respectively, in the high‐nutrient environment. Endophyte infection can increase the performance of the host grass, but the positive effects depend on the host species, the species composition and soil nutrient availability.     

Leaf dynamics of timothy and meadow fescue under Nordic conditions

Leaf production dynamics of timothy ( Phleum pratense L. cv. Tarmo) and meadow fescue ( Festuca pratensis Huds. cv. Kalevi) were studied in Finland under field conditions in mixed swards containing timothy, meadow fescue and red clover ( Trifolium pratense L. cv. Björn) harvested by cutting. Fifteen randomly selected tillers of both species were marked and leaf number, leaf appearance rate (LAR), leaf elongation rate (LER), leaf senescence rate (LSR) and leaf live span (LLS) were followed through weekly observations. Concurrently another set of fifteen tillers of each species was dissected on each occasion to determine the number of unemerged leaves and development stage of the apex. The first observation period was between 26 April and 22 June and the second between 22 July and 22 September.
The species differed clearly for most of the measured parameters but most differences were dependent on the season. In spring, timothy started growth earlier, had higher LAR (+57%), produced more leaves (+83%) with higher LER_gross (+58%), higher LSR (+61%), higher LER_net (+54%) and produced larger tillers (+193% in leaf area) than meadow fescue. In autumn, timothy still had a higher LAR (+80%), produced more leaves (+60%) with higher LER_gross (+36%), but had also a higher LSR (+77%) and thus similar LER_net (0%) to meadow fescue. Timothy tillers were only slightly larger than meadow fescue tillers (+20% in leaf area). The consequences of these findings on competition in mixed swards and implications for grazing theories are discussed.     

Some differences between eight grasses within the Lolium-Festuca complex when grown in conditions of severe water shortage

Differences between eight related grasses within the Lolium-Festuca complex in conditions of severe water shortage were investigated. Transparent covers were used to restrict the access of rain to field plots of pure stands of Westerwolds ryegrass ( Lolium multiflorum var. Westerwoldicum), Italian ryegrass ( L. multiflorum ), hybrid ryegrass ( L. multiflorum × L. perenne ), perennial ryegrass ( L. perenne ), meadow fescue ( Festuca pratensis ), Italian ryegrass × meadow fescue, perennial ryegrass × meadow rescue, tall fescue ( Festuca arundinacea ) and three mixtures (Italian ryegrass with perennial ryegrass; Italian ryegrass with tall fescue; perennial ryegrass with tall rescue) grown in west Wales. Westerwolds ryegrass and Italian ryegrass died out after 12–15 months of exclusion of rain; meadow fescue, meadow rescue × ryegrass hybrids and perennial ryegrass died out after ≊ 2 years of exclusion of rain; tall rescue was still alive after 4 years of exclusion of rain. During the 2 years after the year of sowing, the loss of water from the top 1 m of soil was greater with tall rescue than with the other grasses. Tall fescue and perennial ryegrass had a low rate of leaf expansion, a small increase in length of exposed leaf sheath and a high weight per unit area of emerging leaf blade. Tall rescue had a greater number and weight of roots than the other grasses, particularly in the 50–100 cm layer of soil, on an adjoining, uncovered site. The order of suitability of the eight grasses for conditions of severe water shortage appears to be tall fescue > perennial ryegrass > perennial ryegrass × meadow rescue, meadow fescue, Italian ryegrass × meadow rescue, hybrid ryegrass > Italian ryegrass > Westerwolds ryegrass.     

COMPARATIVE PRODUCTIVITY OF HERBAGE VARIETIES ON UPLAND PEAT

The productivity and persistence of 27 varieties of herbage plants were compared, following sowing on a blanket peat after ploughing. Yields were estimated from 3–5 cuts per year for 5 growing seasons. The top yields in each year ranged from 2510 lb/ac. in the 5th year when no nitrogenous fertilizer was applied up to 4700 lb/ac. in the 3rd year when given 174 lb N/ac. Such levels are slightly less than half the level expected on lowland, but livestock output on hill land is far lower than on lowland. Varietal differences were significant, with red fescues, especially S59 smooth-stalked meadow grass, S170 tall fescue, S143 cocksfoot, Yorkshire fog and S53 meadow fescue giving top yields and persistence. Aberystwyth varieties were in all cases superior to other varieties. S50 timothy and S23 perennial ryegrass were persistent but production was low. The conclusions are compared with those of other investigators.     

THE EFFECT OF COMPANION GRASS AND SEED RATE ON THE PRODUCTIVITY OF A TALL-FESCUE SWARD

The effects of sowing from 10 to 50 lb/acre of tall-fescue seed on plant establishment and on the annual and seasonal distribution of DM yield of tall fescue and its companion grasses were measured through two growing seasons. Increasing the seed rate increased plant numbers per unit area and decreased per cent establishment. Adding Scots timothy as a companion grass depressed both plant number and per cent establishment. S37 cocksfoot and New Zealand perennial ryegrass reduced both still further. Total annual dry-matter production was relatively unaffected by varying either the seed rate or the companion grass. The contribution of tall fescue to total production was highest when sown alone and was successively reduced by timothy, perennial ryegrass and cocksfoot. When sown alone there were only slight differences in contribution of tall fescue due to seed rate. With any companion grass the contribution from tall fescue increased with increasing seed rate. Early growth was not reduced by modifying seed rate but was reduced by all the companion grasses. It is concluded that increasing the seed rate of tall fescue above 30 lb is not justified, that Scots timothy can be sown at 3 lb/acre with tall fescue without affecting total production or early growth of the mixture, with the advantages of control of unsown species and improved palatability of the herbage.     

GRAZING MANAGEMENT FOR SEED PRODUCTION IN LEAFY STRAINS OF GRASSES

Leafy strains of five grasses were grown for seed and subjected to various cattle grazing treatments between October and April for three harvest years. The grasses were: S.143 cocksfoot, S.215 meadow fescue, S.170 tall fescue, S.59 red fescue and S.23 perennial ryegrass. All but S.59 red fescue (row crop) were studied as row and broadcast crops. The plots were sown under an arable silage crop and received top dressings of nitrogenous fertiliser every year. Yield of seed, and also quantity of herbage in winter, were measured.
October grazing in the seeding year reduced the first crop of seed in all species except ryegrass. Grazing in December improved the yield of meadow fescue throughout the experiment, and of cocksfoot, tall fescue and red fescue after the first year. Several factors might operate to bring about this effect; suggestions are made for further investigation. Repeated grazing from December to March tended to reduce vigour, and so to offset the advantage of removing autumn-grown herbage. Grazing at intervals from December to late April seriously reduced yield in all species. Tall fescue and red fescue, early flowering species, were most seriously affected, meadow fescue and perennial ryegrass least. Cocksfoot and tall fescue yielded more seed when grown in 2-ft. rows than when broadcast. Meadow fescue and perennial ryegrass did not. The yield of meadow fescue was less affected by adverse conditions than cocksfoot.     

Factors influencing the proportion of natural hybrids between Lolium perenne L. and Festuca pratensis Huds. in permanent pastures

Hybrids between ryegrass and meadow fescue form rarely but occur widely in old pastures. In one experiment six different hybrid genotypes were able to compete, on equal terms, with members of their parental species. In two others the proportion of the total yield due to one hybrid genotype was increased by added fertilizer or by reduced water supply.     

THE EFFECTS OF WHITE CLOVER AND APPLIED NITROGEN ON THE NITROGEN CONTENT OF VARIOUS GRASS/CLOVER MIXTURES

Percentages and yields of N in various grasses and grass mixtures with clover and in associated clover grown at various levels of N application over 5 years are given. Grasses varied consistently in N content, percentages being highest in meadow fescue (cv. S53) and lowest in perennial ryegrass (cv. S24). N application did not significantly affect the N percentage in clover, but the associated grasses had a highly significant effect Percentages were highest in clover associated with meadow fescue and lowest in clover associated with ryegrass. The apparent transfer of N to grass per Ib N in associated clover was calculated for various treatments each year. Reasons for the changing levels of apparent transfer from year to year are considered.     

Establishment and production of common sainfoin (Onobrychis viciifolia Scop.) in the UK. 2. Effects of direct sowing and undersowing in spring barley on sainfoin and sainfoin-grass mixtures

The dry matter (DM) yield and herbage quality of swards of sainfoin ( Onobrychis viciifolia ), meadow fescue ( Festuca pratensis ,) and tetraploid perennial ryegrass ( Lolium perenne ) grown in monocultures and in four sainfoin:grass mixtures (0·33 sainfoin:0·66 meadow fescue, 0·66 sainfoin:0·33 meadow fescue, 0·33 sainfoin:0·66 perennial ryegrass and 0·66 sainfoin:0·33 perennial ryegrass), established by direct sowing or undersowing in spring barley, were investigated over 3 years in a field experiment in the UK. Direct sowing produced a mean yield across all species and mixtures of 1·8 t DM ha⁻¹ in the establishment year, whereas undersowing produced no measurable yield except for that of the spring barley. Undersowing reduced the yields of sainfoin and sainfoin-grass mixtures in the first full-harvest year but not in the second. The annual yield of a monoculture of sainfoin was 7·53 t DM ha⁻¹ and that of sainfoin-grass mixtures was 8·33 t DM ha⁻¹ averaged over 3 years. Both sainfoin and the sainfoin-grass mixtures had higher annual DM yields than the grass monocultures. The mixture of 0·66 sainfoin:0·33 meadow fescue gave the highest mean annual yield (9·07 t DM ha⁻¹) over the 3 years. There was a higher proportion of sainfoin maintained in mixtures with perennial ryegrass than with meadow fescue. The proportion of sainfoin in sainfoin–meadow fescue mixtures declined from 0·62 in the first year to 0·32 in the third year, whereas the proportion in sainfoin–perennial ryegrass increased from 0·48 in the first year to 0·67 in the second year and remained stable in the third year.     

THE INFLUENCE OF SEASONAL CHANGES IN LIGHT ENERGY ON LEAF AND TILLER DEVELOPMENT IN RYEGRASS, TIMOTHY AND MEADOW FESCUE

The importance of leaf area as a determinant of yield in herbage crops is now widely realized. Selection for a rapid increase in leaf area might therefore be a useful approach to the problem of breeding for yield.
The present paper describes the seasonal changes in two important components of leaf area, rate of leaf and tiller production, in contrasting varieties of ryegrass, timothy and meadow fescue. The regular seasonal variation in these components in the glasshouse is shown to be determined primarily by available light energy, the species and varieties differing in their response to changes in light energy. The possibilities of selection for these components of leaf area are discussed, but it is pointed out that individual leaf size must also be considered.     

COMPARATIVE STUDIES OF PERENNIAL RYEGRASS, TIMOTHY AND MEADOW FESCUE

The growth of calves grazing pure swards of perennial ryegrass, timothy and meadow fescue at low grazing pressure was measured. Organic-matter intake was estimated on three occasions during the grazing season. In 1963 rest periods of 7 days in April and May and of 21 days in June-Octoher were used hetween 7-day grazing periods. In 1964 the effect of long (35–55 days) and short (15–25 days) rest periods hetween 3.5-day grazing periods were compared. There were no measurable differences in intake hetween the calves on the three grass species in 1963. In 1964 faecal production was higher with timothy than with the other species and with small differences in digestihility, the intake on this species was higher. The faecal production and intake were higher, hut not significantly so, with the long-rest management in the second year. The liveweight gains of the calves grazing the three species were similar in 1963; in 1964 gains with meadow fescue were lower than with the other species from 13 April to 22 June but this was the only significant difference. The results obtained on slaughtering the calves at the end of each summer are presented.     

Endophyte infection, nutrient status of the soil and duration of snow cover influence the performance of meadow fescue in sub-artic conditions

Neotyphodium endophytic fungi are beneficial for certain cultivated grass species by increasing the stress tolerance and herbivore resistance of their host plants. These grass endophytes have been widely studied in temperate regions. Endophyte-infected grasses are also cultivated in sub-artic conditions, but studies on the effects of endophyte infection on grasses under conditions of prolonged snow cover are scarce. A 2-year field experiment was conducted with two originally endophyte-infected meadow fescue ( Schedonorus pratensis ) cultivars. The growth and reproduction of endophyte-infected (E+) and endophyte-free (E−) plants of two cultivars at two sites, differing in nutrient status and duration of snow cover, were compared at the latitude of the Arctic Circle in Finland. Endophyte infection enhanced plant performance most at the site with the higher nutrient status and the shorter duration of snow cover. The difference in growth between E+ and E− grasses increased during the second growing season and was most pronounced in the reproductive investment of grasses, but the benefits of endophyte infection for reproductive mass were cultivar-dependent. At the site with the lower nutrient status, costs of endophyte infection on the vegetative growth of grasses were found. The results suggest that Neotyphodium infection may benefit meadow fescue in sub-arctic conditions but the extent of the effects is dependent on the cultivar and the environment.     

A comparison of perennial ryegrass‐ and tall fescue‐based swards with or without a cropping component for dairy production: Animal production,herbage characteristics and financial performance from a 3‐year farmlet trial

Perennial ryegrass/white clover swards have some limitations in temperate grazed dairying systems. This study tested the hypothesis that farmlets based on alternative species would be equally or more profitable than those perennial ryegrass‐based, and would produce more herbage in summer‐dry conditions. Six farmlets were established; three with perennial ryegrass‐ and three with tall fescue‐based swards. For each grass species, one farmlet was solely based on grass‐clover swards while the other two had either chicory‐red clover or lucerne crops planted on 20%–25% of the area. Animal‐ and herbage performance‐related variables were measured for 3 years, and calculated financial performance was evaluated. Using tall fescue improved total annual herbage yield compared with perennial ryegrass, but animal production and operating profit were lower. This was likely due to the reduced yield and nutritive value of tall fescue during spring and an associated decline in daily milksolids production. The deficit in spring milksolids production was never recovered, despite greater herbage production from tall fescue during summer/autumn. Incorporating chicory‐red clover or lucerne crops reduced both annual herbage and milksolids production. This reduced farm income, while increasing operating expenses as the farmlets required crop renewal and more purchased supplementary feed to maintain feed supply. Under the conditions of this study (i.e., partial irrigation, high nitrogen supply), changing the forage base from perennial ryegrass to tall fescue did not improve animal production or profitability, nor did incorporation of crops on 20%–25% of the farmlet area.     

Concentration of ergot alkaloids in Czech ecotypes of Lolium perenne and Festuca pratensis

The content of ergot alkaloids (ergovaline and chanoclavine), and their production in October 1996 and during the period May–September 1997, were investigated in seventeen ecotypes of perennial ryegrass ( Lolium perenne L.) and in nineteen ecotypes of meadow fescue ( Festuca pratensis Huds.), naturally infected with Neotyphodium spp. The ecotypes were collected in the north-eastern part of the Czech Republic. In 1996 the content of ergovaline in the ecotypes of perennial ryegrass ranged from 0·00 to 2·73 μg g^–1 dry matter (DM) (one cut), and in 1997 from 0·00 to 4·65 μg g^–1 DM (five cuts). In meadow fescue the content of ergovaline varied from 0·00 to 0·61 μg g^–1 DM (one cut) in 1996, and in 1997 from 0·00 to 2·31 μg g^–1 DM (five cuts). The content of chanoclavine (investigated in 1997 in four cuts only) in perennial ryegrass ranged between 0·00 and 3·39 μg g^–1 DM, and in meadow fescue between 0·00 and 2·26 μg g^–1 DM. Most ecotypes of L. perenne reacted to the high temperature and heavy rainfall in June and July of 1997 with an enhanced production of ergovaline, whereas the content of chanoclavine was not changed. Such reaction to stress conditions was not observed in the ecotypes of F. pratensis. Large differences in the production of both ergot alkaloids between different ecotypes of both plant species were observed.