Plant indicators of wheat and soybean crop water stress

Summary The onset of water stress within a crop is defined as the time at which the rate of water loss declines below that of a well watered crop in the same locality. The relation to the onset of water stress and soil water status of several readily measured plant parameters was investigated in crops of wheat and soybeans over three years. Evapotranspiration ET was monitored with weighing lysimeters. A noticeable decline in the rate of ET for both wheat and soybeans was detected once 20% to 30% of the total plant available water PAW remained in the 1 m deep lysimeter soil profile. Extension growth of wheat declined when PAW was 33% and 34% in two years of measurement. In soybeans, the decline in the rate of leaf extension coincided with the decline in the rate of ET. Midmorning measurement of exposed leaf water potential _L, covered leaf water potential _CL and covered plant leaf water potential _CP yielded similar results for both wheat and soybeans. Day-to-day variability was least in _CP and most in _L. Values of _CP, _L and _CL decreased rapidly with PAW < 30%. Daily values of leaf diffusive conductance were variable but there was a general decline in conductance with PAW < 30%. It is suggested that _CL may be the easiest and most reliable parameter to monitor as a means of detecting the onset of stress. The results indicated that PAW levels in the root zone of 50% for wheat and 30% for soybean probably do not affect extension growth or plant water status parameters and can thus be used as criteria for irrigation scheduling.Seconded from the Water Research Commission, Pretoria; present address: CSIRO, Division of Irrigation Research, Griffith, N SW 2680, Australia     

Relationships between crop temperature,grain yield,evapotranspiration and phenological development in two hybrids of moisture stressed sorghum

Summary Recent studies have shown that the grain yields of corn (Zea mays L.) and wheat (Triticum aestivum L.) are related to the degree of water stress they undergo. The purpose of the study reported here was to establish relationships between crop temperature and the grain yields, phenological development, evapotranspiration rates (ET) and leaf water potential ( _l ) of two hybrids of grain sorghum (Sorghum bicolor L. Moench) subjected to varying levels of plant water stress. The study was conducted at the University of Nebraska Sandhills Agricultural Laboratory in 1978 on a Typic Ustipsamment (Valentine fine sand) soil. The sorghum hybrids used were RS 626 and NB 505. Four irrigation treatments were applied in order to subject the crops to varying levels of water stress during each of three major growth stages. Soil moisture was monitored with a neutron probe. ET was estimated with the water balance technique. Crop temperature was measured with an IR thermometer and leaf water potential was measured with a Scholander pressure bomb. Grain yields were reduced by water stress occuring at anytime during the growing season. Yield reductions were largest when stress occurred during only the grainfill period and were least when stress occurred during the entire growing season. The percentage reduction in sorghum grain yield can be described by an index involving the seasonal accumulation of the daily mid-day temperature differences between well-watered and stressed crops ( TSD). As TSD values increased, ET decreased. However, the correlation of ET with TSD was relatively low (R² = 0.60) probably due to the limited amount of data available for analysis and inaccuracies in the soil water balance method used to estimate ET. The mid-day temperature of well-watered rows ranged between 18.0 and 32.8 °C with a mid-day temperature range of about 0.5 °C between the well-watered rows in various plots for several days following an irrigation. However, in certain instances, the mid-day temperature range increased to 1–2 °C for a few days before irrigation. This suggests that certain of the rows experienced water stress and should have been irrigated earlier. Yield data support that conclusion. Range in crop temperature within a field appeared to be a sensitive indicator of crop water stress in sorghum. No significant difference in the phenological development of sorghum resulted from water stress except in one NB 505 plot in which plants were stressed throughout the entire season. In that plot, the stressed plants lagged in development behind non-stressed plants by approximately ten days. The differences in mid-day leaf water potentials ( _l ) and crop temperatures (T) between stressed and non-stressed vegetation were examined. As T increased up to about 4 °C, _l , also increased. Beyond that point, _l decreased while T continued to increase. This behavior was attributed to stomatal closure which permitted an increase in _l of the stressed plants (hence reducing _l ) even as T continued to increase.Published as Paper No. 6551, Journal Series, Nebraska Agricultural Experiment Station. The work reported was conducted under Regional Research Project 11–33 and Nebraska Agricultural Experiment Station Project 11–50. The work upon which this publication is based was supported in part by funds provided by the Office of Water Research and Technology B-044-NEB, US Department of the Interior, Washington, DC, as authorized by the Water Research and Development Act of 1978. This article was sponsored in part by the Nebraska Water Resources Center, Institute of Agriculture and Natural Resources, University of Nebraska-LincolnResearch Assistant, Associate Professor, Research Assistant, and Associate Professor, University of Nebraska, Lincoln. Contents of this puplication do not necessarily reflect the views and policies of the Office of Water Research and Technology, US Dept. of the Interior, nor does mention of trade names or commercial products constitute their endorsement or recommendation for use by the United States Government     

Effect of soil water osmotic potential on growth and water relationships in barley during soil water depletion

Summary Barley plants (Hordeum distichum, L., cv. Zita) grown in a sandy soil in pots were adjusted during a pretreatment period of 5 days to three levels of soil water osmotic potential by percolating 61 of a nutrient solution with additional 0, 22.3 and 44.6 mM KCl. A drying cycle was then started and the plants were harvested when the soil water matric potential had decreased to –1.4 MPa, respectively 6, 7 and 8 days later.No significant differences in dry matter yields, transpiration coefficients and wilting percentages were found between treatments.During the drying cycle leaf water potential ( _l ) decreased concomitantly with decrease in soil water potential ( _s ) with almost constant and similar differences ( _l – _s ) for all treatments despite differences in levels of potentials. The concomitant decrease in leaf osmotic potential () was due partly to dehydration (58%) and partly to increase in leaf solute content (42%) independent of treatment. The part of total osmotic solutes due to K decreased relatively during the drying cycle.Close relationships were found between and _l as functions of relative water content (RWC). Identical curves for the two levels of salt treatment agree with similar concentrations of K, Cl, and ash found for salt treated plants indicating that maximum uptake of macro nutrients may have been reached.During the main part of the drying cycle the turgor potential as function of RWC was higher and decreased less steeply with decreasing RWC in the salt treated than in the non-salt treated plants.In the beginning of the drying cycle additions of KCI lowered the transpiration rates of the salt treated plants resulting in a slower desiccation of the soil and hence an increased growth period. A delay in uptake from a limited soil water supply may be advantageous during intermittent periods of drought.     

Photosynthesis,leaf conductance,and water relations of cowpea under saline conditions

Summary Cowpea (Vigna unguiculata L.), grown widely under both irrigated and dryland conditions, is well adapted to drought and high temperature and is moderately salt tolerant. Data on photosynthetic response and regulation of water relations in cowpea under salinity stress is lacking. Therefore, in conjunction with a field plot experiment to establish the leaching requirement of cowpea, measurements were made of carbon dioxide assimilation rates (A) by ¹⁴CO₂ uptake, leaf conductances to H₂O (g₁) by tritum uptake, and to CO₂ (g), and leaf total water potential (_t ¹) and osmotic potential ( ¹).Cowpeas, grown in field plots containing Pachappa fine sandy loam (mixed, thermic, Mollic Haploxeraff), were irrigated daily with saline water (1,350 mg 1^–1 total salt concentration) to achieve leaching fractions of 0.17, 0.13, 0.09, 0.07, and 0.02. Cowpea maintained high leaf water potentials, high rates of CO₂ assimilation and high leaf conductances under moderately saline conditions (high leaching). Values of _t ¹ and ¹ for high leaching were consistently 50 to 200 J kg^–1 higher than for low leaching throughout the day. Calculating ¹ at full leaf turgor eliminated diurnal variation in ¹. As leaching decreased, however, A, g₁, and g, decreased significantly. About 45% of the 1°C assimilated by the leaf was incorporated rapidly into ethanol insoluble compounds. The relationship between A and g₁ for cowpea was similar to that reported for other crops.Contribution from the US Salinity Laboratory, USDA-ARS, 4500 Glenwood Dr., Riverside, CA. 92501, USA     

Continuous measurement of plant and soil water status for irrigation scheduling in plum

The usefulness of continuous measurement of soil and plant water status for automated irrigation scheduling was studied in a drip-irrigation experiment on plum (Prunus salicina Black Gold). Two levels of water restriction were imposed at different phenological periods (from pit-hardening to harvest, post-harvest) and compared with a well irrigated control treatment. Soil matrix water potential (_soil) was measured with granular matrix sensors (Watermark); and short-period trunk diameter variation (TDV) was measured with linear variable displacement transformers. The Watermark sensor readings were in reasonable agreement with the irrigation regime and showed a good indication of plant water status across the season (r²=0.62), although they were a better predictor of stem water potential (_stem) in the dry range of _soil. Nonetheless, the most important drawback in their use was the high variability of readings (typical CV of 35–50%). From TDV measurements, maximum daily shrinkage (MDS) and trunk growth rate (TGR) were calculated. Their performance was also compared with _stem, which had the lowest variability (CV of 7%). During most of the fruit growth period, when TGR was minimum, MDS was higher in the less-irrigated treatment than in the control and correlated well (r²=0.89) with _stem. However, after harvest, when TGR was higher, this correlation decreased as the season progressed (r²=0.73–0.52), as did the slope between MDS and _stem, suggesting tissue elasticity changes. Later in the season, TGR was better related to plant water status. These observations indicate some of the difficulties in obtaining reference values useful for irrigation scheduling based exclusively on plant water status measurements.     

Pressure chamber procedures for leaf water potential measurements of cotton

Summary Measurement of leaf water potential ( _l ) with a pressure chamber is usually regarded as a reliable and practical field technique. However, recent evidence indicates that results depend on the measurement techniques employed. Field experiments were conducted to identify the magnitude and sources of error affecting pressure chamber measurements of _l in cotton (Gossypium hirsutum L.) and to develop an accurate and operationally flexible procedure. Water potential of bare cotton leaves was about 0.2 MPa less than aluminum foil wrapped leaves when the elapsed time between excision to chamber pressurization was less than 30 s. The water potential of intact leaves increased 0.3 MPa after 15 s of enclosure in aluminum foil. 5 to 30 min of enclosure were sufficient to reach equilibrium between _l and water potential within the plant stem. Aluminum foil wrapped leaves maintained their _l for 2 h stored in a humid, dark box at 21–28 °C while wet wrapped (cheesecloth) or bare leaf _l increased after one hour because of hydration. An accurate and operationally flexible _l measurement procedure, suitable for large scale sampling, was defined.Contribution from the USDA-ARS, Water Management Research Laboratory, 2021 S. Peach, Fresno, CA 93727 and the Dept. of Land, Air and Water Resources, University of California, Davis, CA 95616, USA     

Yield and vegetative growth as related to plant water potential of cotton irrigated with a moving sprinkler system at different frequencies and wetting depths

Summary Seed-cotton yield, yield components and vegetative growth were determined under different irrigation frequencies and wetting depths with a self-propelled moving-irrigation-system (MSIS) in 1986 and 1987. Irrigation timing was determined in both years by pre-irrigation, mid-day plant water potential (_w). The amount of water to be applied was determined by measuring the soil moisture deficit. In 1987, the effect of a change from one irrigation frequency and wetting depth to another at mid-flowering was also examined. Linear responses of relative seed-cotton yield to the amount of evapotranspiration (ET) were found for both years with similar slopes but different intercepts. Significant positive regressions were obtained between pre-irrigation plant _w and relative seed-cotton yield, and vegetative growth during the linear growth stage. Seed-cotton yield was affected by both wetting depth and pre-irrigation plant _w. The deeper the irrigation the higher was the seed-cotton yield for each pre-irrigation plant _w. Irrigation frequencies which maintained plant _w above -1.5 MPa during vegetative growth, flowering and boll-filling resulted in maximum production. The boll filling stage appeared to be a very sensitive one, as boll weight was found to be the main yield component responding to irrigation treatments. At a wetting depth of 120 cm, higher seed-cotton yields were obtained than at a more shallow wetting. Different irrigation managements resulted in different turgor potentials (_t) mainly during mid-day. Both leaf water vapour conductance and net assimilation rate were sensitive to leaf _w.Contribution from the Agricultural Research Organization, The Volcani Center, Bet Dagon, Israel, No. 2903-E, 1990 series. Research was supported by the U.S.-Israel Binational Agric. Res. and Develop. Fund.     

Salinity and drought

Summary Water deficit (water stress — WS) and excess salt (salt stress — SS) evoke similar plant responses, yet clear differences have been observed. The effect of the two forms of stress applied consecutively to cotton (Gossypium hirsutum) and pepper (Capsicum annuum) was studied in a growth chamber (29/20°C day/night temperature, 50% RH, 12-h photoperiod) in 2.5-liter containers packed with a silt loam soil.Leaf water potential () under increasing WS [soil water potential decrease from –0.16 to –1.10 MPa] of transpiring cotton and pepper plants declined to lower levels than under equivalent SS. The decline of leaf solute potential ₀ on the other hand, was less under WS than under SS, resulting in reduced turgor potential ( _p ), in contrast with turgor maintenance under SS. Predawn turgor potential of WS plants was maintained at all levels of soil water potential. Transpiration, CO₂ assimilation and light period leaf extension rate were higher under low soil water potential produced by salinity than an equivalent value produced by water deficit.The more severe effect of WS was attributed to incomplete osmotic adjustment — the reduction in solute potential did not keep pace with the reduction in leaf water potential, and to increased root interface resistance in the dry soil.The leaf sap of cotton under WS had a higher proportion of sugars (65%) than electrolytes, compared to SS. When WS was converted to SS and plant solute potential decreased, electrolytes were taken up at the expense of a reduction in the sugar concentration. Water stress and salt stress may have an additive effect in depressing growth. But at equivalent levels, the relative magnitude of the effect of low soil matric potential (WS) on plant growth was twice as great as that of low soil solute potential (SS).     

Effects of irrigation regimes on the yield and water use of strawberry

Summary Strawberry plants (Fragaria x annanasa D. cv Chandler) were grown in field plots and in drainage lysimeters under controlled soil moisture regimes. Four irrigation treatments were established by watering the plants when soil water potential reached -0.01, -0.03,-0.05 and -0.07 MPa. The maximum yield was attained at -0.01 MPa soil water potential. Differences in yield were caused by both changes in the number of fruits per plant and in the fresh weight per fruit. Yield reductions were associated with reductions in total assimilation rate resulting from the decreased assimilatory surface area in plants irrigated at lower soil water potentials. The crop water production function calculated on a fruit fresh weight basis resulted in a yield response factor (K _y) of 1.01.     

Crop coefficients and water requirements of irrigated wheat (Triticum aestivum L.) in the Nigerian savannah zone

Summary Daily water use of irrigated wheat (Triticum aestivum L., var. Siete Cerros) was determined for three seasons between 1983 and 1986 using a hydraulic weighing lysimeter at Kadawa in the Kano River Irrigation Project of Kano State, Nigeria. Crop coefficients were determined for various time intervals during each growing season using the lysimeter data and a grass-based reference crop evapotranspiration estimated with Class A pan evaporation data. Mean crop coefficients for each ten-day period of crop growth were then determined. Observed length of a season ranges between 110 and 120 days. Seasonal crop water use ranges between 395 mm and 456 mm. Wheat crop water requirements (CWR) were then estimated for major irrigation projects in the Nigerian Sudan and Sahel savannah zones, between latitudes 10°N and 14°N, using the crop coefficients obtained and long term mean climatic data. The estimated CWR agreed with values obtained from the lysimeter experiments. Seasonal CWR values estimated by designers of three existing irrigation projects in the area agreed with the experimental results, but the designers' short term, ten-day period estimates differed from the results obtained.Contribution from the Irrigation Research Programme, Institute for Agricultural Research, Ahmadu Bello University, Zaria, Nigeria     

Establishment of shallow and restricted root systems in cotton and its impact on plant response to irrigation

Summary The effects of frequent and shallow soil wetting by surface drip irrigation on root growth, morphology, and location, and their impact on plant sensitivity to irrigation management were studied in cotton (Gossypium hirsutum L.). Daily drip irrigation, which wetted the 0 to 40-cm soil depth, encouraged root development mainly around the drippers. Water extraction took place mostly from 0 to 20 cm below the drippers, where the roots were concentrated. Shallowness of root growth was not altered by the expansion and deepening of the wetted soil zone which resulted from an increase in amount of irrigation water. The shallow and restricted root system was characterized by a high fraction of thin roots (less than 1 mm dia.) which comprised almost 90% of the root dry matter. Root proximity to the drippers and the limited amount of water in the rooted soil led to a sensitive and quick response of the plants to small amounts of irrigation. A supply of 1.0 mm H₂O given at midday to 70 day-old plants resulted in a leaf water potential (L _w) increase from –1.64 to –1.32 MPa over a 20-min period. This amount of irrigation comprised 15% of the average daily quantity. A 24 h delay in irrigation to 80 dayold plants was enough to decrease L _w from –1.41 to –2.42 MPa. This decrease was caused by a soil water deficit of less than 6 mm H₂O. Extending the irrigation delay to 72 h affected yield and earliness, although the deficient amount of water was supplied over the several days after the treatment. A strong response to minor, but continuous, differences in the daily irrigation amount was detected. Differences in irrigation of less than 1 mm H₂O per day applied during the whole growth season substantially affected L _w, yield and earliness. It was concluded that the establishment of a shallow and restricted root system resulted in strong dependence of the plants on frequent and sufficient supply of water, and temporary minor changes in irrigation affected plant water status and productivity.     

Water relations,growth and yield of Fino lemon trees under regulated deficit irrigation

Fino lemon trees (Citrus limon L. Burm. fil.) on sour orange (Citrus aurantium L.), growing on a low water retention capacity soil, were submitted to three different irrigation treatments over four years: 100% ETc all year (T-0), 25% ETc all year except during the rapid fruit growth period when 100% ETc was applied (T-1) and 100% ETc all year, except during the rapid fruit growth period when 70% ETc was applied (T-2). A water saving of 30 and 20% was achieved in the T-1 and T-2 treatments, respectively. The plant responses to irrigation treatments were similar in all the years studied. Leaf water potential decreased during deficit irrigation periods in T-1 and T-2 treatments. Larger differences were found in values taken at predawn ( _pd) than at midday ( _md), indicating that _pd is a more useful indicator of plant water status. There was neither osmotic nor elastic adjustment in response to deficit irrigation treatment. A clear separation between the main periods of shoot and fruit growth was found, which can be considered an advantageous characteristic in applying regulated deficit irrigation strategies. Onset of the critical period of rapid fruit growth could be determined precisely by considering the decrease in relative fruit growth rate values. T-2 treatment did not induce a significant reduction in total yield, but it caused a delay in reaching marketable lemon fruit size. T-1 treatment did not affect total yield, with a reduction in yield on the first pick occurring in only one year. Chemical characteristics of lemon fruit were not significantly modified by irrigation treatment.     

Irrigation scheduling in a mature peach orchard using tensiometers and dendrometers

Three trickle irrigation schedules, two of which were scheduled according to soil water potential ( soil) (tensiometer method) and daily stem contraction (DSC) (dendrometer method) respectively and the other one was a schedule of restricted water supply, were applied to a mature peach orchard.The annual water application based on soil was greater than that based on DSC. However, tree growth, fruit size and leaf water potential (leaf) on the trees in the dendrometer scheduling plot did not differ from those in the tensiometer scheduling plot while the premature fruit drop and fruit bud initiation were greatly different. The restricted water supply treatment limited significantly both tree and fruit growth. In addition, the lower leaf was observed on the trees in this plot.Further study shows that use of the dendrometer method for scheduling irrigation satisfies the water needs of the plant and that the tensiometer method is less accurate.Abbreviations leaf leaf water potential - soil soil water potential - DSC daily stem contraction - LVDT linear variable displacement transducer - PET potential evapotranspiration     

Yield and growth responses of kenaf (Hibiscus cannabinus L.) in a semi-arid tropical environment to irrigation regimes based on leaf water potential

Summary The growth response of kenaf (Hibiscus cannabinus L.) to four irrigation schedules based on leaf water potential _l was evaluated in a semi-arid tropical environment. Total dry matter production was unaffected by regimes in which the mean value of leaf water potential _l (mean of solar noon and dawn value) did not fall below –1.26 MPa. Stem elongation was more sensitive than dry matter accumulation to plant water stress. — The economic yield for paper pulp production (i. e. total plant dry matter production minus that of the foliage and upper 60 cm of stem) increased with the frequency of irrigation. — Growth recovery by kenaf following a period of water stress was examined. Alleviation of water stress 10 weeks after irrigation, when _l was –1.60 MPa, produced stem elongation rates that were greater than those of plants previously receiving irrigation. This ability to withstand water stress and partially compensate in growth following alleviation of the stress indicates that the kenaf crop has stress response features suitable for rainfall only production under semi-arid tropical conditions. — Irrigation schedules based on _l resulted in water applications tailored to crop requirements in that water use increased, and the time interval between irrigation decreased, with increasing canopy development as well as with increasing evaporative demand. However, erratic fluctuations in _l between irrigations make scheduling by this method difficult and the use of daily mean, dawn or noon values of _l for scheduling irrigation of kenaf cannot be recommended in environments of high evaporative demand. The factors contributing to these fluctuations in (_l) are discussed.     

Root system parameters determining water uptake of field crops

Summary The distribution of a crop rooting system can be defined by root length density (RD), root length (RL) per soil layer of depth z, sum of root length (SRL) in the soil profile (total root length) or rooting depth (z _r . The combined influence of these root system parameters on water uptake is not well understood. In the present study, field data are evaluated and an attempt is made to relate a daily maximum water uptake rate (WU_max) per unit soil volume as measured in different soil layers of the profile to relevant parameters of the root system. We hypothesize that local uptake rate is at its maximum when neither soil nor root characteristics limit water flow to, and uptake by, roots. Leaf area index and the potential evapotranspiration rate (ET _p ) are also important in determining WU_max, since these quantities influence transpiration and hence total crop water uptake rate. Field studies in Germany and in Western Australia showed that WU_max depends on RD. In general, there was a strong correlation between the maximum water uptake rate of a soil layer (LWU_max) normalized by ET _p and RL normalized by SRL. The quantity LWU_max · ET _p ^-1 was linearly related to (RL/SRL)^1/2. The data show that the single root model will not predict the influence of RD on WU_max correctly under field conditions when water-extracting neighboring roots may cause non-steady-state conditions within the time span of sequential observations. Since the rooting depth z _r was linearly related to (SRL)^1/2, the relation: LWU_max · ET _p ^-1 = f (RL^1/2/z _r ) holds. Furthermore it was found that the maximum specific uptake rate per cm root length UR_max was inversely related to RD^1/2 and to SRL^1/2 or z _r of the profile. Observed high specific uptake rates of shallow rooted crops might be explained not only by their lower RD-values but also by the additional effect of a low z _r . The relations found in this paper are helpful for realistically describing the sink term of dynamic water uptake models.Growing plants extract water from the soil to meet transpiration needs. Rates of transpiration and of water uptake are set by evaporative demand and by plant and soil factors which influence capacity to meet that demand. These factors include crop canopy size and leaf characteristics, root system characteristics and hydraulic properties of the soil and the soil-root interface. Soil and root system properties vary with depth and all factors vary in time, so that parameters related to them require constant updating over a crop season.Dynamic simulation models describe water uptake by root systems under field conditions as a function of soil depth and time. Many of these simulation approaches are based on Gardner's (1960) single root model (Feddes 1981). These simulation procedures follow the assumption that water uptake is proportional to a difference in water potential between the bulk soil and the root surface or the plant interior, to the hydraulic conductivity of the soil-plant system and to the effectiveness of competing roots in water uptake. The effectiveness factor accounts more or less empirically for the influence of various root system parameters on water uptake such as percentage of active roots absorbing water, root surface permeability, root length density determining the distance between neighbouring roots, or total root length and depth of the root system. Such models however, will not always reflect correctly the influence of root system characteristics on water uptake since these assumptions have rarely been tested under field conditions. In many instances, there is better agreement between simulated and measured total water use of plants than between predicted and observed water depletion by roots within individual layers of the soil profile (Alaerts et al. 1985).Water uptake by an expanding root system as a function of depth and time has been studied under field conditions for several crops (listed in Herkelrath et al. 1977a; Feddes 1981; Hamblin 1985). They show that the dynamics of water uptake depend on root length density and the availability of soil water. However, the combined influence of root length density, total root length and rooting depth on the water uptake pattern has not been assessed. An evaluation of root system parameters with respect to soil water extraction should aid our understanding of how roots perform under field conditions and may assist our efforts to formulate the water uptake function of roots in dynamic simulation studies more realistically.The aim of the present investigation is to develop an approach that relates measured water uptake rates to relevant parameters of the root systems. This approach will be confined to situations where water uptake in a soil layer is not restricted by unfavorable soil conditions, such as in wet soil, by insufficient aeration and, in dry soil, by reduced water flow towards roots or by increased contact resistance (Herkelrath et al. 1977b). We will define a maximum water uptake rate WU_max that is neither soil-limited nor appreciably limited by the decreasing permeability of aging roots. This WU_max will be related to relevant root system parameters as they exist when WU_max is observed. Hence, water uptake by roots in a very wet, as well as in a dry soil, has been excluded from consideration.     

Canopy temperature and water stress of cotton crops with complete and partial ground cover

Summary The use of canopy and air temperature differences to compute a crop water stress index (CWSI) for assessing plant water status was investigated using cotton crop canopies that either fully or partially covered the ground. The complete ground cover canopy condition was studied in a well watered moisture regime in a rainout shelter with measurements made on six Texas cotton race stocks. The partial ground cover canopy situation was investigated in a well watered moisture regime of a commercial cotton variety Paymaster 266 grown in the field. The slope of the nonstressed baseline of the CWSI for a cotton canopy with about 50% ground cover was approximately one-half that reported for full canopies. Values of CWSI calculated with theoretical and empirical procedures agreed more closely under a complete canopy condition than under a partial canopy situation. Values of aerodynamic resistance (r _a ) and canopy resistance for well watered soil moisture conditions (r _ep )were estimated in order to use the theoretical procedure of computing CWSI. Values of r _a ranged from 10 to 15 sm^–1 and r _cp from 50 to 60 sm^–1. Both the theoretical and empirical procedures showed much promise, but more information is needed to develop techniques for evaluating r _a and r _cp under differing canopy and environmental conditions.     

Use of stem diameter variations to detect plant water stress in tomato

The sensitivity of stem diameter variations (SDV) measured with linear variable transducer (LVDT) sensors as indicators of plant water status in tomato was evaluated. Two tomato crops were grown sequentially in a sandy loam soil in an unheated plastic greenhouse. These were an autumn–winter tomato crop (autumn crop) and a spring–summer tomato crop (spring crop). One drying cycle of 61 days was imposed to the autumn crop in winter at 92 days after transplanting (DAT). Two drying cycles, each of 29 days, were applied to the spring crop, to young (58 DAT) and mature plants (121 DAT). For each drying cycle, four replicate plots did not receive irrigation, and four were well watered. During each drying cycle, LVDT sensors continuously measured SDV, and daily measurements were made of leaf (Ψ _leaf) and stem water potential (Ψ _stem). SDV data was interpreted using the SDV-derived indices, maximum daily shrinkage (MDS) and stem growth rate (SGR). The response of SDV-derived indices to water deficit differed with (1) climatic conditions during stress imposition and (2) crop age. In the winter drying cycle of the autumn crop, the responses of the SDV-derived indices to soil drying were relatively small and slower than Ψ _leaf and Ψ _stem. Under warmer conditions, the SDV-derived indices were much more responsive to soil drying. In rapidly growing young plants, where SDV was characterized by high SGR and small MDS, SGR was the most sensitive SDV-derived index. In more mature plants with little stem growth, MDS was the most sensitive SDV-derived index. In mature plants grown in warm to hot conditions, MDS (1) responded at a similar time or earlier than Ψ _leaf and Ψ _stem and (2) had larger “signal” values (ratio of values from unwatered to control plants) than Ψ _leaf and Ψ _stem. However, there was appreciably more “noise” (coefficient of variation, CV) associated with the SDV-derived indices, giving lower “sensitivity” values, determined from “signal” to “noise” ratios, than for Ψ _leaf and Ψ _stem. Regression analysis between MDS of well-watered plants and climatic variables gave best results for a linear relationship between MDS and daily maximum vapor pressure deficit. There were strong linear relationships between MDS and Ψ _leaf for each drying cycle. The slopes of these relationships differed with crop age indicating that there was no constant relationship between MDS and Ψ _leaf for a whole season. Overall these results demonstrated that MDS and SGR can be sensitive indicators of the water status of tomato crops under conditions of moderate to high evaporative demand. However, the variability associated with the SDV-derived indices and the changing MDS–Ψ _leaf relationship with crop age represent major issues regarding the development of irrigation scheduling protocols for tomato.     

Crop coefficient, yield response to water stress and water productivity of teff (Eragrostis tef (Zucc.)

In the semi-arid region of Tigray, Northen Ethiopia a two season experiment was conducted to measure evapotranspiration, estimate yield response to water stress and derive the crop coefficient of teff using the single crop coefficient approach with simple, locally made lysimeters and field plots. During the experiment we also estimated the water productivity of teff taking into account long-term rainfall probability scenarios and different levels of farmers’ skills. During the experimental seasons (2008 and 2009), the average potential evapotranspiration of teff ranged from 260 to 317 mm. The total seasonal water requirement of teff was found to lower in contrast to the assumptions of regional agronomists that teff water requirement is comparable to that of wheat and barley (375 mm). The average single crop coefficient values (k_c) for the initial, mid and late season stages of teff were 0.8-1, 0.95-1.1 and 0.4-0.5, respectively. The seasonal yield response to water stress was 1.04, which indicates that teff exhibits a moderately sensitive and linear response to water stress. The results suggest that teff is likely to give significantly higher grain yield when a nearly optimal water supply is provided. The study showed that, in locations where standard equipment is not affordably available, indicative (rough) crop evapotranspiration values can be obtained by using field plots and employing locally made lysimeters. The difference in economic water productivity (EWP) and the crop water productivity (CWP) for teff were assessed under very wet, wet, normal, dry and very dry scenarios. In addition two groups of farmers were evaluated, a moderately (I) and a highly skilled (II) group. The results showed that higher EWP and CWP were obtained under very wet scenario than very dry scenario. There was also a 22% increase in EWP and CWP under group II compared to group I farmers. The increase was due to a 22% reduction in unwanted water losses achieved through use of improved technology and better irrigation skills. Both EWP and CWP can be used to evaluate the pond irrigation water productivity (IWP) for a given climate, crop and soil type, and skill and technology level of the farmer. For special crops like teff extra criteria may be needed in order to properly evaluate the pond irrigation water productivity. During the experimental seasons, a high IWP for teff was attained when about 90% of the optimal water need of the crop was met. IWP can be used as an indicator as how much supplementary irrigation has to be applied in relation to the rainfall and other sources of water supply in order to assure greatest yield from a total area. However, the supplemental irrigation requirement of the crops may vary with season due to seasonal rainfall variability.     

Effects of a shading screen on microclimate and crop water requirements

Despite the steadily increasing area under protected agriculture there is a current lack of knowledge about the effects of the 30% black shading screen on microclimate and crop water requirements. Meteorological and lysimeter measurements inside a screenhouse planted with sweet pepper were compared to external reference data. Irrigation water use efficiency (IWUE) was calculated from yield records Y and water meter readings I _applied. Shading reduced mean global radiation R _G by more than 40%, and the screen transmissivity τ _screen was shown to vary with solar elevation angle β. Wind speed inside the screenhouse u _in was reduced by more than 50%. Crop water requirements ET _c were 38% lower than estimates for an open field crop, suggesting a significant water saving potential when using screenhouses. However, the screen did not significantly modify maximum temperature T _max and daily vapor pressure deficit. The FAO-Penman–Monteith approach based on meteorological measurements in the screenhouse accurately predicted daily crop evapotranspiration, and was in close agreement with lysimeter measurements. IWUE was relatively high (10.7 kg m⁻³ in 2004 and 13.5 kg m⁻³ in 2005), but additional research is required to quantify the effect of shading on yield as well as to determine the water saving potential of other commonly used screens. Contribution no. 603/06 from the Agricultural Research Organization An erratum to this article can be found at     

Comparative performance of different crop production functions for wheat (Triticum aestivum L.)

Summary Experiments were conducted for wheat (Triticum aestivum L.) grown in the lysimeters with controlled irrigation at Berlin (FRG) and in field plots at Hisar (India) under different climatic conditions. Crop production functions, relating crop yield with evapotranspiration with and without the consideration of the time of water deficit during crop growth period, were derived. The derived functions and those previously obtained by different workers were tested against the experimental data. There was a significant correlation of seasonal evapotranspiration with yield that was slightly higher for dry matter than for grain. The correlation, which was quite high for linear functions, was slightly greater for nonlinear functions. The correlation was also slightly improved if evapotranspiration for different growth stages was considered rather than total seasonal evapotranspiration. Since there apears to be no single equation that fits all of the conditions studied and since different equations lead to different conclusions, it is suggested that the crop production functions considering evapotranspiration at different growth stages be used with caution. In general the degree of sensitivity of grain yield and dry matter yield to water stress for wheat cultivar WH 283 grown in field experiments under semi-arid climate decreased in the order of crop growth sub-periods I (Sowing to heading), III (Milk ripe to ripe), and II (Heading to milk ripe). However, the sensitivity of the grain yield and dry matter yield to water stress for wheat cultivar Kolibiri grown in lysimeter experiments under humid climate decreased in the order of the growth sub-periods i.e. I, II, and III.