Effect of trees on the yield of wheat crop

A study was conducted at the National Agricultural Research Center (NARC) of Pakistan Agricultural Research Council (PARC) by planting trees of four different species: Eucalyptus camaldulensis, Albizia procera, Morus alba and Leucaena leucocephala along the boundary of wheat fields in a randomized complete block design. Data on crop yield for each tree species and control (no trees) at different distances viz 2, 4, 6, 8, 10 and 12 meter (m) from the tree bases and control were collected and analyzed. The statistical analysis did not show any significant difference in the wheat yield among different tree species. However, the wheat yield was numerically lowest at 2 m distance in case of all the four tree species and control. In case of mulberry, it was lowest statistically also from other distances. Numerically higher wheat yield values were noted at later distances (8, 10 and 12 m) in case of all tree species including control except for siris where numerically highest value was found at 6 m distance. Therefore, it can be generalized that tree's impact on wheat yield can be experienced up to 2 m distance, there is little, if any, impact up to 6 m distance and almost no impact at 8, 10 and 12 m distances.     

Wheat cultivation in association with Acacia nilotica (L.) Willd ex. Del. field bund plantation — a case study

The paper presents the results of a case study conducted to find out the influence of single row bund plantation of Acacia nilotica var. juquemontii on the growth and yield of associated wheat crop under irrigated conditions in Haryana, India. The indications are that the tree line does affect all crop parameters like height growth, shoot numbers, ear length, grain number and grain yield in the vicinity of trees upto 4 m distance from the tree line and establishes that as the distance from the tree line increases the growth and yield of wheat crop also improves. The effect on wheat crop was found more pronounced in the plots laid out towards the middle of the tree line as compared to plots towards the outer border.     

Improving the Traditional Acacia Senegal-Crop System in Sudan: The Effect of Tree Density on Water Use, Gum Production and Crop Yields

The traditional Acacia senegal bush-fallow in North Kordofan, Sudan, was disrupted and the traditional rotational fallow cultivation cycle has been shortened or completely abandoned, causing decline in soil fertility and crop and gum yields. An agroforestry system may give reasonable crop and gum yields, and be more appealing to farmers. We studied the effect of tree density (266 or 433 trees ha⁻¹) on two traditional crops; sorghum (Sorghum bicolor) early maturing variety and karkadeh (Hibiscus sabdariffa), with regard to physiological interactions, yields and soil water depletion. There was little evidence of complementarity of resource sharing between trees and crops, since both trees and field crops competed for soil water from the same depth. Intercropping significantly affected the soil water status, photosynthesis and stomatal conductance in trees and crops. Gum production per unit area increased when sorghum was intercropped with trees in low or high density. However, karkadeh reduced the gum yield significantly at high tree density. Yields of sorghum and karkadeh planted within trees of high density diminished by 44 and 55% compared to sole crops, respectively. Intercropping increased the rain use efficiency significantly compared to trees and field crops grown solely. Karkadeh appears to be more appropriate for intercropping with A. senegal than sorghum and particularly recommendable in combination with low tree density. Modification of tree density can be used as a management tool to mitigate competitive interaction in the intercropping system.     

Light distribution,photosynthetic rate and yield in a Paulownia-wheat intercropping system in China

Measurements of photosynthetically active radiation (PAR), leaf photosynthesis, canopy leaf area index (LAI) and crop yield were carried from flowering to maturity to study the effects of tree shading on crop yield in a Paulownia-wheat intercropping system in China. We found that the tree shading reduced the amount of incoming PAR within the intercropping system by 22%, 44% and 56% during flowering, grain-filling and maturing, respectively. The amount of PAR intercepted by the wheat crop inside the intercropping system was 34%, 55% and 68% less than the crop outside the intercropping system during flowering, grain-filling and maturing. Estimates of PAR-saturated leaf photosynthetic rate were not affected by tree shading, and the differences between the wheat yield inside the intercropping system and outside the intercropping system can be explained by the difference in the amount of PAR intercepted. Total grain numbers and grain dry weight per 1000 grains were linearly correlated with the amount of PAR intercepted during 7 days prior to anthesis and during anthesis and grain-filling, respectively. As compared with the wheat crop outside the intercropping system, grain numbers and grain dry weight were reduced by 36% and 25%, respectively. As a result wheat yield inside the intercropping system was 51% lower than that outside the intercropping system. Our results suggest a much higher reduction in crop yield and much smaller economic gain of the Paulownia-wheat intercropping system than previous studies on the similar Paulownia-wheat intercropping systems in China.     

Influence of direction and distance from trees on wheat yield and photosynthetic photon flux density (Qp) in a Paulownia and wheat intercropping system

A Paulownia-winter wheat intercropping experiment with the object of quantifying photosynthetically active radiation (PAR) and its effect on wheat yield was conducted 60 km south of Zhengzhou (35°N 113°E), Henan Province, PR China, from September 1991 to July 1992 using a tree and crop interface approach. The middle row of three 240 m long rows of 11-year-old trees was studied for its effects on the yield of irrigated and fertilized winter wheat. Photosynthetic photon flux density (Q_p) was quantified using a split-plot design with four blocks. There were four distance (subplot) treatments (2.5 m, 5 m, 10 m and 20 m) and two direction (main plot) treatments laid out to the east and west of a north-south tree line. Results showed no difference in direction effects but Q_p did affect total grain weight (P = 0.0047) between 2.5 m and 20 m. A regression equation was fit using the mean for each distance treatment: Y = 391.7 + 4.57X with r² = 0.9310 indicating a yield increase of 4.57 g m⁻² (45.7 kg ha⁻¹) over a distance of 2.5 m to 20 m from the trees.     

An evaluation of the Acacia albida-based agroforestry practices in the Hararghe highlands of Eastern Ethiopia

Growing Acacia albida as a permanent tree crop, on farmlands with cereals, vegetables and coffee underneath or in between, is an indigenous agroforestry system in the Hararghe highlands of Eastern Ethiopia. However, there is practically no systematic record or data on the merits and benefits of this practice.The paper presents the results of an investigation into the effects of the presence of A. albida on farmlands on the yield of maize (Zea mays L.) and sorghum (Sorghum bicolor L. Moench). Twenty seven plot pairs each consisting of one plot underneath the A. albida foliage cover and the other in the open, away from the tree-on farmers' fields, in a 40 km radius around the Alemaya College of Agriculture, were sampled and the yield components analyzed. A statistically significant increase in crops yields by 56% on average was found for the crops under the tree canopies compared to those away from the trees. This increase was caused by the improvement in 1000 grain weight and number of grains of plants under the tree, indicating that the trees enhanced the fertility status of the soil and improved its physical conditions in terms of crop growth.Additional benefits from the A. albida trees include supply of fuelwood and fodder. Quantitative estimates of these outputs as well as their monetary values are presented in the paper. However, in order to realize these benefits to a discernible extent, higher stand densities of the tree than at present are required.Based on an enquiry about the farmers attitude towards A. albida, the prospects for an extension of this promising agroforestry technique are discussed against the background of the state and trends of development of agriculture in the area. It is surmised that despite some shortcomings like the relatively slow and highly variable growth of A. albida and a conflict with the spreading cultivation of Ch'at (Catha edulis Forsk.), the prospects of extension of this technique are good. It is recommended that its propagation should be incorporated into the programmes of the extension agencies of the various governmental agencies concerned with land use.     

Productivity and nutrient cycling in young Acacia senegal farming systems on Vertisol in the Blue Nile region, Sudan

Acacia senegal, the gum arabic-producing tree, is the most important component of traditional dryland agroforestry systems in the␣Sudan. The spatial arrangement of trees and the type of agricultural crop used influence the interaction between trees and crops. Tree and crop growth, gum and crop yields and nutrient cycling were investigated over a period of 4 years. Trees were grown at 5 × 5 m and 10 × 10 m spacing alone or in mixtures with sorghum or sesame. No statistically significant differences in sorghum or sesame yields between the intercropping and control treatments were observed (mean values were 1.54 and 1.54 t ha⁻¹ for sorghum grain and 0.36 and 0.42 t ha⁻¹ for sesame seed in the mixed and mono-crop plots, respectively). At an early stage of agroforestry system management, A. senegal had no detrimental effect on crop yield; however, the pattern of resource capture by trees and crops may change as the system matures. A significant positive relationship existed between the second gum picking and the total gum yield. The second gum picking seems to be a decisive factor in gum production and could be used as an indicator for the prediction of the total gum yield. Soil organic carbon, N, P and K contents were not increased by agroforestry as compared to the initial levels. Soil OC was not increased by agroforestry as compared to sole cropping. There was no evidence that P increased in the topsoil as the agroforestry plantations aged. At a stocking density of 400 trees ha⁻¹ (5 × 5 m spacing), A. senegal accumulated in its biomass a total of 18.0, 1.21, 7.8 and 972 kg ha⁻¹ of N, P, K and OC, respectively. Agroforestry contributed ca. 217 and 1500 kg ha⁻¹ of K and OC, respectively, to the top 25-cm of soil during the first four years of intercropping.     

Crop yields of rice and wheat grown in rotation as intercrops with three tree species in the outer hills of Western Himalaya

Studies on tree crop interaction under rainfed condition in Dehradun valley were conducted for 13 years during 1977 to 1990. Grewia optiva (Bhimal), Morus alba (Mulbery) and Eucalyptus hybrid were tried along with rice (CV: Akashi) — wheat (CV: RR-21) rotation. One-year-old tree seedlings of the above tree species were planted in line, 5 m apart in N-S direction, in July 1977, in the middle of the plot (size 20 m × 20 m). Eucalyptus was first harvested in 1987. Grewia optiva, Morus alba and coppice of Eucalyptus were harvested in 1990. All tree species had depressing effect on crop yields. Eucalyptus had maximum effect in depressing crop yield till the first harvest and had least effect thereafter. From 1987 onwards, Morus alba affected rice most, while wheat was mostly affected by Grewia optiva. The depressing effect on an average varied from 28 to 34% depending upon the species.Distance of tree line from the crop significantly affected the crop yield upto a distance of 5 m and there was 39% decrease in crop yield upto 1 m, 33% from 1–2 m, 25% from 2–3 m and 12% from 3–5 m distance. Annual removal of lops and tops from trees partly compensated the deficit. Grewia optiva could produce 1.08 t ha^–1 yr^–1 of branches and 0.26 t ha^–1 yr^–1 of leaves (air dry) and 1.28 t ha^–1 yr^–1 of branches and 0.28 t ha^–1 yr^–1 of leaves were obtained from Morus alba. Wood (ADT) produced by the trees was 33.6 t ha^–1 from Eucalyptus, 9.5 t ha^–1 from Grewia optiva and 11.6 t ha^–1 from Morus alba.     

The effect of young Leucaena leucocephala (Lam) De Wit hedges on the growth and yield of maize,sweet potato and cowpea in an agroforestry system in Sierra Leone

An investigation was undertaken at Senehun — Kamajei, a high rainfall region in Sierra Leone, to assess the effects of Leucaena leucocephala, on the growth and productivity of maize, cowpea and sweet potato. The experiment was laid out in a completely randomised design with four replicates of each treatment. The treatments and controls were: (i) pure crops of maize, cowpea, sweet potato, Leucaena clean weeded and unweeded and (ii) intercrops of Leucaena with the food crops, both (iii) with and without applied fertilizers after the first year.The growth of Leucaena was slow but nevertheless tended to reduce grain yields of maize, and tuber and vine yields of sweet potato in the rows in the immediate vicinity of the trees, especially in the nitrogen-treated plots. The maize, in particular, caused an improvement in the early height growth of the trees.     

Agroforestry for soil and water conservation in the western Himalayan Valley Region of India 2. Crop and tree production

A ten-year-study (1983 to 1992) conducted on nine 15 × 90 m runoff plots at 4% slope compared production efficiency of Leucaena leucocephala and Eucalyptus hybrid based agroforestry as well as monocropping landuse systems in the warm, subhumid climate of the western Himalayan region of India. Treatments for the first sequence were: monocropping systems of leucaena, eucalyptus, Chrysopogon fulvus grass and maize – wheat rotation, and alley cropping systems of grass and crops at 4.5 and 10.5 m alley widths with paired contour tree rows of leucaena and eucalyptus. In the second sequence, alley width increased to 22.5 m in 1989, grass was replaced by turmeric Curcuma longa and paired contour rows of leucaena hedges were introduced in monocropping systems of grain crops and turmeric. Integration of leucaena and eucalyptus trees with crops caused severe reduction of crop yields ranging from 21 to 92% for wheat grain, 59 to 69% for maize grain, 60 to 67% for dry grass and about 50% for turmeric rhizome depending upon the age of trees and alley width. The grain yield of crops stabilized at about 50% reduction with 22.5 m alley width. Total crop biomass (grain + straw) also revealed a similar trend; however, its magnitude of reduction was less severe than for grain. Production of biomass was much lower near the tree rows than in mid alleys. Managing leucaena as contour hedgerows eliminated crop yield reduction in alleys. Performance of grass and turmeric in alleys was not found to be satisfactory. Biomass produced from trees adequately compensated the crop yield reduction. Land equivalent ratios of agroforestry landuses were comparable or even better than monocropping systems indicating suitability of these systems for the western Himalayan valley region. This revised version was published online in June 2006 with corrections to the Cover Date.     

Yield components and phenology of durum wheat in a Mediterranean alley-cropping system

It is often claimed that agroforestry could increase the total productivity per land unit compared to monocropping systems. The aim of this study was to evaluate, in a sub-humid Mediterranean climate, the behavior of the yield components, phenology, LAI and NDVI of durum wheat in an alley-cropping system. Our hypothesis was that the microclimate changes in agroforestry could change the devlelopment and yield of cereals. Two different experiments were carried out: in 2015 under 16-year old poplars in East–West lines and in 2016 under 21-year-old ash trees in North–South lines. In each experiment, 12 genotypes of durum wheat were sown. The grain yield was not significantly different in agroforestry and full sun conditions in 2015; however, both systems in this experiment had a particularly low yield (≈ 10% of the historical average yield of the plot). In 2016, the grain yield was significantly lower in agroforestry in comparison with full sun conditions. In both experiments, the most impacted yield component by agroforestry was the number of grains per spike. Similarly, in both experiments, the number of grains per spike was the only yield component impacted by the position within the alley inside agroforestry. Surprisingly, in 2016 the grain yield was higher in the West than in the center position of the alley. In both experiments, agroforestry delayed the maturity of the crop. The use of standard growing degree days was not sufficient to explain the difference in phenology between agroforestry and full sun conditions.     

Productivity and nutrient uptake of newly released wheat varieties at different sowing times under poplar plantation in north-western India

Standardisation of cultural practises is one of the primary objectives to make the system ecologically sustainable and economically viable. In this context, the present study was conducted to optimise the time of sowing in relation to newly released wheat varieties under 4–6 year old poplar block plantation. Six widely grown wheat varieties (PBW 502, PBW 343, WH 542, PDW 274, PBW 509 and PBW 373) were intercultivated during three consecutive crop growth seasons at three times of sowing (mid November, late November and mid December) under block plantation of poplar (Populus deltoides Bartr.) clone G-48. The grain yield and nutrient uptake of wheat varieties was higher in open conditions than under the trees. The wheat variety PBW 502 out yielded the rest of wheat varieties over different sowing times. The highest grain yield was recorded when crop was sown during mid November over the 3 years. Thus adoption of PBW 502 under poplar plantation would substantially improve the overall productivity of the system without any additional input cost. All the varieties performed better when sown early compared to one-month delay. The various growth parameters like tiller height and number, spike length and 100 grain weight contributed significantly towards higher grain yield in early sown conditions (mid November) under poplar irrespective of its age of plantation. The nutrient uptake (N, P and K) by wheat straw and grain was higher in early sown crop with longer growth span than the late sown crop. The higher quantity of N, P and K was removed from soil by PBW 509, PBW 373 and PBW 343, respectively.     

The spatial and temporal effects of paulownia intercropping: The case of northern China

We present a financial analysis and further statistical tests of the spatial and temporal effects of intercropped paulownia (Paulownia elongata) trees on crop yields. The results provide evidence that growing paulownia trees in farm fields can either increase or decrease crop production, depending on the manipulation of the tree density and rotation length. A 60–100% reduction of crop yields can result from a higher tree density and/or a later stage of the rotation, but the pattern of reduction is different for the first (wheat) and second crops (corn, bean or cotton). However, the reduction in crop yield is accompanied by gains of timber, fuel, fodder, and other goods and services. Thus, although the accumulated net returns from crops decline as tree density increases, the combined net returns of crops and trees are generally higher than those of the control, ranging from 50% to 100%. This revised version was published online in June 2006 with corrections to the Cover Date.     

Moisture extraction and crop yield as a function of distance from a row of Eucalyptus tereticornis

A study was made on the effect of an east-west tree line of years old Eucalyptus tereticornis on soil physical parameters and adjoining mustard and wheat crops as a function of distance from the tree-crop interface in north- and south-facing sides. Maximum soil temperature, pan evaporation, moisture extraction, number of plants/m², and yields of mustard and wheat did not differ significantly on the two sides of plantation. Plant growth parameters decreased linearly with distance from the plantation. Mustard yield, Y (kg/ha) was linearly related to moisture extraction, ME (mm) as Y = 1914–120.3 ME. Eucalyptus tereticornis was found to extract 5 times more water than mustard from 0–150cm soil profile. Mustard and wheat yield reductions in adjoining 10 meter strips on both sides of the plantation were 47 and 34 per cent, respectively. Eucalyptus tereticornis is, therefore, not a suitable species for row plantations in agro-forestry systems in deep water table conditions of semi-arid regions.     

Effects of trees on the yield of irrigated wheat crop in semi-arid regions

A study was conducted in the semi-arid regions of Haryana, in Northern India, to see the effect of Azadirachta indica, Prosopis cineraria, Dalbergia sissoo and Acacia nilotica on the yield of irrigated wheat crop. Data on crop yield for each tree species at different distances (1, 3, 5 and 7 m) and four directions (east, west, north and south) from the tree bases and control (no trees) were collected. Results indicate that A. indica and P. cineraria did not show any significant difference in the wheat yield while the other two species (D. sissoo and A. nilotica) showed a reduction in wheat yield. A. nilotica had the most significant and prominent effect, and a reduction of nearly 40 to 60% wheat yield was observed. The effect of this tree species was observed even beyond the spread of the crown. D. sissoo reduced yield by 4 to 30% but the reduction was only up to a distance of 3 m. In general, the impact of trees on wheat yield was observed up to 3 m distance and there is little, if any, impact up to 5 m distance and almost no impact at 7 m distance. In all the tree species, the wheat yield was reduced to a maximum on the north side of the trees and had almost no effect in the southern direction. Crop maturity was observed to be delayed by three weeks under A. nilotica, by 9–10 days under D. sissoo, and only by 6–7 days under P. cineraria and A. indica.     

The effects of between-row (alley widths) and within-row spacings ofGliricidia sepium on alley-cropped maize in Sierra Leone

The effects of spacings between hedgerows (alley widths) and the spacings of trees within hedgerows ofGliricidia sepium on growth and grain yield of maize were investigated at Senehun in southern Sierra Leone. Four between-row spacings (2, 4, 6 and 8 m) were combined with three within-row spacings (0.25, 0.50 and 1.00 m) in a split block design. Maize, at densities of 20,000, 40,000 and 53,333 plants ha^–1, was established in the alleys and also as pure crops. N, P and K fertilizers were applied to all plots before pruning of the trees began. When pruning started, only the pure maize plots received fertilizer; prunings from the hedgerows were returned to the appropriate alleys in the other plots.Plots with the highest maize populations consistently gave the best yields before pruning started, but lower populations gave improved yields after pruning. Yields of maize increased with increasing alley widths before the start of pruning, after which the narrower alleys of 2 and 4 m outyielded the wider ones by almost double, probably because of the large amount of nutrients applied in prunings. Lack of light limited grain yields before the start of pruning, when there was some shading by the hedgerows. Alleys of 2–4 m wide, planted no closer than 0.50 m within rows, resulted in more than twice the yields of maize than in the 8-m alleys planted at 0.25 m within rows, once the hedgerows were well established and were being managed.     

Fruit production ofAcacia tortilis andA. nilotica in semi-arid Ethiopia

Dry, dehiscent fruits ofAcacia tortilis provide important fodder for pastoral livestock in dry seasons on the central Borana Plateau, fruits ofA. nilotica may also be useful during drought. Information was needed on fruit yield to assess what these species could contribute to improved calf feeding systems based on local resources. Fruit production of 10 mature trees per species was measured at five sites for seven months during 1988–9 (n=50 per species). Fruit yields varied according to site, season and species x site (each atP<0.001), but there was no main effect of species (P=0.13). Yields were not correlated with trunk diameter at breast height (DBH) or canopy area within or across species (P>0.05 in all cases). Yields ranged from 0 to 40 kg DM per tree overall, with an average of 5.3 kg DM per tree (or 65 g DM/m² of canopy area). This average tree had a DBH of 26 cm and a canopy area of 81 m². Low and highly variable fruit yields appear to constrain enhanced use of these species here. These species warrant further attention in research and development, however, given their strategic value as forage resources in pastoral systems and their ability to persist in variable environments.     

Pinus contorta growth in northern Sweden as affected by soil scarification

The aim of this study was to quantify the effects of different soil scarification methods on tree growth. Soil scarification influenced stem volume and stem biomass yield of lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) in a 17-year-old field trial in boreal Sweden. Soil scarification (disc trenching, mounding and ploughing) resulted in an average stem volume yield of 3.1 and 34.2 m³ ha^–1 on the poor and intermediate sites, respectively, while corresponding values for no soil scarification were 0.9 and 16.7 m³ ha^–1. In comparison to no scarification, ploughing increased volume yields by 500% on the poor site and by 200% on the intermediate sites. The ranking according to stem volume yield was ploughing > disc trenching = mounding no soil scarification. Averaged over the two sites, the mean annual increment of stem biomass was 219% and 145% higher (in d.w., 0.26 kg and 0.34 kg per sample tree) after ploughing compared with no soil scarification, for the average and dominant sample trees, respectively. Although not significant, the increased growth rate after soil scarification decreased the average stem basic wood density of the sample trees with 1.6% and 5.3%, at the poor and intermediate sites, respectively. In conclusion, soil scarification significantly increased the 17-year stem volume yield compared with no scarification. The results also indicate that the difference in stem biomass yield between ploughing and the other methods, especially no soil scarification, will increase even more in the near future.     

Trees on farms in Bangladesh: 5. Growth of top- and root-pruned trees in wetland rice fields and yields of understory crops

Growth of trees and seasonal yields of understory crops were measured over a 5-year period for 4 crops grown under 17 tree species at 8 × 8 m spacing in wetland rice fields. All tree species grew well in rice fields, at rates comparable to their growth in forest plantations. Top and root pruning reduced average tree girths by up to 19% and average tree volumes by up to 41%, depending on intensity of pruning. The crops monitored were Oryza sativa, Triticum aestivum, Corchorus olotorius, and Lens culinaris. Crop yields under the trees averaged 93% of the corresponding yields outside the tree canopy. The most important factor affecting the yields of undercrops was tree size (height and/or girth). Differences among tree species and the interaction with species of undercrops were not significant after controlling for tree size. Pruning of roots and branches significantly improved crop yields under trees by amounts proportional to the intensity of root or top pruning. This revised version was published online in June 2006 with corrections to the Cover Date.     

Growth and yield of maize and timber trees in smallholder agroforestry systems in Claveria,northern Mindanao,Philippines

On-farm experiments were conducted in the Philippines to study over a 4-year period the growth of two timber trees, gmelina (Gmelina arborea R. Br.) and bagras (Eucalyptus deglupta Blume), and their impact on the grain yield of intercropped maize. The experiment consisted of maize monocropping plots (control) and maize intercropped between trees planted in block (2 × 2.5 m), and hedgerow arrangement (1 × 10 m). Three maize crops were planted in the block plots before canopy closure, and seven maize crops were planted in the hedgerow and monocropping plots. Maize grain yield in the hedgerow and in the block arrangement with gmelina were respectively 37% (16.58 tons ha⁻¹) and 68% (8.3 tons ha⁻¹) lower than in monocropping (26.21 tons ha⁻¹). In the plots with bagras, maize grain yield in hedgerow and in block arrangement were respectively 19% (24.8 tons ha⁻¹) and 66% (10.4 tons ha⁻¹) lower than in monocropping (30.6 tons ha⁻¹). For both tree species, the diameter at breast height (dbh) was greater in hedgerow than in block arrangement, with the difference being more pronounced with age. It was estimated that gmelina planted in hedgerows would produce 6–8 m³ ha⁻¹ of merchantable volume more than if planted in block. The study verifies the hypothesis that intercropping between widely-spaced trees rows (planted at 10 m or more) is more profitable and feasible to smallholders than either maize monocropping or woodlots, and concludes with recommendations on how to further improve the productivity of tree-intercropping systems.