South-west England grassland survey 1983

A survey of grassland on 127 farms in south-west England was conducted in order to investigate the cutting and grazing management of grassland, botanical composition and the use of fertilizer N, and to quantify recent trends in reseeding and the age structure of swards. Twenty-six percent of the grassland surveyed was classified as arable grassland (in rotation with crops) and 74% as permanent grassland. Only 37% of the established grassland on dairy farms was aged over 20 years compared with 53% on livestock farms. Mowing was practised on 52% of the grassland, two-thirds of it every year, and virtually all grassland was grazed for part of the year. The average fertilizer N input was 168 kg ha⁻¹ with considerable variation with sward age (average 113 kg ha⁻¹ for over-20-year-old swards) and between dairy farms (average 217 kg ha⁻¹) and livestock farms (100 kg ha⁻¹)- The average proportion of sown species (mainly Lolium perenne) ranged from 90% in young swards to 36% in swards aged over 20 years, with considerable variation within age groups. L. perenne was associated with high inputs of fertilizer N, good drainage, sheep grazing and organic manures, e.g. among swards aged over 20 years it contributed 38% on average where >250 kg N ha⁻¹ were applied, but only 12% where no fertilizer N was applied. A continuous network of Trifolium repens occurred in 27% of the grassland–this was associated with low inputs of fertilizer N, good drainage and sheep grazing. Agrostis spp. were the most abundant unsown species and Cirsium arvense, Ranunculus spp. and Rumex spp. the most common broad-leaved weeds. Features of the sample are compared with those in the south-west England region and in England and Wales as a whole.     

The comparative performance of Holcus lanatus and Lolium perenne under sheep grazing in the Scottish uplands

An experiment was conducted over 3 years (1983-85) to assess the performance of Holcus lanatus German Commercial and cv. Massey Basyn compared with Lolium perenne cv. Perma on a gley soil under sheep grazing in the Scottish uplands. All grasses were sown together with Trifolium repens cv. Grasslands Huia. The swards were rotationally grazed at similar herbage allowances. During the first harvest year, the swards were grazed hard to a low mass (500 kg DM ha⁻¹). In the second harvest year, post-grazing herbage masses of 500 and 1000 kg DM ha⁻¹ were compared.
Perma ryegrass had a higher level of herbage production than both H. lanatus cultivars in the first and second harvest years after sowing and hence had a greater number of sheep grazing days. The ryegrass sward consisted of 25% more green sown grass (85 cf. 60%) but 12% less white clover (4 cf. 16%) compared with both H. lanatus cultivars in the establishment and first harvest years. The persistence of all three grasses was poor although ryegrass had a higher presence (36%) than either Massey Basyn (22%) or German Commercial (13%) at the end of the. second harvest year.
At a similar herbage allowance, there were no significant differences in the herbage intake and liveweight gain of sheep. Ryegrass had a higher organic matter digestibility and lower neutral and acid detergent fibre and lignin contents than either of the H. lanatus cultivars.
In the second harvest year, although herbage production was greater at the higher herbage mass, there was no difference in the proportion of sown grass.
It was concluded that ryegrass is a superior grass to H. lanatus on upland soils with high N status, moderate P status and a high pH.     

Effects of field drainage on the growth of herbage and the liveweight gain of grazing beef cattle

A long-term field grazing experiment was begun in 1982 to examine the impact of efficient field drainage on herbage and animal production from swards on an impermeable clay loam in the south-west of England. Drained and undrained lysimeter plots (each of 1 ha) were established on the existing permanent sward and received annual applications of fertilizer N of 200 or 400 kg ha⁻¹. Similar plots were initially ploughed and reseeded with Lolium perenne (cv. Melle), and received fertilizer N at an annual rate of 400 kg ha⁻¹. All plots were continuously stocked by beef cattle and stock numbers were adjusted to maintain a constant sward height and to avoid poaching damage. Results for the first 5 years show that the benefits from drainage were modest and, for beef production, unlikely to pay for its costs over the shorter term. The main benefit was in spring when herbage dry matter yield was 11% greater on the drained plots, but with no significant interaction with fertilizer N level or sward type. This benefit was reduced to 3% on an annual basis, due to the effect of the larger soil water deficits sustained by the drained swards in mid-season. Drainage increased the annual liveweight gain per ha by the grazing cattle by 11%. Possible mechanisms accounting for these effects are discussed in relation to the influences of seasonal patterns of weather.     

The effects of change in herbage mass on rates of herbage growth and senescence in mixed swards

A study was conducted to determine the influence of a change in herbage mass on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in swards of Lolium perenne L., Poa annua L. and Trifolium repens L. under continuous stocking with sheep. Swards were maintained at either high (1700–1900 kg OM ha⁻¹; H) or low (700–900 kg OM ha⁻¹; L) herbage mass from April to 20 August. From 20 August to 13 September half of the L sward was permitted to grow until a herbage mass similar to that of the H sward was achieved (LH), and at the same time half of the H sward was grazed down until a herbage mass similar to that of the L sward was achieved (HL). The other half of each sward was held at constant mass (LL and HH). Estimates of G, S and NP were obtained for each plant species over a 2-week period from 17 September to 1 October and estimates of species population densities were also made.
The combined species rates of G and S per unit area were highest in treatment LH and lowest in treatment HL, whilst the rates for the LL and HH swards were intermediate. The rate of NP was similar in the LL, HH and LH treatments (34.5, 29.3, 33.6 ± 6.2 kg DM ha⁻¹d⁻¹) but was lower in the HL sward (100 ± 6.2). The effects of alteration of herbage mass on individual tiller or stolon G and S rates were rapid but population density changes were slower.
Within the limited conditions of this trial it was not possible to increase NP by manipulation of herbage mass under continuous stocking management but the results indicate that NP can be reduced in the short term if a sward of high herbage mass and low population density is grazed hard.     

Response of permanent and reseeded grassland to fertilizer nitrogen. 2. Effects on concentrations of Ca, Mg, K, Na, S, P, Mn, Zn, Cu, Co and Mo in herbage at a range of sites

The concentration of mineral elements in herbage from permanent swards of mixed species composition (predominantly Agrostis spp., Holcus lanatus, Festuca rubra, Lolium perenne, Poa spp., etc.) was compared with that from sown perennial ryegrass ( Lolium perenne ) at two fertilizer rates (0 and 300kg N ha⁻¹ year⁻¹, both with P and K) and with L. perennel Trifolium repens (at 0 N with P and K). The investigation was conducted over four years at sixteen contrasting sites in England and Wales. Herbage samples for mineral analysis were taken at first and fourth harvests (May and August) each year from plots under 4-weekly cutting. There was a wide degree of variation between sites, particularly for concentrations of Ca, Mg, Na and all trace elements analysed. Herbage from L. perenne reseeds had significantly higher concentrations of Ca, Mg, Na, Co and S, but K, Mo, Zn, Cu and P were generally lower than in the permanent swards. L. perennel T. repens swards had the highest concentration of major cations, but trace element levels were generally intermediate between those for 0 N permanent and L. perenne swards. Fertilizer N reduced concentrations of Ca, Mn, Mo and S, and increased Mg, Na and Zn, with no consistent effects on K, Co and Cu. Herbage at harvest 4, compared with harvest 1, had higher Ca, Mg, Na, Mn, Zn and S. Permanent and L. perenne swards responded similarly to fertilizer N and between harvest dates. Results are discussed in relation to previous reports of botanical and fertilizer effects on mineral concentrations, and the mineral requirements of livestock. Changes in mineral concentration resulting from reseeding permanent swards are considered unlikely to increase mineral-related livestock disorders, and in many cases could be beneficial, though problems might arise on sites where some elements, e.g. Cu and Mg, are low.     

The effects of stocking rate and nitrogen fertilizer on a perennial ryegrass-white clover sward

The effects of stocking rate and N fertilizer on a mixed sward of perennial ryegrass ( Lolium perenne) and white clover ( Trifolium repens ) set-stocked with sheep were examined. Sward production and composition, and sheep production were studied.
Increasing the stocking rate over the range 25–55 yearling sheep ha⁻¹ reduced herbage accumulation by about 40%, whether or not N fertilizer was applied. Increasing the stocking rate increased the density of ryegrass tillers, but reduced the density of clover stolons and the clover content of the swards. Applications of N fertilizer (200 kg N ha⁻¹ a⁻¹) increased herbage accumulation by about 20% but substantially reduced the clover content.
Liveweight gain per animal and per unit area were greater at the lower stocking rates where the clover content and nutritive value of the diet were greatest. Wool growth per unit area was greater at the higher stocking rates. Applications of 1M fertilizer increased liveweight gain at stocking rates above 25 sheep ha⁻¹, but had no effect on wool production at any stocking rate.
The results demonstrate that a stable and productive grass-clover association was maintained under conditions of set-stocking at around 23 yearling sheep ha⁻¹and that at this stocking rate, which appears to be about the biological optimum, there was no advantage in using N fertilizer.     

Herbage productivity of a range of grass species in association with white clover

Red fescue ( Festuca rubra L.) cvs Reptans, Dawson and Ruby, perennial ryegrass ( Lolium perenne L.) cvs Perma and Sprinter, crested dogstail ( Cynosurus cristatus L.) cv. Southlands, smooth meadow-grass ( Poa pratensis L.) cvs Parade and Arena, Yorkshire fog ( Holcus lanatus L.) commercial types I and II, creeping bent ( Agrostis stolonifera L.) cv. Penncross and rough meadow-grass ( Poa trivialis L.) cvs Omega Øtofte, Dasas and Sabre were each sown with white clover ( Trifolium repens L.) cv. Donna and the productivity of the fourteen mixtures assessed under simulated grazing (six harvests per year) for 3 years. A single fertilizer-N rate (50 kg ha⁻¹) in early spring was applied to all treatments each year.
Over the 3 years, mean annual dry matter (DM) production of the swards ranged from 5·25 t ha⁻¹ (creeping bent) to 8·71 t ha⁻¹ (red fescue cv. Reptans), with the species ranking order of: red fescue > perennial ryegrass > crested dogstail > smooth meadow-grass > Yorkshire fog > creeping bent. Rough meadow-grass only persisted over the first harvest year. Mean annual organic matter digestibility (OMD) values of the swards ranged from 0·712 (creeping bent) to 0·782 (ryegrass cv. Sprinter). Mixtures with the lowest proportion of grass (crested dogstail and smooth meadow-grass) generally gave the highest proportions of white clover. The creeping bent mixture had exceptionally low production of both components.
Red fescue, crested dogstail and smooth meadow-grass showed compatibility with clover and, therefore, have potential for use in grass/clover swards. Yorkshire fog had lower compatability with clover. Some breeding effort to improve selected characteristics of promising species, e.g. improvement of low OMD in red fescue, seems warranted.     

Assessment of contrasting perennial ryegrasses, with and without white clover, under continuous sheep stocking in the uplands. 3. Herbage production, quality and intake

Herbage characteristics were monitored over 3 years (1985–1987) in three perennial ryegrass ( Lolium perenne L.) varieties as grass/only (200 kg N ha⁻¹) and grass/clover ( Trifolium repens L.) (75 kg N ha⁻¹) swards when continuously stocked with sheep. Although mean total annual herbage production was similar from the grass varieties, growth of Aurora, a very early-flowering variety, was higher than that of Aberystwyth S23 and Meltra (tetraploid) late-flowering types in spring. Herbage production from grass-only swards was 15% higher than from grass/clover swards.
In vitro organic matter digestibility (OMD) of Aurora and Meltra was 34 g kg ⁻¹ and 26 g kg ⁻¹ higher ( P < 0·01) than that of S23. Averaged over varieties, the OMD of grass/clover was 25 g kg ⁻¹ higher ( P < 0·05) than grass-only during the post-weaning period. Individual lamb digestible organic matter intake (DOMI) was 22% higher ( P < 0·05) on grass/clover than on grass-only during this period. Water-soluble carbohydrates content of Aurora was 42% higher ( P < 0·001) than that of both Meltra and S23.
Herbage characteristics were related to lamb performance reported previously. Lamb production was positively correlated with DOMI. However, the magnitude of differences in lamb performance between treatments was much greater than the extent of differences detected in herbage characteristics.     

Frequency of stocking rate adjustments required on contrasting upland perennial ryegrass pastures continuously grazed to a sward height criterion from May to July

The use of sward height as a criterion for determining the time and extent of stocking-rate changes on continuously grazed swards was investigated over a 2-year period (1985–86) in a sheep production experiment. Swards of three contrasting perennial ryegrass ( Lolium perenne L.) varieties were established with and without Aberystwyth S184 small-leaved white clover ( Trifolium repens L.) at an upland site (310–363 m) in mid-Wales. From spring (late April) until weaning (mid-July) the pastures were continuously stocked with Beulah Speckled Face ewes and Suffolk cross lambs. During this period sward heights of 4 ± 0.5 cm were obtained and maintained by regular adjustment of animal number on individual paddocks. Grass-only swards received 160 and 200 kg N ha⁻¹ and the grass clover swards were given 80 and 75 kg N ha⁻¹ in 1985 and 1986 respectively.
Differences were observed between the treatments in sward height profiles over the season necessitating contrasting adjustments to stocking rates. Mean stocking rate necessary on early flowering Aurora (22 6 ewes ha⁻¹) was respectively 27% and 17% higher than on late-flowering Aberystwyth S23 and Meltra (tetraploid) ryegrasses; mean stocking rate on grass-only swards was 19% higher than on the grass-clover pastures.
It is concluded that sward height is a useful criterion on which to make adjustments to stocking rates to compare the potential performance of contrasting swards, under continuous grazing. The infrequent adjustments required to maintain a constant sward height, especially on the late flowering diploid perennial ryegrass variety on which many upland pastures are based, suggest that the criterion of sward height could be successfully employed on farms as an aid to efficient grazing management.     

Dry matter and quality of herbage harvested from Holcus lanatus and Lolium perenne grown in monocultures and in mixtures

The herbage dry matter (DM) harvested, N response and herbage quality of monocultures and mixtures of Holcus lanatus (local ecotype) and Lolium perenne (cv. S24) were compared in a field experiment. The species were seeded in the proportions 100:0, 75:25, 50:50, 25:75 and 0: 100; three rates of N were applied (150, 300 and 450 kg ha⁻¹) and five or six cuts were taken in each of four harvest years.
H. lanatus established poorly in the seeding year and in the first full harvest year L. perenne contributed 1, 54, 76, 89 and 99% of the total herbage DM harvested from the five sward types (mean of N treatments). Only a small decline in the L. perenne content of the swards occurred during the experiment.
Total amounts of herbage harvested were significantly lower for H. lanatus monoculture than for the other sward types in three of the four harvest years However, L perenne monoculture yielded significantly more total DM than mixtures with H lanatus in only one of the four harvest years. The five sward types did not differ overall in response to N fertilization. Nitrogen, phosphate and potassium content of the herbage was greatest for H. lanatus monoculture and decreased with increasing proportions of L perenne present in the mixtures.
Thus the presence of a high proportion of H. lanatus in mixture with L. perenne cv. S24 had     

Effects of spring fertilizer nitrogen and sward height on production from perennial ryegrass/white clover swards grazed by beef cattle

An experiment was carried out to compare the effects of two compressed sward height treatments, each at two fertilizer nitrogen (N) treatments (0 and 50 kg ha⁻¹ in spring), on the date of turnout and liveweight gain of steers grazing a perennial ryegrass/white clover ( Lolium perenne/Trifolium repens ) sward sited on clay loam in south-west England. The sward height treatments were 6 cm all season, and 4 cm in the spring rising to 6 cm in June; these were maintained using continuously variable stocking with Hereford × Friesian steers. Cattle were turned out on average 11·5 days earlier on the 4-cm sward height compared with the 6-cm sward height treatment. Liveweight gain early in the season was lower on the 4-cm swards than on the 6-cm swards. Liveweight gain ha⁻¹ over the whole season was similar for the two sward height treatments. Fertilizer N did not affect turnout date or liveweight gain.     

Animal production evaluation of herbage varieties. 1. Comparison of Aurora with Frances, Talbot and Melle perennial ryegrasses when grown alone and with clover

Comparative herbage characteristics and sheep production (ewes and lambs until weaning and lambs thereafter) from swards of Aurora (very early flowering), Frances (early flowering), Talbot (intermediate flowering) and Melle (late flowering) varieties of perennial ryegrass ( Lolium perenne ) were studied under a continuous variable stocking management based on sward surface height guidelines. The varieties were assessed as grass-only (215 kg N ha⁻¹ fertilizer) and grass/clover (83 kg N ha⁻¹ fertilizer) pastures.
Over 2 years (1985-86) total annual lamb production per hectare from grass-only swards of Aurora was 19% more than that from Frances despite similar herbage productivity. Differences between the varieties in lamb output were more pronounced as grass/clover swards, with Aurora producing 29 and 18% more lamb than Melle and Talbot respectively and with Frances also giving 16% more than Melle. Overall lamb production from grass/clover swards was 10% more than that from the grass-only pastures, mainly due to 69% advantage in individual lamb growth rates after weaning. Herbage organic matter digestibility, during the post-weaning period, was higher on grass/clover than on grass-only swards, and similarly with Aurora versus the other varieties. Under a frequent cut simulated grazing regime the relative herbage productivity of the four varieties differed, with Aurora 13% less productive than Frances.
The results are discussed in relation to the limitations of assessing performance under cutting, and the significance of the interaction in animal productivity between the two sward types.     

Assessment of contrasting perennial ryegrasses, with and without white clover, under continuous sheep stocking in the uplands. 5. Herbage production, quality and intake in years 4–6

Herbage characteristics were studied over years 4–6 (1988–90) in three perennial ryegrass ( Lolium perenne L.) varieties as grass-only (200 kg N ha^-1) and grass/clover ( Trifolium repens L.) swards which received 75kg N ha^-1 in 1988 and 0kg N ha^-1 in 1989 and 1990 when continuously stocked with sheep. Mean total annual herbage production of Aurora, a very early flowering variety, was 11% more than that of late-flowering Aberystwyth S23 due to 21% higher growth as grass/clover pasture. The grass/clover sward of Meltra, a tetraploid late-flowering variety, out-yielded S23/clover by 17%. Herbage production of grass/clover was 86% of that of grass only in 1988 but only 54% of the grass-only swards averaged for 1989 and 1990.
In vitro organic matter digestibility (OMD) of Meltra was 38g kg^-1 OM and 27g kg^-1 OM higher than that of S23 and Aurora respectively. OMD of grass/clover was 15g kg^-1 OM higher than that of grass only during the post-weaning period. Herbage intake was positively correlated with OMD of herbage.
The herbage attributes were related to lamb performance reported previously. Lamb output was positively correlated with intake of digestible organic matter.
Differences between the three varieties in herbage characteristics were greater as grass/clover than as grass-only swards, reflecting their compatibility with white clover. In this respect Meltra was the best and S23 the poorest variety.     

The utilization of N fertilizer, applied to perennial ryegrass/white clover pasture growing on a humus iron podzol in N.E. Scotland

Applications of N fertilizer (0.40, 80, 120 and 240 kg N ha⁻¹ year⁻¹ in dressings of 40 or 80 kg N ha⁻¹) were made to a perennial ryegrass/white clover ( Lolium perenne L. Trifolium repens L.) pasture growing on a humus iron podzol reclaimed from heather ( Calluna vulgaris L.) moor in 1982 and 1983. Where no N was applied, estimates of Ni fixation and mineralization were almost equal, being approximately 50 kg N ha⁻¹ year⁻¹ from each source. Apparent efficiencies of fertilizer use were generally low; for each dressing they ranged between –0·7 and 25·5 kg dry matter (DM)kgN⁻¹. Also, responses to N fertilizer were affected by previous dressings. The net N recovery in harvested herbage from application of 120 kg N ha⁻¹ year⁻¹ was 30 kg N ha⁻¹ year⁻¹.     

The influence of sward condition on rates of herbage growth and senescence in mixed swards under continuous stocking management

A study was conducted to determine the influence of herbage mass under continuous stocking management with sheep on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in a mixed-species sward of Lolium perenne. Poa annua and Trifolium repens. Plots were maintained in as nearly constant herbage mass as possible at 500, 700, 1000 and 1700 kg organic matter (OM) ha⁻¹ by continuous but variable stocking with sheep for the period May to July inclusive. Estimates of G, S and NP were obtained for each species from repeated measurements over two 2-week periods on individually identified grass tillers or clover stolons. Estimates of species population densities enabled estimates of G. S and NP per unit area to be made.
Rates of G were higher and of S were lower for Lolium tillers and Trifolium stolons than for Poa tillers. The laminae of Poa tillers occupied inferior positions in the canopy of the swards maintained at higher herbage masses and were less accessible to the grazing animals, so that Poa tissue was less well utilized.
The total population densities of tillers and stolons declined at herbage masses both above and below 700 kg OM ha⁻¹. The rate of G for the three species combined increased with herbage     

1986 survey of upland grassland in England and Wales: changes in age structure and botanical composition since 1970–72 in relation to grassland management and physical features

A sample of 10000 ha of grassland in seven upland districts originally surveyed in 1970–72 was re-surveyed in 1986. Forty percent of the area was above 240 m elevation, 38% had gradients above 81/2°, 29% had poor or bad drainage and 54% was classified as having difficult or severe limitations to management. Twelve percent had been reseeded within the previous 5 years and 39% within 20 years; 6% was classified as arable grassland (in rotation with crops). Thirty-four percent was usually mown, two-thirds of it for silage. In most years 72% of grassland received fertilizer N (average application 123 kg N ha⁻¹) and 35% received organic manure; nearly 20% received neither. Older swards tended to receive lower rates of fertilizer N than reseeds, hay fields and grazed fields less than silage fields, and fields on beef/sheep farms less than those on dairy farms. The proportion of sown species in swards averaged 35%, of which Lolium perenne accounted for 23%. A good contribution of Trifolium repens was recorded in 15% of the grassland. Herb species were recorded in 30% of the grassland, mostly in situations with low fertilizer N under hay or grazing. Agrostis spp., Poa spp., Holcus lanatus and Festuca rubra were the main unsown grasses. The average contribution of L. perenne had increased since 1970–72; it was also present in two-thirds of the area from which it was absent in 1970–72, whilst the contribution of T. repens had decreased. The incidence of Cirsium arvense and Ranunculus spp. had also decreased, whilst that of Pteridium aquilinum had increased.     

Effect of fertilizer nitrogen rate and timing on herbage production and nitrogen use efficiency for first-cut silage

Eight field-plot experiments were carried out on established grassland swards between 1984 and 1988 to examine the effects of date and rate of application of calcium ammonium nitrate (CAN) on herbage dry matter (DM) yield and apparent efficiency of nitrogen (N) use at first-cut silage.
CAN application significantly increased ( P <0 ·001) the mean yields of herbage and N uptakes by herbage in all experiments. Herbage yields were similar ( P > 0·05) with N rates of 100 kg ha⁻¹, 125 kg ha⁻¹ or 150 kg ha⁻¹ in five experiments but in the other three there were increases above 100 kg ha⁻¹. Date of N application had a significant effect on DM yield in three experiments; this effect was inconsistent for both single and split dressings. Lower production was associated with reduced uptake of N, a trend that primarily reflected lower DM yields and not wide herbage N content variation.
It is concluded that selection of the date on which to apply fertilizer N in early spring to obtain optimum herbage yields at first-cut silage often required little precision. The use of fertilizer N rates >100 kg ha⁻¹ should be questioned where there are likely to be appreciable quantities of available N derived from non-fertilizer sources.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

The effects of paraquat and glyphosate residues in sprayed herbage on the development of seedlings of a normal and a paraquat-tolerant variety of Lolium perenne

Paraquat and glyphosate were sprayed on to simulated swards in a glasshouse experiment. Residues of these herbicides depressed emergence, growth and survival of Lolium perenne seedlings sown after spraying.
Residues of paraquat applied at a rate (1·5 kg ha^-1) within the range normally used for sward destruction killed nearly all seedlings of a normal L. perenne variety, Barlenna, but had much less effect on the paraquat-tolerant variety, Causeway. It was estimated that Causeway was almost nine times as tolerant of paraquat residues as Barlenna. Sowing at a depth of 20 mm gave some protection against paraquat residues. A delay of 10 d between spraying and sowing had equivocal effects. It was concluded that residues of paraquat used for killing swards before direct drilling may hinder the establishment of normal L. perenne varieties but not of paraquat-tolerant varieties.
Residues of glyphosate had equal effects on the two varieties and the effects were severest on seedlings sown on the soil surface immediately after spraying. It is thought to be unlikely that glyphosate residues in sprayed herbage would be a problem in direct drilling.     

The effect of drainage on herbage growth and soil development

The effects of drainage on the yield of perennial ryegrass ( Lolium perenne ) swards grown for silage, and on soil development under grassland, were studied between 1982 and 1985.
Over the 4 years of the experiment, drainage resulted in a mean increase in herbage production of 1-5 t dry matter (DM) ha⁻¹, 16% above the yield recorded on the undrained site. Drainage also led to increased herbage nitrogen contents at every cut except one, so that total nitrogen recovery was increased by an average of 24-8 kg N ha⁻¹ (15%) when drainage was installed. These total differences were mainly due to differences that were present at the first cut taken in June each year; drainage led to an average 24% increase in yield, and an average 27% increase in nitrogen recovery at the first cut.
Detailed sward measurements taken in 1985 endeavoured to explain these differences; drainage increased depth to free water over winter, enabling increased root efficiency and nitrogen availability, greater tillering and more rapid leaf extension in early spring. There was some suggestion that drainage also led to an improved topsoil structure in terms of improved aggregation and aeration. This is important for long-term soil development as well as short-term herbage yield.