Research note: Grazing‐index method procedures of vegetation surveys

Abstract

In the past, veld condition in the Karoo was assessed using the ecological index method. This recently changed to the grazing‐index method on account of the differently estimated grazing‐index values being used. The principles governing the method of survey remain the same. The method employs canopy‐spread‐cover strikes to record the species composition of different sites. By appropriate management of these line‐point data, different sites can be compared with one another by employing the Z‐index of agreement (similarity).     

Effect of stocking density on vegetation and animal performance within the semi-arid shrublands of the Karoo,South Africa

As the primary land use in the Karoo, extensive small-stock farming contributes significantly towards the agricultural economy of the region. However, the sustainability of this practice has been questioned for many years. This led to the establishment of a long-term stocking-density trial in 1988 at the Carnarvon Research Station. The trial was designed on a three-camp rotational grazing system under four stocking densities (8, 7, 5.5 and 4 ha SSU^–1). Following continuous treatment over 28 years, the results from a once-off assessment in February 2016 portrayed no significant differences between treatments with regard to plant height, cover, species diversity, ecological and grazing index scores. Animal production varied among treatments where the mean production per hectare was found to be the highest under the high stocking-density treatment. Due to the absence of historical data, the results from this study are not conclusive with regards to changes in the vegetation dynamics over time and therefore does not lend itself to making conclusions concerning stocking densities. It can, however, be concluded that the vegetation of the Western Upper Karoo is remarkably resilient to the perceived higher stocking densities set at the time, on condition that an appropriate grazing management strategy is applied.     

Examining the evidence for ecologically sustainable ostrich breeding practices on natural veld in the Little Karoo,South Africa

Land degradation in the Little Karoo is extensive. Overstocking of breeding ostriches on natural veld has been among the main causes of this. The National Department of Agriculture has set a general stocking rate of 60 ha LSU^?1 as a guideline for livestock on natural veld in the Little Karoo, which equates to 22.8 ha ostrich^?1. The aim of this review is to examine the scientific principles, data and assumptions the current recommended stocking rate for breeding ostriches on natural veld in the Little Karoo is based on and to investigate if there is a stocking rate (or range of stocking rates) that has been demonstrated to be ecologically and/or economically sustainable. We found no evidence that the recommended stocking rate of 22.8 ha ostrich^?1 is economically or ecologically sustainable in the Little Karoo. Most studies only addressed a single dimension of sustainability, and stocking rates deemed to be economically sustainable were several times higher than ecologically based recommendations. Given that economic and ecological objectives appear irreconcilable on open veld, an industry switch from extensive to intensive breeding practices may be a solution. Further research is required that integrates the socio-economic and ecological aspects of ostrich farming in the Little Karoo.     

Development of veld management research in the karoo region

The development of veld research in the semi‐arid Karoo areas since 1934 is outlined. It entailed key experiments on different types of veld, and the synthesis of grazing systems and their testing in experimental camps and co‐operatively on a practical farming scale. The experiments were primarily conducted with Merino sheep but Angora goats, Boer goats, cattle and Persian sheep were also employed. A large number of rotational grazing systems, special treatments and methods have been set out for the entire Karoo Region. The group camp approach serves as a basis for complete veld management planning in the Karoo Region.

Supporting research covered plant physiological, phenological and ecological studies; the determination of selective grazing habits of livestock; production of karoo bushes; and the development of research techniques especially in connection with quantitative and qualitative vegetation surveys.     

The relation between canopy spread cover and the aboveground available phytomass of Nama Karoo subshrubs and grasses

The canopy spread cover of Karoo bushes and grasses in the Nama Karoo was measured and related to the aboveground phytomass available to the grazing animal. A correlation coefficient of r = 0.7456, calculated for Karoo bushes, indicates that this method can be used to non-destructively estimate available phytomass. This in turn can be used for the prediction of current grazing capacity in areas where Karoo bushes are dominant in the vegetation. In the case of the Karoo grasses, a rather loose definition and interpretation of the canopy spread cover, led to a low correlation coefficient, r = 0.5662. This renders the present method largely unsuited for available phytomass estimations to be used in further predictions of grazing capacity in the mountainous areas where the studied grasses are dominant in the vegetation. In the more arid areas, however, the method can be used to advantage.     

Growing Degree Day: Noninvasive Remotely Sensed Method to Monitor Diet Crude Protein in Free-Ranging Cattle

The daily nutritional balance of free-ranging cattle is the net result of intake from available forage biomass and nutritive value weighed against the nutritional requirements of the animal. Plant phenology influences nutritive value. Plant phenology is dictated by time of year and an accumulation of photosynthetically active days. Growing degree day (GDD) is a concept that quantifies this relationship and has been used to predict nutritive value in perennial range grasses. GDD could be substituted for chemical analysis to inform grazing animal nutritional monitoring efforts. We hypothesized that in C4 grass-dominated rangelands, a cumulative GDD calculation would correlate with diet crude protein (CP) predictions obtained by fecal near infrared spectroscopy (FNIRS) from free-ranging cattle. Therefore, the objectives of our research were to evaluate the effectiveness of GDD to predict FNIRS-derived determinations of grazing cattle diet CP in 1) two groups of three individual animals grazing a small native pasture and 2) large commercial-scale herds grazing expansive rangelands. For the first objective, cumulative GDD and FNIRS-predicted diet CP were strongly correlated (r² = 0.76; P < 0.01). Relationships between cumulative GDD and FNIRS-predicted diet CP for the second objective varied considerably among ranches, ranging from a low r² of 0.05 (P = 0.871) to a high r² of 0.78 (P < 0.049). Similar values for individual ranch/year combinations were stronger; ranging from a minimum r² of 0.44 (P = 0.556) to a maximum of 0.95 (P = 0.051). The aggregate relationship between GDD and FNIRS-predicted CP for all ranch/year combinations was highly significant (r² = 0.37; P < 0.001), but the standard error was 1.86% CP. The noninvasive remotely sensed grazing animal nutritional monitoring method described here was accurate enough to inform tactical rangeland diet quality assessments but was not accurate enough to inform operational-scale grazing management decisions.     

基于生态安全的高寒牧区生态承载力评价

生态承载力指在一定社会经济条件下,自然生态系统维持其服务功能和自身健康的潜在能力。本研究以甘南藏族自治州（以下简称甘南州）ETM（Enhanced Thematic Mapper Plus）和TM（Thematic Mapper）遥感影像及统计年鉴数据为依据,应用生态安全评价模型PSR（压力一状态一响应）选取平均植被覆...     

Long-Term Production and Profitability From Grazing Cattle in the Northern Mixed Grass Prairie

Conventional wisdom among rangeland professionals has been that for long-term sustainability of grazing livestock operations, rangeland should be kept in high good to low excellent range condition. Our objective was to analyze production parameters, costs, returns, and profit using data generated over a 34-yr period (1969–2002) from grazing a Clayey range site in the mixed-grass prairie of western South Dakota with variable stocking rates to maintain pastures in low–fair, good, and excellent range condition classes. Cattle weights were measured at turnout and at the end of the grazing season. Gross income · ha^?1 was the product of gain · ha^?1 and price. Prices were based on historical National Agricultural Statistics Services feeder cattle prices. Annual variable costs were estimated using a yearling cattle budget developed by South Dakota State University agricultural economists. All economic values were adjusted to a constant dollar using the Bureau of Labor Statistics' Consumer Price Index. Stocking rate, average daily gain, total gain, net profit, gross revenue, and annual costs · ha^?1 varied among range condition classes. Net income for low–fair range condition ($27.61 · ha^?1) and good range condition ($29.43 · ha^?1) were not different, but both were greater than excellent range condition ($23.01 · ha^?1). Over the life of the study, real profit (adjusted for inflation) steadily increased for the low–fair and good treatments, whereas it remained level for the excellent treatment. Neither drought nor wet springs impacted profit differently for the three treatments. These results support generally observed rancher behavior regarding range condition: to maintain their rangeland in lower range condition than would be recommended by rangeland professionals. Ecosystem goods and services of increasing interest to society and associated with high range condition, such as floristic diversity, hydrologic function, and some species of wildlife, come at an opportunity cost to the rancher.     

The influence of tree density on the grass layer of three semi-arid savanna types of southern Africa

The influence of bush density on the grass component of the herbaceous layer was investigated for three savanna veld types (Molopo Thornveld, Mixed Vaalbos Thornveld, and the Eastern Grass and Bushveld) in the Northern Cape and North-West Province of South Africa. Relationships between bush density and the density, species composition and phytomass production of the grass component were determined over a four-year period. Bush encroachment had an adverse influence on the density and production of the grass component, but not on the species composition. Fluctuations in seasonal rainfall appeared to be the most important factor governing changes in the species composition of the grass component. Bush enchroachment (2 500 tree equivalents ha^?l) decreased the potential grazing capacity by as much as 331%, 149% and 58% in the Molopo Thornveld, the Mixed Vaalbos Thornveld and the Eastern Grass Bushveld, respectively, In comparison to sites with tree densities of less than 400 tree equivalents ha^?1. In the encroached condition grazing capacities of 15, 13 and 10 ha LSU^?1 are recommended for the Molopo Thornveld, the Mixed Vaalbos Thornveld and the Eastern Grass and Bushveld, respectively. The implementation of bush control measures in all problem areas is proposed in order to restore the productivity of these areas in the long term.     

Long-Term Vegetation Trends on Grazed and Ungrazed Chihuahuan Desert Rangelands

Long-term information on the effects of managed grazing versus excluded grazing effects on vegetation composition of desert rangelands is limited. Our study objectives were to evaluate changes in frequency of vegetation components and ecological condition scores under managed livestock grazing and excluded livestock grazing over a 38-yr period at various locations in the Chihuahuan Desert of southwestern New Mexico. Sampling occurred in 1962, 1981, 1992, 1998, 1999, and 2000. Range sites of loamy (1), gravelly (2), sandy (2), and shallow sandy (2) soils were used as replications. Black grama (Bouteloua eriopoda Torr.) was the primary vegetation component at the seven locations. Dyksterhuis quantitative climax procedures were used to determine trends in plant frequency based on a 1.91-cm loop and rangeland ecological condition scores. Frequency measures of total perennial grass, black grama, tobosa (Hilaria mutica Buckley), total shrubs, honey mesquite (Prosopis glandulosa Torr.), and other vegetation components were similar on both grazed and ungrazed treatments (P > 0.1) at the beginning and end of the study. The amount of change in rangeland ecological condition scores was the same positive increase (14%) for both grazed and ungrazed treatments. Major changes (P < 0.1) occurred within this 38-yr study period in ecological condition scores and frequency of total perennial grasses and black grama in response to annual fluctuations in precipitation. Based on this research, managed livestock grazing and excluded livestock grazing had the same long-term effects on change in plant frequency and rangeland ecological condition; thus, it appears that managed livestock grazing is sustainable on Chihuahuan desert rangelands receiving over 25 cm annual precipitation.     

Livestock grazing has minimal effect on the species richness and diversity of semi-arid shrubland in the Western Little Karoo,South Africa

The Succulent Karoo, one of two arid biodiversity hotspots in the world, is known for its high plant species richness, but little is known about the influence of topography and how it mediates the potentially deleterious effects of grazing. Changes in vegetation species composition, cover and species diversity were examined along piosphere gradients on northerly slopes, bottomlands and low-lying plains on 45 farms. Landscapes differed in the plant speciesand life-form composition, species richness and cover, with the more species-rich, heterogeneous grassy northerly slopes distinct from the plains and bottomlands, which were dominated by Pteronia pallens, Psilocaulon junceum and Drosanthenum spp. with more annual and ephemeral species. Overall, species richness declined linearly with decreasing rangeland condition. A weak grazing effect was detected only on the north-facing slopes, where shrubs and grass decreased with intense grazing. Years of overgrazing have resulted in the widespread dominance of P. pallens, especially on the plains and bottomlands, rendering them insensitive to grazing. The current condition of the northern slopes should be maintained and managed such that palatable species can spread to more degraded areas of the landscape.     

Grazing Density Effects on Cover,Species Composition,and Nitrogen Fixation of Biological Soil Crust in an Inner Mongolia Steppe

Biological soil crusts (BSCs) are important in many arid and semiarid ecosystems for their abilities to fix atmospheric nitrogen (N) and stabilize surface soil. Grazing disturbance has a profound influence on abundance, species composition, and ecological functioning of BSCs. To elucidate the effects of grazing on BSCs in Inner Mongolia grasslands, an investigation was conducted in a typical steppe that had previously been subjected to long-term grazing with six grazing densities (control: 0 sheep · ha^?1, very light: 4 sheep ·ha^?1, light: 8 sheep · ha^?1, medium: 12 sheep · ha^?1, heavy: 16 sheep · ha^?1, and very heavy: 20 sheep · ha^?1). Cover, species composition, potential N-fixing activity, and potential N input of BSC indicate that long-term grazing significantly reduced the importance of BSC in N input and soil stabilization. Such reductions were highly related to grazing density. Very light grazing had no significant effect on the role of BSC in soil stabilization, but resulted in a 13.3% reduction in BSC N input potential. Heavy and very heavy grazing led to a decrease of potential N input by one order of magnitude, and a decline of BSC function via a shift from high coverage of an attached group–dominated BSC community to a low coverage of a vagrant group–dominated community. Constraining grazing level to a very light density—and to a lesser extent, a light grazing density—is likely a preferred practice for conserving BSC and the ecological services it provides in N fixation and soil stabilization.     

Habitat Effects on Condition of Doe Mule Deer in Arid Mixed Woodland-grassland

Productivity of mule deer (Odocoileus hemionus Raf.) populations is closely linked to individual nutritional condition. We modeled body fat of individual does as a function of vegetation cover, composition, and water characteristics of their annual, summer, and winter home ranges in north-central New Mexico. We also modeled home range size as a function of the same characteristics. Levels of body fat were most closely and negatively related to the amount of pinyon-juniper in an individual deer's annual home range (F_1,21 = 7.6; P = 0.012; r² = 0.26). Pinyon-juniper types provided little (combined ground cover of preferred forbs and shrubs = 5.7%) mule deer forage but were included in home ranges in excess of their availability on the landscape, likely because of security cover attributes. Proportion of grasslands in home ranges was most strongly related to both annual (F_1,23 = 4.9; P = 0.037; r² = 0.18) and summer (F_2,25 = 5.7; P = 0.009; r² = 0.31) home range sizes, and home ranges increased as the grassland component increased, indicating that this habitat type was providing little value to mule deer. Grassland (0.2% combined cover of preferred forb and shrub) and montane conifer (3.2% ground cover of preferred forb and shrub) habitat types similarly lacked preferred mule deer food, and grasslands also lacked cover. Most immediate gains in mule deer habitat in north-central New Mexico may be attained by management of pinyon-juniper communities to increase forage quantity and quality while maintaining cover attributes. Gains can also be realized in grasslands, but here management must establish both cover and forage.     

Measuring the impact of indigenous herbivores: reply to Kerley andLandman

Pteronia paniculata is an indigenous, unpalatable shrub that invades mismanaged Karooveld, resulting in degraded rangelands with low species diversity and grazing potential. We conducted a series of trials in the Succulent Karoo Randteveld near Barrydale to determine if the uniform defoliation of P. paniculata dominated vegetation at two heights (0.05 m and 0.20 m above ground level) using a brush-cutter and, in one trial, application of a second cut will improve the plant species composition, productivity and grazing capacity of the veld. Brush-cutting treatments and the uncut control all resulted in a change in species composition towards greater species diversity and more palatable species and an average increase of 540 kg ha^?1 (28%) in above-ground biomass over four years. It appears that there was a pervasive improvement in species composition associated with a general decline in the cover and abundance of P. paniculata over the time-scale of the present study that was not influenced by the defoliation treatments, except for the 1996-cut treatment where the cover of P. paniculata increased. The absence of propagules of palatable species in the soil seed bank and competition from P. paniculata (a long-lived, perennial shrub) are assumed to be among the main reasons for the lack of response of the vegetation community to the defoliation treatments. Brush-cutting (in the absence of reseeding), aimed at reducing the dominance of unpalatable karoo shrubs, was more costly but not significantly better than long-term resting in improving veld composition or forage production.     

Evaluation of shrub species for sheep grazing in the Strandveld of the Cape West Coast of South Africa

The production of three exotic shrub species, Atriplex nummularia (old man saltbush), A. canescens (four‐winged salt‐bush) and Medicago arborea (tree lucerne) and one indigenous evergreen shrub, Chrysanthemoides monilifera subsp. pisifera (west coast bitou), was evaluated in the Strandveld of the Cape West Coast in terms of grazing capacity (ssu ha^?1) and average daily gain (ADG) (g ssu^?1 day^?1) using mature Dorper sheep. Medicago arborea died out during the first season. The other three shrubs, however, survived and over a seven‐year period mean grazing capacities of 1.7, 1.6 and 1.0 ssu ha^?1 were obtained, respectively, for C. monilifera, A. nummularia and A. canescens, which is in excess of the area norm of 5 ha ssu^?1 for natural veld. Positive ADGs of 80 and 30 g ssu^?1 day^?1 were obtained on A. nummularia and A. canescens, respectively, but not on C. monilifera (‐180 g ssu^?1 day^?1).     

Impacts of non-selective grazing on cover,composition, and productivity of Nama-karoo grassy shrubland

The non-selective rotational grazing system has undergone a long and controversial development. The merits of this grazing system, where relatively large numbers of livestock are herded into numerous small paddocks for short timespans with long rests between grazings, have not been formally evaluated in a long-term monitoring experiment in the Karoo. In this study we used exclosures (controls) on a 7 000 ha farm in the Central Lower Karoo, camped into approximately 50-ha paddocks, to evaluate the impact of this grazing system on certain vegetation parameters. We report on the first four years of monitoring, after each of four replicate paddocks had received four treatments (one treatment = 40 - 60 Large Stock Unit grazing days per hectare over a period of 2 - 16 days). Concentrated defoliation with concomitant trampling, dunging, and urinating did not influence the perennial species composition, and cover of this grassy, semi-arid shrubland. Changes over time in plant composition, and cover are explained by annual, and short-term (e.g. quarterly) rainfall rather than by grazing impacts. Ephemerals were not favoured by this grazing system, but litter was more abundant in the treatment than the control areas. The dominant grass, Eragrostis lebmanniana, and shrub, Pentzia incana, are resilient to this defoliation regime, and show signs of compensatory growth. As yet there is no evidence that non-selective grazing increases diversity, but the severe defoliation and trampling may enhance the biomass turnover rate, resulting in more vigorous, and productive plants in the grazed areas. Diversity is unlkely to change rapidly in response to grazing, largely because of the persistence of grazing-tolerant perennials. Concentrating mixed herds of livestock onto small areas with lengthy rests can be a useful tool for 'kick-starting' moribund karoo veld into greater productivity. Several of the impacts hold potential for restoring the rangeland quality of degraded areas, but this needs to be tested.     

Effects of Grazing Intensity,Precipitation, and Temperature on Forage Production

Questions have been raised about whether herbaceous productivity declines linearly with grazing or whether low levels of grazing can increase productivity. This paper reports the response of forage production to cattle grazing on prairie dominated by Kentucky bluegrass (Poa pratensis L.) in south-central North Dakota through the growing season at 5 grazing intensities: no grazing, light grazing (1.3 ±  animal unit months [AUM] · ha^-1), moderate grazing (2.7 ±  AUM · ha^-1), heavy grazing (4.4 ±  AUM · ha^-1), and extreme grazing (6.9 ±  AUM · ha^-1; mean ± SD). Annual herbage production data were collected on silty and overflow range sites from 1989 to 2005. Precipitation and sod temperature were used as covariates in the analysis. On silty range sites, the light treatment produced the most herbage (3 410 kg · ha^-1), and production was reduced as the grazing intensity increased. Average total production for the season was 545 kg · ha^-1 less on the ungrazed treatment and 909 kg · ha^-1 less on the extreme treatment than on the light treatment. On overflow range sites, there were no significant differences between the light (4 131 kg · ha^-1), moderate (4 360 kg · ha^-1), and heavy treatments (4 362 kg · ha^-1; P &spigt; 0.05). Total production on overflow range sites interacted with precipitation, and production on the grazed treatments was greater than on the ungrazed treatment when precipitation (from the end of the growing season in the previous year to the end of the grazing season in the current year) was greater than 267.0, 248.4, 262.4, or 531.5 mm on the light, moderate, heavy, and extreme treatments, respectively. However, production on the extreme treatment was less than on the ungrazed treatment if precipitation was less than 315.2 mm. We conclude that low to moderate levels of grazing can increase production over no grazing, but that the level of grazing that maximizes production depends upon the growing conditions of the current year.     

Winter Grazing and Rainfall Synergistically Affect Soil Seed Bank in Semiarid Area

The soil seed bank is an important ecological component of grassland restoration and renewal. In semiarid regions, grassland restoration and renewal are highly affected by annual variations in precipitation and grazing activity because these variations can affect the composition, density, richness, and diversity of seeds in the soil. This study aimed to characterize and compare these parameters of the germinable seed bank under different stocking rates in a winter grazing system in a semiarid area of China in 2015 and 2016 (dry and near-average rainfall condition, respectively). The composition, density, richness, and diversity of seeds were determined by the method of seedling emergence. The results showed that a total of 18 species belonging to nine families germinated from the soil. Drought significantly reduced the density, richness, and diversity of the soil seed bank, but grazing was able to significantly increase the richness and diversity of the soil seed bank by increasing the richness and diversity of the aboveground vegetation. The similarity between the soil seed bank and aboveground vegetation was influenced by the rainfall conditions: in the dry year, it was higher at the lower stocking rates (0 and 0.4 animal unit months [AUM] ha^–1), and in the near-average rainfall condition year, it was higher at the higher stocking rates (0.8 and 1.3 AUM ha^–1).     

A study of animal production under different grazing regimes in the False Upper Karoo at Grootfontein

Three systems of animal production on the fragile Karoo veld, i.e. multi-paddock rotational grazing, pauci-paddock rotational grazing and continuous grazing were studied. Of these systems, multi-paddock rotational grazing gave the best results with continuous stocking resulting in the worst. All three systems studied had long histories of both continuous stocking- and of pauci-paddock rotational grazing. The veld, under both these previous systems of management has stabilised, but at different seral stages. The continuously stocked range stabilised in the early developmental seral stage, consisting mainly of annual grasses. The veld under pauci-paddock rotational management stabilised in the early to mid-developmental seral stage, consisting mainly of perennial grasses. Experimentation on veld in a stable condition is a prerequisite if valid conclusions in respect of rotational grazing and continuous stocking are to be drawn. Rotational grazing on overgrazed veld will result in poor animal performance, while it is generally accepted that continuous stocking on pristine veld will result in excellent animal performance for a number of years. Multi-paddock rotational grazing outperformed both pauci-paddock rotational grazing as well as continuous stocking on both types of veld. This is surprising taking into account the mid-seral stage condition of the vegetation. This supports the argument that with improved veld management, an improved production potential is possible on veld even in a degraded or current condition.     

Grazing-Induced Modifications to Peak Standing Crop in Northern Mixed-grass Prairie

Selective grazing can modify the productive capacity of rangelands by reducing competitiveness of productive, palatable species and increasing the composition of more grazing-resistant species. A grazing system (season-long and short-duration rotational grazing) × stocking rate (light: 16 steers · 80 ha^-1, moderate: 4 steers · 12 ha^-1, and heavy: 4 steers · 9 ha^-1) study was initiated in 1982 on northern mixed-grass prairie. Here, we report on the final 16 years of this study (1991–2006). Spring (April + May + June) precipitation explained at least 54% of the variation in peak standing crop. The percentage of variation explained by spring precipitation was similar between stocking rates with short-duration grazing but decreased with increasing stocking rate for season-long grazing. April precipitation explained the greatest percentage of the variation in peak standing crop for the light stocking rate (45%), May precipitation for the moderate stocking rate (49%), and June precipitation for the heavy stocking rate (34%). Peak standing crop was 23%–29% greater with light (1 495 ± 66 kg · ha^-1, mean ± 1 SE) compared to moderate (1 218 ± 64 kg · ha^-1) and heavy (1 156 ± 56 kg · ha^-1) stocking rates, which did not differ. Differences in peak standing crop among stocking rates occurred during average and wet but not dry springs. Neither the interaction of grazing system and stocking rate nor grazing system alone affected standing crop across all years or dry, average, or wet springs. Grazing-induced modification of productive capacity in this northern mixed-grass prairie is attributed to changes in species composition with increasing stocking rate as the less productive, warm-season shortgrass blue grama (Bouteloua gracilis [H.B.K.] Lag. ex Griffiths) increases at the expense of more productive, cool-season midheight grasses. Land managers may need to substantially modify management to offset these losses in productive capacity.