枫香优树14年生子代遗传变异及选择

在福建省洋口国有林场,对来自安徽省南部黄山地区的53个枫香家系进行子代测定,研究家系的遗传变异规律,开展优良家系及单株选择.结果表明:子代试验林14年生时,树高、胸径和单株材积平均值分别为8.05m、6.30cm和0.014 93 m3,生长性状的家系遗传力为0.437～0.576,受中强度的遗传控制,单株遗传力为D.113～0.195,受较弱的遗传控制;木材材性的家系遗传力为0.507～ 0.591,单株遗传力为0.360～0.500,均受中强度的遗传控制.以生长量为主要指标,选出速生优良家系9个,其树高、胸径和单株材积平均值分别为8.85 m、7.22 cm和0.021 49 m3,遗传增益分别为2.86％、5.47％和13.4％;从木材基本密度大于0.500 g·m-3的家系中选出7个生长材性兼优家系,树高、胸径、单株材积和木材基本密度的平均值分别为8.34m、6.79 cm、0.018 58 m3和0.518 g·m-3.所选出的速生优良家系和生长材性兼优家系可作为枫香育种材料在生产上推广应用.     

火炬松半同胞家系遗传测定与早期性状评定

对15年生火炬松半同胞家系的生长和材性性状测定表明:火炬松家系树高、胸径、材积、木材基本密度、纤维素含量、管胞长度与宽度等具有显著的差异;在家系水平上,木材基本密度、纤维素含量、管胞长度与宽度和材积生长相关不明显．遗传力测定表明:家系生长性状与材性性状平均遗传力均较高,达到中等程度以上．对树高生长、胸径生长、木材基本密度、纤维素含量、管胞长度与宽度等性状进行年度相关、早晚期选择研究表明:火炬松家系的生长相关主要由遗传决定;树高从第5年开始,胸径从第4年开始与以后各年的相关系效均在0．7以上,表明以此为最低年龄进行早期选择有较大的可靠性;木材基本密度10年生与其后各年龄级的相关系数值趋于稳定,术材密度最低选择年龄为10年;木材纤维素含量11年生与其后各年龄级的相关系效值趋于稳定,木材纤维素含量的最低选择年龄为11年．     

Variation analysis and fine family selection on half-sib progenies from clonal seed orchard of Pinus massoniana

Variation of traits that include height, diameter at breast height (DBH), stem volume, crown width, straightness, wood basic density, tracheid length and width of open-pollinated progenies from clonal seed orchard of Pinus massoniana was analyzed. Differences in traits such as growth, tree form and wood quality among families was significant, which indicated that the improvement potential was notable. Heritability of height and wood basic density reached 0.691 and 0.640, respectively. According to a correlation analysis between traits, stem straightness could be improved indirectly when growth traits are improved; growth traits were significantly correlated with wood basic density; and wood basic density was not significantly correlated with length and width of the tracheid. Under the selection ratio of 10%, genetic gains were 19.74% and 19.32% respectively, and selected families from the progeny test stands at the age of five years and eight years were the same. Genetic correlation of height, DBH and stem volume was also significant at the level of 0.01. These show that selection of the short rotation of P. massoniana for pulp and paper use would be efficient for five-year-old trees. According to the comparison of the heritability of all the traits and taking into account the result of canonical correlation analysis, height could be considered as the most important index for selection. Given wood quality of all the families, and flowering and fruit settings of the clones in a seed orchard, twelve fine families were selected at 10% selection ratio of height growth and stem volume. Translated from Scientia Silvae Sinicae, 2005, 41(6): 43–49 [译自: 林业科学]     

Growth and wood basic density of acacia hybrid clones at three locations in Vietnam

Field trials testing a total of 27 clones of the interspecific hybrid Acacia mangium × A. auriculiformis and seedling controls of the parental species were established at Ba Vi and Yen Thanh in the north of Vietnam and Long Thanh in the south. At both Ba Vi and Yen Thanh there were significant (P < 0.001) differences in height and diameter at breast height (DBH) among 22 tested clones at 4 years. At Long Thanh, twelve hybrid clones did not differ significantly in DBH at age 3 years, but did (P < 0.001) at age 5 years. At the two northern sites the acacia hybrid clones had significantly greater DBH than control seedlots of the parental species. At Long Thanh, DBH of the hybrid clones and A. mangium was similar, with a genetically improved seedlot of A. mangium displaying the best DBH. Mean wood basic density at breast height of the acacia hybrid clones was 539 kg m⁻³ at Yen Thanh at age 8 years, and 473 kg m⁻³ at Long Thanh at age 5 years; density for A. mangium at Long Thanh was only slightly lower than the hybrid clones at 461 kg m⁻³. Linear regressions of Pilodyn penetration (PP) at breast height on wood basic density explained 60% of the variance in density of treatments (clones and control seedlots) at Yen Thanh and 36% at Long Thanh. There were significant differences between hybrid clones in PP at all three trial sites. Clonal DBH performance was not strongly correlated across the three trial sites; Pearson correlations of clone mean DBH between pairs of sites ranged from −0.47 to 0.20. Clonal rankings for PP were more stable, with Pearson correlations between pairs of sites ranging from r = 0.71 to 0.78.     

25年生马尾松生长和木材基本密度家系变异与选择

以福建省华安县西陂林场的25年生来自省内不同产地的46个马尾松自由授粉家系试验林为试材,研究了达主伐年龄时马尾松生长、干形和木材基本密度的家系差异以及性状间的相关性,并依据不同培育目标选择优良家系。结果表明,马尾松生长、树干圆满度和木材基本密度皆存在显著的家系差异,其中生长和树干通直度变异相对较大,而树干圆满度和木材基本密度变异较小,变异系数在10%以下。生长和干形性状受到弱至中度的家系遗传控制,家系遗传力为0.23 0.46;木材密度受到的遗传控制相对较强,家系遗传力为0.46。性状遗传相关分析结果显示,选择速生种源可同时改良树干圆满度,但对树干通直度和木材基本密度影响不大。根据不同用材对材质材性要求,选出24个纸浆纤维材优良家系和6个锯材优良家系,平均木材干物质量和胸径生长皆为对照的1.16倍。     

长白落叶松纸浆材优良家系多性状联合选择研究

采用胸径木芯与伐倒木取样方法,对24年生长白落叶松优树子代测定林14个处理的生长性状、木芯材质性状、解析木材质性状及其制浆造纸特性进行了研究,结果表明:生长性状、木芯基本密度、木芯管胞性状均存在丰富的变异;家系间生长性状差异极显著;木芯基本密度、木芯管胞性状差异显著,家系遗传力在0.56 0.80之间;解析木基本密度、早材微纤丝角、管胞长度、早材壁腔比、木质素、阿拉伯糖含量、抗张强度、撕裂度等差异极显著,家系遗传力在0.64 0.86之间;晚材率、管胞长宽比、综纤维素含量等差异显著,家系遗传力在0.51 0.61,家系水平的材性改良潜力较大。生长性状与木芯管胞长度呈极显著正相关,与木芯基本密度、管胞长宽比相关不显著;木芯基本密度、管胞长度、管胞宽度以及管胞长宽比与解析木相应性状呈极显著正相关关系,且回归模型较为理想,可以利用胸径木芯各性状值预测其单株值,间接选择和评价纸浆材优良家系。通过生长性状、木芯基本密度和木芯管胞长宽比综合分析选出166、169为优良家系,根据解析木生长、材质以及纸张物理性状选出的优良家系与其结果一致。优良家系材积、木芯基本密度和木芯管胞长宽比的遗传增益(超过对照)分别为48.34%(38.80%)、14.01%(3.63%)和19.89%(6.42%)。     

杉木2代种子园子代主要经济性状遗传变异及单株选择

(Cunninghamialanceolata(Lamb.)Hook.)作为我国南方林区重要的用材树种,经过几十年的遗传改良已进入到高世代时期,遗传改良工作取得了较大进展,林木生长量也已获得较大增长.20世纪90年代初浙江省从杉木1代种子园家系子代测定林、杂交试验林和种源试验林中选择出优良材料营建了2代种子园[1],对新建的种子园材料进行子代测定是林木遗传改良的重要工作,不仅可了解其种子园的改良效果和主要经济性状遗传变异规律,也可为高世代种子园营建提供选择材料.     

Genetic and environmental variation in wood properties of <Emphasis Type="Italic">Acacia melanoxylon</Emphasis>

Context  

Australian blackwood (Acacia melanoxylon), is a fast-growing, high-quality, appearance-grade timber species native to eastern Australia. Some of its key wood properties are percentage heartwood, heartwood colour, basic density, wood stiffness and green moisture content. Variation in these properties affects log value, processing and marketing.     

马尾松无性系种子园半同胞子代变异分析和家系选择

以福建省漳平五一林场马尾松无性系种子园自由授粉子代半同胞家系为研究对象,分析其树高、胸径、材积、冠幅、通直度、高径比、冠径比、木材基本密度、管胞长度和宽度、管胞长宽比等性状,发现家系间除管胞长宽比性状差异显著,其余性状均达差异极显著,暗示着马尾松种子园半同胞家系间存在较为丰富的变异,具选育潜力。8年生林分内各性状的家系遗传力以树高性状为最高(0·691),其次是木材基本密度(0·640)。各性状间的相关性分析结果表明,生长量性状的改良可间接改善通直度,且生长量性状与木材基本密度存在显著正相关关系,木材基本密度与管胞长度和宽度不相关。采用10%入选率,发现5年生林分和8年生林分入选家系一致,材积遗传增益分别达19·74%和19·23%,且5年生林分和8年生林分生长量性状的遗传相关极显著,认为马尾松制浆造纸材短轮伐期的初选年龄可确定为5年生;根据性状遗传力的相对大小和典型相关分析的结果,认为马尾松家系选择可适当注重树高生长量指标。以树高和材积为选择指标,按照10%入选率,兼顾材性和种子园内无性系的开花结实情况,筛选出制浆造纸材短轮伐期优良家系12个。     

蓝桉、直干桉木材密度在家系内和株内的变异

在蓝桉、直干桉子测林中各抽3个家系，从不同的区组按径阶等距抽样测定木材的密度，研究家系内木材密度的变异；在种子园中抽取蓝桉30株、直干桉36株作试材研究木材密度株内变异。结果表明蓝桉家系内不同径阶的木材密度差异不显著，直干桉极显著；蓝桉的木材密度在区组和家系间的差异都显著，而直干桉都不显著。株内不同方向的木材密度差异两树种均不显著，而不同高度和不同年龄的木材密度差异都极显著；在亲缘关系混杂的群体中两树种的树高和胸径与木材密度均呈负相关，而在3个蓝桉家系内，胸径与密度呈正相关，直干桉3个家系中两个呈正相关，1个呈负相关，但都不显著。为将这两个树种纸浆材的主伐年龄定为6年提供了充分的科学依据,且在2年生时对蓝桉、直干桉作早期选择可取得满意的效果，对直干桉尤为明显。     

尾叶桉×细叶桉木材密度与生长的联合选择

[目的]评估Resistograph钻刺法间接测定尾叶桉×细叶桉木材密度的可靠性,检测杂交亲本对子代表型的效应以及生长与木材密度的相关,评选速生、优质的尾细桉杂种。[方法]基于10株尾叶桉与10株细叶桉不完全析因交配产生的56个杂交组合的7.5年生试验林,利用79株分析容积法与Resistograph钻刺法测定的木材密度的相关,通过方差分析检测亲本对杂种生长和木材密度的效应,结合多重比较和独立淘汰法进行材积和木材密度的联合选择。[结果]容积法与Resistograph钻刺法测定的木材密度的表型相关系数为0.52(P0.001),遗传相关系数为0.55(P0.05);树高、胸径及材积的母本间和父本间均呈极显著差异(P0.001或0.01),但母本×父本互作的效应不显著;对钻刺木材密度,父本间呈极显著差异(P0.001),母本×父本互作显著(P0.05),但母本间差异不显著;树高、胸径和材积间的表型相关和遗传相关均极显著(P0.001),其与钻刺木材密度的表型相关极显著(P0.001),但遗传相关不显著;评选出速生、木材密度较高的杂交组合14个、单株17株。[结论]Resistograph钻刺法是一种间接测定尾细桉木材密度的简便、经济和可靠的方法;母本和父本选择以及母本与父本的组配对培育速生、材质优良的尾细桉杂种均较重要;尾细桉生长与木材密度的遗传相关不显著,需要对这两类性状分别进行选择;评选的尾细桉杂交组合和单株为培育速生、优质的桉树良种提供了有用的材料。     

Genetic variation in wood specific gravity among half-sib families of chir pine (<Emphasis Type="Italic">Pinus roxburghii</Emphasis> sargent)

Chir pine (Pinus roxburghii) is an important tree species that grows all along the Himalayas. Wood specific gravity of chir pine before and after removal of extractives from wood increment cores was assessed from a 22-years-old progeny test in Himachal Pradesh, India. These values averaged 0.433 and 0.425, respectively. A large amount of genetic variation among the 58 half-sib families was found as indicated by range of values, additive coefficient of variation, variance estimates and narrow-sense heritability values for these traits. Moisture and extractive content averaged 86.259% of oven-dry weight and 2.003% of extractive-free oven-dry weight, respectively. Wood extractive content was highly variable and the family differences were highly heritable (h _f² = 0.5831). There was wide variability in moisture content, but a large portion of it was due to environmental or non-additive component of genetic variation. Heritability on family mean basis was found to be lower than that on individual tree basis for each trait. Estimated gain in specific gravity resulting from 30 to 50% family selection ranged from 0.0080 to 0.0127. Growth data and specific gravity were not significantly correlated implying that selection for higher growth rate would not necessarily result in reduction in wood specific gravity in chir pine.     

Wood properties of <Emphasis Type="Italic">Pericopsis mooniana</Emphasis> grown in a plantation in Indonesia

The relationships between growth characteristics and wood properties were investigated for a threatened species, Pericopsis mooniana, to promote the establishment of plantations of this species in the tropics. Growth characteristics (diameter and height) and stress-wave velocity (SWV) of trees were measured for 22-year-old P. mooniana trees planted in Indonesia. The trees were categorized into three groups, fast-growing, middle-growing, and slow-growing trees, to investigate the effect of growth rate on the wood properties. In addition, radial variation of anatomical characteristics and wood properties were determined. No significant correlation was found between growth characteristics and SWV. The values for the vessel diameter, cell wall thickness of wood fibers, wood fiber length, basic density, modulus of elasticity, and modulus of rupture from wood at the bark side were higher than those at the pith side. On the other hand, vessel frequency gradually decreased from pith to bark. These results suggested that low-quality wood, such as juvenile wood, existed near the pith area.     

Geographic variation in tree growth and wood density of <Emphasis Type="Italic">Guazuma crinita</Emphasis> Mart. in the Peruvian Amazon

Guazuma crinita Mart. is a fast-growing timber tree in the Peruvian Amazon. A trial of 11 Peruvian provenances was established in three planting zones. Tree growth at 6, 12, 18 and 30 months and wood density at 32 months were analyzed across and within zones. Zones accounted for more variation than provenances. Density varied significantly due to provenances in the analysis across zones, whereas growth exhibited strong provenance by zone interactions. The local provenance grew well and produced relatively dense wood in all zones. The relative magnitude of variation due to provenances depended on the planting zone. Density was greater in the lower than in the upper stem. Density increased with longitude from the more humid to the drier part of the sample region. In general, correlations indicated that larger trees had lower density and a larger difference in density between the lower and upper stem, but the strength of these relationships depended on the provenance and zone. Results suggest that fast-growing provenances can be selected at an early age without significantly reducing wood density. The local provenance is recommended for reforestation pending future research demonstrating the superiority of foreign provenances. Some practical implications for tree-improvement programs are discussed.     

马尾松自由授粉家系生长和材质的遗传分析及联合选择*

对９年生８１年自由授粉家系的研究发现，马尾松高，径生长，树干通直度，分枝性状和木材基本密度在家系间差异显著，具有适度的遗传力，而未发现管胞长度显著的家系效应，由于高、径生长和基本密度呈微弱的正相关至微弱的负相关，进行生长迅速家系的选择并不会至致使幼龄材材性的变劣，利用通直度和分枝习性等指标可以对高，径生长和木材密度进行间接选择，分枝角可能是一个独特的指标，它独立于其它多数性状。借助于约束或无约束选     

Impact of Ceratocystis wilt on eucalyptus tree growth and cellulose pulp yield

In this study, we determined the effects of Ceratocystis fimbriata wilt on volumetric growth and cellulose pulp yield of eucalyptus. Seven‐year‐old healthy and infected trees were separated into different classes of severity, and the individual volume of each whole tree with bark was determined. Volumetric losses were found regardless of the class of disease severity, resulting in reduction in volumetric growth of the trees from 65% to 87%, depending on the infection level. Two approaches were used for evaluating the effects of the disease on cellulose pulp yield: (i) evaluation of wood basic density and chemical composition, contents of ethanol/toluene extractives and Klason lignin, and the alkaline load and screen yield across Kraft pulping to kappa number 18 ± 0.5 for both healthy and discoloured materials; (ii) determination of the alkaline load and yield across Kraft pulping to kappa number 18 ± 0.5 of woods with different levels of discoloration (0, 25, 50, 75 and 100% of discoloured wood). Discoloured wood presented lower basic density, higher ethanol/toluene extractives and Klason lignin, higher alkaline load and lower pulp screened yield at kappa number 18 ± 0.5 compared with the healthy ones. Increasing the fraction of infected wood in chip furnishes resulted in proportionally increased alkaline load demand and decreased screen yields. Adding 50% of infected wood in the chip furnish resulted in 23.8% increase in the alkali load and 13.7% reduction in the pulp screen yield at kappa number 18 ± 0.5. The results obtained have proved that the disease reduces volumetric growth, increases wood consumption and reduces yield in eucalyptus pulp production.     

Cellulose content as a selection trait in breeding for kraft pulp yield in Eucalyptus urophylla

? The aim of this study was to investigate the effectiveness of using cellulose content, measured by the diglyme-HCl method, as a selection trait in breeding programs for kraft pulp yield in Eucalyptus urophylla.

? A total of 275 trees from sixty-two families were sampled from a thinned progeny trial of E. urophylla in northern Vietnam to evaluate cellulose content from breast-height increment cores. Among those, twenty unrelated trees were felled to evaluate cellulose content and pulp yield from breast-height disk samples.

? The regression of pulp yield of disk samples on cellulose content was strong either from disks (R ² = 0.83) or increment cores (R ² = 0.69). There was no significant difference in cellulose content between the provenances. The narrow-sense within-provenance heritability of cellulose content was 0.50 and the coefficient of additive genetic variation was 3.9%. Genetic correlations between cellulose content and growth (0.28–0.45) or wood basic density (?0.02) were not significantly different from zero.

? Breast-height increment core cellulose content measured by diglyme-HCl method is under strong genetic control and can be used to rank trees for pulp yield in E. urophylla plantations. Selection for increased cellulose content would have only minor effects on growth and wood basic density.

     

How dense is my blue gum? prediction of whole-tree basic density of <Emphasis Type="Italic">Eucalyptus globulus</Emphasis>

Accurate prediction of whole-tree wood basic density (BD) from indirect density measures is important for estimating Eucalytpus globulus breeding values and genetic gain. We measured 100 E. globulus trees at 6.5 years of age on each of a high productivity site (Albany) and a low productivity site (Esperance), sampling 16 full-sib families and seven open-pollinated ‘check’ seedlots. On each tree we assessed pilodyn penetration at 1.3 m and BD of: (a) the outer 10 mm which was penetrated by the pilodyn pin of a bark-bark core, (b) a whole bark-bark core at 1.3 m, (c) a cross-sectional disk at 1.3 m and (d) the whole tree, represented by cross-sectional disks at five heights. Site effects and genetic values for families and checks were determined by mixed model analysis. A trend of increasing BD from pith to bark was more pronounced at Albany, where wood formed before age 3.5 years was 12 kg/m³ less dense than at Esperance (P = 0.01) and wood formed from 3.5 to 6.5 years was 27 kg/m³ denser than at Esperance (P < 0.001). Mean whole-tree BD was 19 kg/m³ greater at Albany (P < 0.001). Pilodyn penetration was well-correlated with BD of the 10 mm sample of penetrated wood and this relationship was not greatly affected by site variation. Genetic values for pilodyn penetration formed precise relationships with BD of breast-height discs and whole-tree BD although they were site-specific. Pilodyn penetration was a better predictor for genetic values of whole-tree BD than were bark-bark cores.     

Photosynthesis and canopy characteristics in genetically defined families of silver birch (Betula pendula)

Net photosynthetic rates (A) of leaves in upper and lower crown layers (A(upper) and A(lower)), leaf area index (LAI), mean tilt angle (MTA), several leaf characteristics, and volume growth were observed in fast- and slow-growing families of a 14-year-old full-sib and half-sib family progeny test of Betula pendula Roth. Each measure of net photosynthetic rate was calculated after correcting measured net photosynthesis for the effects of environmental variables. The differences in A(upper) and LAI among families were significant. The proportions of the total variance assigned to family for A(upper), A(lower) and LAI were 33.64, 28.93 and 54.99%, respectively. The mean A(upper) and LAI of the fast-growing families were significantly higher than those of the slow-growing families, whereas the mean A(lower) of the fast-growing families was significantly lower than that of the slow-growing families. There were also significant differences among families in leaf size, leaf shape, and the ratios leaf fresh weight/area and leaf dry weight/area. Between 27.55 and 54.55% of the total variance in these characteristics could be assigned to the family effect. Volume growth was positively correlated with A(upper) and LAI, but it was most strongly correlated with A(upper) x LAI.     

日本落叶松自由授粉家系纸浆材材性遗传变异的研究

对12年生日本落叶松自由授粉子代测定林纸浆材材质性状进行了遗传变异分析、遗传参数估计、相关分析及综合评价和选择。结果表明:家系间综纤维素含量、基本密度、早材和晚材纤维长度、早材和晚材壁腔比差异极显著或显著,10g·L-1NaOH抽出物差异也达10%水平显著。各性状受强度或中等强度的遗传控制,对日本落叶松进行家系水平的材性改良具有很大的潜力。木材密度与早、晚材壁腔比、10g·L-1NaOH抽出物呈显著正相关,与综纤维素含量呈显著负相关,与早、晚材纤维长度相互独立;晚材纤维长度与早材纤维长度存在紧密正相关,纤维长度与早材壁腔比存在显著负相关关系,与晚材壁腔比、综纤维素含量和10g·L-1NaOH抽出物相互独立;综纤维素含量与早材壁腔比呈显著负相关,与晚材壁腔比相关不显著;10g·L-1NaOH抽出物与综纤维素含量、早、晚材壁腔比均相互独立。根据对日本落叶松材质性状的综合评价结果,选出最适合作纸浆林培育的249、340、346、217和364号家系。