Dry matter production and nutrient cycling in agroforestry systems of cardamom grown underAlnus and natural forest

A study on dry matter production and nutrient cycling in agroforestry systems of cardamom grown under N₂-fixingAlnus and mixed tree species (non-N₂-fixing) was carried out in the Sikkim Himalaya. The stand total biomass, and tiller number, basal area and biomass of cardamom crop was much higher under the influence ofAlnus. Annual net primary productivity ofAlnus trees was slightly higher than mixed tree species in spite of lower stand density ofAlnus. The agronomic yield of cardamom increased by 2.2 times under the canopy ofAlnus. Litter production and its disappearance rates were also higher in theAlnus-cardamom stand. Nitrogen and phosphorus concentrations of different components ofAlnus were higher than those of mixed tree species, whereas their back translocation from leaf before abscission was lower inAlnus. The cardamom based agroforestry system under the influence ofAlnus was more productive with faster rates of nutrient cycling. The poor nutrient conservation and low nutrient use efficiency ofAlnus, and malleability of nutrient cycling under its influence make it an excellent association which promotes higher availability and faster cycling of nutrients.     

Cardamom, mandarin and nitrogen-fixing trees in agroforestry systems in India's Himalayan region. I. Litterfall and decomposition

Litterfall and decomposition were studied in agroforestry systems involving large cardamom (Amomum subulatum) and mandarin (Citrus reticulata) in the Sikkim Himalaya, India. There were stands with N₂-fixing trees (Alnus nepalensis over large cardamom, and Albizia stipulata over mandarin agroforestry) or without them (native non-symbiotic mixed tree species) in both systems. The total annual litter (litter + crop residue) production was higher in the Alnus-cardamom than in the forest-cardamom stand and in the mandarin than in the Albizia-mandarin stand. The ratio of litter production to floor litter was higher in the N₂-fixing stands than in the non-N₂-fixing stands, indicating a faster litter turnover in the former. Tree litterfall occurred throughout the year, but with marked peaks during November to April. Total soluble polyphenolics of fresh litter were higher in N₂-fixing species than in mixed tree species and crops. Half-life values for ash-free mass were shortest in the leaves of N₂-fixing species. N loss was higher from N₂-fixing Alnus and Albizia leaves, whereas P loss was faster and nearly equal in Alnus leaf litter and cardamom residue in cardamom, and Albizia leaf litter and crop residue in mandarin agroforestry systems. The P turnover in N₂-fixing Alnus and Albizia twigs was faster than in the twigs of mixed tree species. The N₂-fixing tree species increased the N and P cycling through production of more above-ground litter and influenced greater release of these nutrients.     

Management of secondary tree species in agroforestry systems to improve production sustainability in Amazonian Ecuador

Three production systems (robusta coffee, cattle pastures and swidden agriculture), all associated with trees, have provided a strategy for regional development in the Ecuadorian Amazon. Feedback from more than 250 on-farm demonstrations, project evaluations and visitors have all contributed to strengthening the ability of the Ecuadorean Ministry of Agriculture's Forestry Directorate to provide colonists and indigenous groups with management guidelines for low-input sustainable agroforestry production. The existing market for 20 fastgrowing native tree species presents an opportunity for the diversification of farm income (currently dependent on robusta coffee) and for the development of a forest industrial base.The collective area of land owned and operated by these small holders constitutes a national resource and a Collective Forest Estate that can be individually managed. Commercial timber production in these extensive secondary forests can help alleviate conversion of primary forest without taking agricultural land out of production.     

Plant growth, biomass production and nutrient accumulation by slash/mulch agroforestry systems in tropical hillsides of Colombia

Planted fallow systems under ‘slash and mulch’ management were compared with natural fallow systems at two farms (BM1 and BM2) in the Colombian Andes. The BM1 site was relatively more fertile than the BM2 site. Planted fallow systems evaluated included Calliandra calothyrsus CIAT 20400 (CAL), Indigofera constricta (IND) or Tithonia diversifola (TTH). During each pruning event slashed biomass was weighed, surface-applied to the soil on the same plot and sub-samples taken for chemical analyses. While Indigofera trees consistently showed significantly greater (p < 0.05) plant height and collar diameter than Calliandra trees at both study sites, only collar diameter in Indigofera was significantly affected at all sampling times by differences between BM1 and BM2. After 27 months, TTH presented the greatest cumulative dry weight biomass (37 t ha^–1) and nutrient accumulation in biomass (417.5 kg N ha^–1, 85.3 kg P ha^–1, 928 kg K ha^–1, 299 kg Ca ha^–1 and 127.6 kg Mg ha^–1) among planted fallow systems studied at BM1. Leaf biomass was significantly greater (P < 0.05) for CAL than IND irrespective of site. However, CAL and IND biomass from other plant parts studied and nutrient accumulation were generally similar at BM1 and BM2. At both sites, NAT consistently presented the lowest biomass production and nutrient accumulation among fallow systems. Planted fallows using Calliandra and Indigofera trees had the additional benefit of producing considerable quantities of firewood for household use. This revised version was published online in June 2006 with corrections to the Cover Date.     

Litter production and nutrient cycling in coffee (Coffea arabica) or cacao (Theobroma cacao) plantations with shade trees

The relative importance of N fixation, organic material inputs and nutrient inputs in litterfall, as justifications for including shade trees in plantations of coffee or cacao, is discussed. According to existing data, N fixation by leguminous shade trees does not exceed 60 kg.N/ha/a. However, these trees contribute 5,000–10,000 kg. organic material/ha/a.Comparisons are made between the leguminous shade tree Erythrina poeppigiana and the non-leguminous timber tree Cordia alliodora. The former, when pruned 2 or 3 times/a., can return to the litter layer the same amount of nutrients that are applied to coffee plantations via inorganic fertilizers, even at the highest recommended rates for Costa Rica of 270 kg.N, 60 kg.P, 150 kg.K/ha/a. The annual nutrient return in this litterfall represents 90–100 percent of the nutrient store in above-ground biomass of E. poeppigiana, and hence the consequences of competition with the crop should not be a serious limitation. In the case of C. alliodora, which is not pruned, nutrient storage in the tree stems, especially of K, is a potential limiting factor to both crop and tree productivity.It is concluded that, in fertilized plantations of cacao and coffee, litter productivity is a more important shade tree characteristic than N fixation.An early version of this review was presented at the CATIE-IUFRO meeting Los Arboles de Uso Multiple en Sistemas Agroforestales, June 1985, Turrialba, Costa Rica.     

印度恒河平原苏打土壤种植的10个多功能树种生物量和生物能产量(英文)

评价了10个多功能树种--阿江榄江树(Terminalia arjuna)、印楝(Azadirechta indica)、牧豆树(Prosopis juliflora)、水黄皮(Pongarnia pin-nata)、木麻黄(Casuarina equisetifolia)、角豆树(Prosopis alba )、阿拉伯金合欢(Acacia nilotica)、细叶桉(Eucalyptus tereticornis)、牛蹄豆(Pithecellobium dulce)、铁刀木(Cassia siamea )等10年间的生物量和生物能产量,这些树种采用单种栽培的农作方式种植于印度北部恒河平原苏打土壤。该地区土壤紧密、盐碱化、透水性差、营养匮乏。10个树种中,细叶桉植株最高,牧豆树和木麻黄次之。阿拉伯金合欢林分基面积为3.04m2·hm-2,植株胸径优于其他树种,牧豆树和木麻黄植株胸径次之。10年间,牧豆树和Acacia nilotica分别产出生物量56.50和50.75 Mg·hm-2,而阿拉伯金合欢、水黄皮和角豆树生物量产出量较小。牧豆树的营养需求和净生物量产出均最大,而且叶片中营养元素(N,P,K,Ca和Mg)含量较高。然而,树种间木质成分差异较小。以每吨木材营养元素产出量计算,氮素在阿拉伯金合欢中产出量最低,磷和钾分别在阿江榄仁树和牛蹄豆中产出最低,钙和镁在牧豆树中产出最低。牧豆树生物质能产量最高1267.75 GJ·hm-2,阿拉伯金合欢次之1206 GJ·hm-2,印楝最低(520.66GJ·hm-2)。图2表7参31。     

Impact of tree coppicing on tree-crop competition in parkland and alley farming systems in semiarid Burkina Faso

The aim of this study was to evaluate the effects of tree coppicing on tree-crop competition in farmed parkland and in alley farming, both in semiarid Burkina Faso.Azadirachta indica A. Juss (neem) was studied in the parklands, while neem,Albizia lebbeck (L.) Benth. (albizia) andLeucaena leucocephala (Lam.) de Wit (leucaena) were investigated in the alley farming system. The crop was sorghum (Sorghum bicolor (L.) Moench) in both cases.Under non-coppiced neem trees in the parkland, the sorghum yield at 0–3 m distance from the tree (under the tree canopy) was 82% of open field yields at 6–9 m distance, but the difference was not significant (p=0.07). Close to trees (0–3 m distance) which had been coppiced before planting, the sorghum yield was 148% of open field yields. In alley farming, early coppicing was superior; sorghum yields were highest close to trees coppiced early and lowest close to trees coppiced late.

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Résumé La présente étude se propose d'évaluer les effets de la taille sur la concurrence entre arbres et cultures dans les parcs arborés et dans les cultures en couloir, en climat semiaride (Burkina Faso). Les espèces étudiées ont été:Azadirachta indica A. Juss (neem) dans les parcs et neem,Albizia lebbeck (L.) Benth. (albizia) ainsi queLeucaena leucocephala (Lam.) de Wit (leucaena) dans les cultures en couloir. Dans les deux cas la plante cultivée concerné a été le sorgho blanc (Sorghum bicolor (L.) Moench).Sous les neems non taillés des parcs, le rendement du sorgho cultivé de 0 à 3 m de l'arbre (sous le bouppier de l'arbre) a atteint 82% du rendement obtenu en plein champ, soit à une distance de 6 à 9 m de l'arbre (différence non significative,p=0.07). Une taille effectuée en début de campagne a entraîné un accroissement du rendement de 148% pour les cultures situées de 0 à 3 m de l'arbre, par rapport aux rendements en plein champ.Dans les cultures en couloir, on a constaté qu'une taille precoce est préférable, c'est à dire que les rendements de sorgho ont été supérieurs dans les lignes proches des arbres taillés tôt que dans celles proches des arbres taillés tard.

     

Dry matter partitions and specific leaf weight of soybean change with tree competition in an intercropping system

In 2004 and 2005, the yield, leaf area, dry weight and dry weight partitions of soybeans were determined at the Agroforestry Research Site (ARS) (est. 1987, Ontario, Canada). Soybean was intercropped with poplar (Populus deltoides x nigra DN-177 L., 556 m³crown tree⁻¹), silver maple (Acer saccharinum L., 308 m³), black walnut (Juglans nigra L., 148 m³) and pecan (Carya illinoensis Wangenh., 114 m³), or grown alone (monoculture). Yield of soybean was not different in either year between the monoculture and the black walnut or pecan intercrops. In the poplar and silver maple treatments, yield was 66 and 85% (2004 and 2005) lower than in the monoculture. Despite the fact that different tree species were used, there was a significant negative linear regression between yield and tree crown volume (R ² = 0.76, P = 0.0049 and R ² = 0.93, P < 0.0001 in 2004 and 2005, respectively). With increasing tree crown volume, soybean tended to partition more dry matter to the photosynthetic and reproductive parts and less to structural tissue and petiole. This demonstrates the phenotypic flexibility of the crop component in agroforestry systems. Contrary to theoretical predictions, soybean leaves were thicker as shade increased (increase by 6.2 × 10⁻⁴–1.2 × 10⁻³ mg cm⁻², per unit of crown volume), pointing to competitive interactions specific to tree-based intercrops.     

Biomass and production of fine and coarse roots of trees under agrisilvicultural practices in north-east India

An understanding of the rooting pattern of tree species used in agroforestry systems is essential for the development and management of systems involving them. Seasonal variation, depth wise and lateral distribution of biomass in roots of different diameter classes and their annual production were studied using sequential core sampling. The investigations were carried out in four tree species under tree only and tree+crop situations at ICAR Research Farm, Barapani (Meghalya), India. The tree species were mandarin (Citrus reticulata), alder (Alnus nepalensis), cherry (Prunus cerasoides) and albizia (Paraserianthes falcataria). The contribution of fine roots to the total root biomass ranged from 87% in albizia to 77% in mandarin. The bulk of the fine roots (38% to 47%) in the four tree species was concentrated in the upper 10 cm soil layer, but the coarse roots were concentrated in 10–20 cm soil depth in alder (46%) and albizia (51%) and at 0–10 cm in cherry (41%) and mandarin (48%). In all the four tree species, biomass of both fine- and coarse-roots followed a unimodal growth curve by showing a gradual increase from spring (pre-rainy) season to autumn (post rainy) season. Biomass to necromass ratio varied between 2 to 3 in the four tree species. The maximum (3.2) ratio was observed during spring and the minimum (2) in the rainy season. In alder and albizia, the fine roots were distributed only up to 1 m distance from the tree trunk but in the other two species they were found at a distance up to 1.5 m from the tree trunk. The annual fine root production varied from 3.6 Mg ha^–1 to 6.2 Mg ha^–1 and total production from 4.2 to 8.4 Mg ha^–1 in albizia to mandarin, respectively. Cherry and mandarin had a large number of woody roots in the surface layers which pose physical hindrance during soil working and intercultural operations under agroforestry. But the high biomass of roots of these two species may be advantageous for sequential or spatially separated agroforestry systems. However, alder and albizia have the most desirable rooting characteristics for agroforestry systems.This revised version was published online in November 2005 with corrections to the Cover Date.     

Crop yields of rice and wheat grown in rotation as intercrops with three tree species in the outer hills of Western Himalaya

Studies on tree crop interaction under rainfed condition in Dehradun valley were conducted for 13 years during 1977 to 1990. Grewia optiva (Bhimal), Morus alba (Mulbery) and Eucalyptus hybrid were tried along with rice (CV: Akashi) — wheat (CV: RR-21) rotation. One-year-old tree seedlings of the above tree species were planted in line, 5 m apart in N-S direction, in July 1977, in the middle of the plot (size 20 m × 20 m). Eucalyptus was first harvested in 1987. Grewia optiva, Morus alba and coppice of Eucalyptus were harvested in 1990. All tree species had depressing effect on crop yields. Eucalyptus had maximum effect in depressing crop yield till the first harvest and had least effect thereafter. From 1987 onwards, Morus alba affected rice most, while wheat was mostly affected by Grewia optiva. The depressing effect on an average varied from 28 to 34% depending upon the species.Distance of tree line from the crop significantly affected the crop yield upto a distance of 5 m and there was 39% decrease in crop yield upto 1 m, 33% from 1–2 m, 25% from 2–3 m and 12% from 3–5 m distance. Annual removal of lops and tops from trees partly compensated the deficit. Grewia optiva could produce 1.08 t ha^–1 yr^–1 of branches and 0.26 t ha^–1 yr^–1 of leaves (air dry) and 1.28 t ha^–1 yr^–1 of branches and 0.28 t ha^–1 yr^–1 of leaves were obtained from Morus alba. Wood (ADT) produced by the trees was 33.6 t ha^–1 from Eucalyptus, 9.5 t ha^–1 from Grewia optiva and 11.6 t ha^–1 from Morus alba.     

Nutrient use efficiency and biomass production of tree species for rotational woodlot systems in semi-arid Morogoro,Tanzania

Frequent nutrient removals accompanying wood and crop harvests from rotational woodlot systems may contribute to declining site productivity and sustainability because of soil nutrient depletion. However, selecting for nutrient-efficient tree species may well sustain productivity under this system. To test this hypothesis, a randomized complete block experiment was adopted to assess effects of five tree species on soil nutrients status, nutrient use efficiency and wood yield in semi-arid Tanzania. After 5 years rotation, top soils under Gliricidia sepium (Jaqua), Acacia polyacantha Willd. and Acacia mangium Willd. were the most fertile with soil organic carbon and exchangeable cation status raised close to those in natural Miombo systems. Soil inorganic N and extractable P levels reached sufficiency levels for subsequent maize culture. Wood productivity in tree fallows averaged three times higher than that of Miombo woodlands indicating the high potential of the woodlot system to supply fuelwood, and consequently relieve harvesting pressures on the natural forests. Acacia crassicarpa A. Cunn. ex Benth. produced the most wood (51 Mg ha⁻¹) at low nutrient “costs” presumably due to high nutrient use efficiency. Wood yield of this species was 42 and 120% greater than that of A. polyacantha and A. nilotica, respectively, but contained comparatively less nutrients (42–60% less for P, K, and Ca). Gliricidia sepium and A. polyacantha returned the largest amount of nutrients through slash at harvests. Of all test species, A. crassicarpa exhibited the most promise to sustain wood production under rotational woodlot systems due to relatively high productivity and low nutrient export at harvest.     

Survival,growth and productivity of tree species under evaluation for agroforestry to control salinity in the Western Australian wheatbelt

Survival and growth were investigated for 10Eucalyptus species and 2 fodder tree species planted for salinity control in the wheatbelt of Western Australia. After two years of growth the trees were harvested to determine fodder biomass production and yields of cineole from the eucalyptus leaf oil. Subsequent harvests were conducted at three and five years after planting.At each harvest, biomass production from fodder species was greater than from most species of eucalypts. Biomass yields from eucalypts were variable, and there were no consistent trends in the productivity of the different species for the three harvests.Leaf cineole concentrations and cineole yields were low after two years of growth, but after three and five years cineole yields were generally higher from all species.E. kochii subspp.plenissima andkochii, E. horistes, E. radiata andE. angustissima produced consistently high cineole yields after three and five years. These species appear to have potential for the production of high grade eucalyptus oil in the wheatbelt of Western Australia.     

Radiation use efficiency in adjacent hardwood and pine forests in the southern Appalachians

The efficiency with which trees convert photosynthetically active radiation (PAR) to biomass has been shown to be consistent within stands of an individual species, which is useful for estimating biomass production and carbon accumulation. However, radiation use efficiency (?) has rarely been measured in mixed-species forests, and it is unclear how species diversity may affect the consistency of ?, particularly across environmental gradients. We compared aboveground net primary productivity (ANPP), intercepted photosynthetically active solar radiation (IPAR), and radiation use efficiency (? = ANPP/IPAR) between a mixed deciduous forest and a 50-year-old white pine (Pinus strobus L.) plantation in the southern Appalachian Mountains. Average ANPP was similar in the deciduous forest (11.5 Mg ha⁻¹ y⁻¹) and pine plantation (10.2 Mg ha⁻¹ y⁻¹), while ? was significantly greater in the deciduous forest (1.25 g MJ⁻¹) than in the white pine plantation (0.63 g MJ⁻¹). Our results demonstrate that late-secondary hardwood forests can attain similar ANPP as mature P. strobus plantations in the southern Appalachians, despite substantially less annual IPAR and mineral-nitrogen availability, suggesting greater resource-use efficiency and potential for long-term carbon accumulation in biomass. Along a 260 m elevation gradient within each forest there was not significant variation in ?. Radiation use efficiency may be stable for specific forest types across a range of environmental conditions in the southern Appalachian Mountains, and thus useful for generating estimates of ANPP at the scale of individual watersheds.     

Comparing genetic diversity in agroforestry systems with natural forest: a case study of the important timber tree Vitex fischeri in central Kenya

It is possible that current tree domestication practices undertaken by farmers reduce the genetic base of tree resources on farms, raising concerns regarding the productivity, sustainability and conservation value of agroforestry ecosystems. Here, we assessed possible changes in genetic variation during domestication in the important and heavily utilised timber species, Vitex fischeri Gürke (syn. Vitex keniensis), by comparing geographically proximate forest and farm material in central Kenya. Employing RAPD analysis, a total of 104 polymorphic markers revealed by five arbitrary primers were scored in a total of 65 individuals, 32 from forest and 33 from farmland. Despite concerns of possible genetic erosion, forest and farm stands did not differ significantly in levels of genetic variation, with H values of 0.278 and 0.269, respectively. However, Mantel tests did reveal greater geographically related associative genetic structure among individuals in farm rather than forest material, with r _M values of 0.217 and 0.114, respectively. A more detailed analysis of structure suggested this could be due to local variation in origin of some on-farm trees. Implications of data for the genetic management of V. fischeri stands during farmer-led tree domestication activities are discussed. At present, there appears little reason to reject on-farm V. fischeri as a source of germplasm for future on-farm planting or for conservation purposes, although this situation may change and will require monitoring.     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica III. Cycles of organic matter and nutrients

Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica. For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered. The implications of the models are discussed in detail. Aspects of net primary production in the systems studied are considered. N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana. For part I see Vol. 4, No. 3, 1986 For part II see this issue Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica III. Cycles of organic matter and nutrients

Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica.For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered.The implications of the models are discussed in detail.Aspects of net primary production in the systems studied are considered.N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana.For part I see Vol. 4, No. 3, 1986 For part II see this issueAgroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica II. Cacao and wood production,litter production and decomposition

During 7 years (1979–1985) cacao harvests (beans and husks) have been recorded for the agroforestry systems ofTheobroma cacao underCordia alliodora andErythrina poeppigiana shade trees. The mean oven dry cacao yields were 626 and 712 kg.ha^–1.a^–1 cocoa beans underC. alliodora andE. poeppigiana respectively. Harvests have gradually increased over the years and the plantation has now reached maturity.Annual extraction of N, P, K, Ca and Mg in fruits, which is relatively small, was calculated on the basis of chemical analyses. The following average values were found (kg.ha^–1.a^–1): At the age of 8 years, theC. alliodora trees have reached 26.7 cm diameter (DBH) and 14.0 m in height. Mean annual growth (from age 5 to 7) is 14.6 m³.ha^–1.a^–1.Natural plant residue production has been measured for 4 years (Nov. 1981–Oct. 1985). UnderE. poeppigiana it has reached a value of 8.91 t.ha^–1.a^–1 and underC. alliodora 7.07 t.ha^–1.a^–1. The shade trees have contributed 57 and 47% respectively. Transference and decomposition rates are high and important in the nutrient cycles.The nutrient content of the litter was analysed and corresponding average yearly transfers were (kg.ha^–1.a^–1): For part I see Vol. 4, No. 3, 1986.Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesselschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica II. Cacao and wood production,litter production and decomposition

During 7 years (1979–1985) cacao harvests (beans and husks) have been recorded for the agroforestry systems ofTheobroma cacao underCordia alliodora andErythrina poeppigiana shade trees. The mean oven dry cacao yields were 626 and 712 kg.ha⁻¹.a⁻¹ cocoa beans underC. alliodora andE. poeppigiana respectively. Harvests have gradually increased over the years and the plantation has now reached maturity. Annual extraction of N, P, K, Ca and Mg in fruits, which is relatively small, was calculated on the basis of chemical analyses. The following average values were found (kg.ha⁻¹.a⁻¹): At the age of 8 years, theC. alliodora trees have reached 26.7 cm diameter (DBH) and 14.0 m in height. Mean annual growth (from age 5 to 7) is 14.6 m³.ha⁻¹.a⁻¹. Natural plant residue production has been measured for 4 years (Nov. 1981–Oct. 1985). UnderE. poeppigiana it has reached a value of 8.91 t.ha⁻¹.a⁻¹ and underC. alliodora 7.07 t.ha⁻¹.a⁻¹. The shade trees have contributed 57 and 47% respectively. Transference and decomposition rates are high and important in the nutrient cycles. The nutrient content of the litter was analysed and corresponding average yearly transfers were (kg.ha⁻¹.a⁻¹): For part I see Vol. 4, No. 3, 1986. Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesselschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Rock phosphate and vesicular-arbuscular mycorrhiza effects on growth and nutrient uptake of Faidherbia albida (Del.) seedlings in an alkaline sandy soil

Response of Faidherbia albida (Del.) to five levels of Kodjari rock phosphate (KRP) application (0, 310, 620, 1240 and 2480 ppm P, equivalent to 0, 775, 1550, 3100 and 6200 kg P/ha) and vesicular-arbuscular mycorrhizal fungi (VAMF) Glomus manihotis Howeler, Sieverding & Schenck or Glomus aggregatum Schenck & Smith emend. Koske was evaluated in an alkaline sandy soil. The F. albida seedlings grew poorly without mycorrhizal colonization and without KRP applications. For non-VAM Faidherbia, the maximum growth response and both P and N uptake in shoots was achieved with the 620 ppm P. However, even without KRP application, VAM plants achieved better results in terms of biomass. VAM plants with G. manihotis and G. aggregatum improved plant growth and increased nutrient contents at any KRP application rate. Although mycorrhizal colonization was comparable at all levels of KRP application, the impact of nutrient content of the shoot varied. Finally, VAM plants did not accumulate more biomass than non-VAM plants at 620 ppm P and above. Growth response and mycorrhizal dependency decreased as KRP applied levels increased. These results suggest that VAM Faidherbia seedlings take up more P from soil and KRP than non-VAM.     

The effects of Douglas-fir on tree-specific arthropod communities in mixed species stands with European beech and Norway spruce

The ecological effects of planting exotic Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] in Central Europe are still poorly understood. The aim of this study was to answer the question of whether Douglas-fir affects tree specific arthropod communities in different mature forest types (Douglas-fir, spruce and beech dominated) in Southern Germany. Therefore, arthropod communities of stem and tree crown strata of Douglas-fir and spruce (Picea abies L.) were sampled in the years 1999–2001 using arboreal photo-eclectors and flight interception traps. Statistical analysis was conducted for all species and focused on conifer specialists at three levels: (1) species diversity, (2) guild structure and (3) community structure. Within the stem stratum, species diversity was significantly higher on spruce than on Douglas-fir independent of year and stand composition. This could not be explained by a single feeding guild, rather by species changing strata during the vegetation period. In contrast, species diversity in tree crowns was approximately the same for both conifer species. However, communities in Douglas-fir crowns were conspicuously different from those in spruce crowns, especially in the Douglas-fir dominated stand type. While zoophagous insects exhibited higher activity on Douglas-fir in 2000, xylophagous beetles were more abundant on spruce in 2001. In European beech stands with widely spaced Douglas-fir trees, the site specific and broad-leaved tree related fauna might be maintained. In addition, Douglas-fir with its resource of Adelges cooleyi and crowns that overtop the broad-leaved tree canopy, offer additional resources for several aphidophagous and thermophile species.